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STOMATAL MOVEMENT IN
SUCCULENTS
SUCCULENTS
• Latin word- sucus,
means juice or sap
• Found in water
arid climates
or soil conditions
• absent, reduced, or
cylindrical-to-spherical
leaves
• reduction in the number
of stomata
• stems as the main site
of photosynthesis, rather than
leaves
• compact, reduced, cushion-like,
columnar, or spherical growth
form
• ribs enabling rapid increases in
plant volume and decreasing
surface area exposed to the sun
• waxy, hairy, or spiny outer surface
to create a humid micro-habitat
around the plant, which reduces
air movement near the surface of
the plant, and thereby reduces
water loss and creates shade
• roots very near the surface
of the soil, so they are able
to take up moisture from
very small showers or even
from heavy dew
• ability to remain plump and
full of water even with high
internal temperatures (e.g.,
52 °C or 126 °F)
• very impervious
outer cuticle (skin)
• mucilaginous substances,
which retain water
abundantly
Sunken stomata in succulents
Crassulacean Acid Metabolism
(CAM) in succulents
• Observed by Botanists Rason and Thomas in
late 1940s
• Its name refers to acid metabolism in
Crassulaceae not the Crassulacean acid.
What is crassulacean Acid
Metabolism?
• It is a carbon pathway
• Also known as CAM photosynthesis
• These plants fix carbon dioxide (CO2) during the
night, storing it as the four carbon acid malate
• The CO2 is released during the day, where it is
concentrated around the enzyme RuBisCO,
increasing the efficiency of photosynthesis.
• The CAM pathway allows stomata to remain shut
during the day, reducing evapotranspiration;
therefore it is espicially common in plants
adapted to arid conditions.
• CAM plants is subset of C4 plants
• The fixation now occurs in mesophyll cells so that
they will be more exposed to the air & in order to
take in more CO2.
• The malate, which is the product of the fixation
process is pumped deeper in the leaf so that it
won’t be exposed to air and to oxygen.
• This is to avoid photorespiration and the wasteful
process since RuBisCo is used in the Calvin cycle
• The process is a lot like the C4 pathway.
Overview of CAM
A two part cycle
During night
• CAM plant’s stomata are open ,allowing CO2
to enter and be fixated as organic acids that are
stored in vacuoles.
• The carbon dioxide is fixed in the mesophyll
cell’s cytoplasm by PEP reaction
• PEP- Phosphoenolpyruvic acid
During day
• During the day the stomata are closed and the
carbon is released to the Calvin cycle so that
photosynthesis may take place.
• The carbon is the organic acids is freed from
the mesophyll cell’s vacuoles and enters the
chloroplast’s stoma and into Calvin cycle .
Comparison chart
Plant characters C3 pathway C4 pathway CAM pathway
Photorespiration
rate
High Low/ negligible Very low/ negligible
Leaf anatomy Typical Kranz Xeromorphic
Typical
environment
All Tropical, elevated
daytime
temperature,
drought
Dry, arid
Stoma open during
the day?
Yes Yes No
No. of steps in
pathway
1 2 2
First molecule
produced in
pathway
3-phosphoglyceric
acid
Malic acid or
aspartic acid
Malate
Uses the Calvin
cycle
yes yes Yes
Stomatal movement in succulents   copy

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Stomatal movement in succulents copy

  • 2. SUCCULENTS • Latin word- sucus, means juice or sap • Found in water arid climates or soil conditions • absent, reduced, or cylindrical-to-spherical leaves • reduction in the number of stomata
  • 3. • stems as the main site of photosynthesis, rather than leaves • compact, reduced, cushion-like, columnar, or spherical growth form • ribs enabling rapid increases in plant volume and decreasing surface area exposed to the sun • waxy, hairy, or spiny outer surface to create a humid micro-habitat around the plant, which reduces air movement near the surface of the plant, and thereby reduces water loss and creates shade
  • 4. • roots very near the surface of the soil, so they are able to take up moisture from very small showers or even from heavy dew • ability to remain plump and full of water even with high internal temperatures (e.g., 52 °C or 126 °F) • very impervious outer cuticle (skin) • mucilaginous substances, which retain water abundantly
  • 5. Sunken stomata in succulents
  • 6. Crassulacean Acid Metabolism (CAM) in succulents • Observed by Botanists Rason and Thomas in late 1940s • Its name refers to acid metabolism in Crassulaceae not the Crassulacean acid.
  • 7. What is crassulacean Acid Metabolism? • It is a carbon pathway • Also known as CAM photosynthesis • These plants fix carbon dioxide (CO2) during the night, storing it as the four carbon acid malate • The CO2 is released during the day, where it is concentrated around the enzyme RuBisCO, increasing the efficiency of photosynthesis. • The CAM pathway allows stomata to remain shut during the day, reducing evapotranspiration; therefore it is espicially common in plants adapted to arid conditions.
  • 8. • CAM plants is subset of C4 plants • The fixation now occurs in mesophyll cells so that they will be more exposed to the air & in order to take in more CO2. • The malate, which is the product of the fixation process is pumped deeper in the leaf so that it won’t be exposed to air and to oxygen. • This is to avoid photorespiration and the wasteful process since RuBisCo is used in the Calvin cycle • The process is a lot like the C4 pathway.
  • 9. Overview of CAM A two part cycle
  • 10. During night • CAM plant’s stomata are open ,allowing CO2 to enter and be fixated as organic acids that are stored in vacuoles. • The carbon dioxide is fixed in the mesophyll cell’s cytoplasm by PEP reaction • PEP- Phosphoenolpyruvic acid
  • 11.
  • 12. During day • During the day the stomata are closed and the carbon is released to the Calvin cycle so that photosynthesis may take place. • The carbon is the organic acids is freed from the mesophyll cell’s vacuoles and enters the chloroplast’s stoma and into Calvin cycle .
  • 13.
  • 14. Comparison chart Plant characters C3 pathway C4 pathway CAM pathway Photorespiration rate High Low/ negligible Very low/ negligible Leaf anatomy Typical Kranz Xeromorphic Typical environment All Tropical, elevated daytime temperature, drought Dry, arid Stoma open during the day? Yes Yes No No. of steps in pathway 1 2 2 First molecule produced in pathway 3-phosphoglyceric acid Malic acid or aspartic acid Malate Uses the Calvin cycle yes yes Yes