2. Environmental stress factors
Types of environmental factors
Biotic
Abiotic
Nutrient stresses come under abiotic factor.
Stress reactions resulting from sub-optimal and damaging
quantities.
Nutrient Stress deviation from the normal physiological
processes as a result of sub-optimal or toxic
concentrations of nutrition.
3. Environmental noxae: stress factors which trigger stress
reactions when applied in any concentration, e.g. UV-B,
Ozone, ionizing radiations, heavy metals, aluminum etc.
Specific and un-specific reactions to stress: un-specific
reactions includes e.g. modification of basic metabolism.
specific reactions include e.g. the production of heat
shock proteins.
currently 15-17 elements are known to be essential for
plant growth, but the role in the physiological metabolism
is not known in detail for all of them.
4. General effects of available nutrient
concentration
The different ranges can be:
a) Extreme deficiency range
b) severe deficiency range
c) moderate deficiency range
d) luxury range
e) Toxic range
Critical concentration range: it occurs between deficiency
and luxury consumption.
This information is useful for the measurement of the
degree of stress in a plant.
5. Conditions for nutrient deficiency
stress
a) Amount and concentration of nutrients in the soil.
b) Form of the soil.
c) The contents of the soil solutions.
d) Soil pH.
Nutrient concentration in plants:
these vary with
a) Plant age
b) Plant part
c) Plant species
d) Soil type
6. Deficiency symptoms
Symptom: is any perceptible change in known structure,
appearance or function.
These include:
Yellowing (chlorosis)
Death (necrosis)
Lesions
Malformations
Malfunctions
Reduced growth and yield
7. Confusion:
Occurs when one realizes that similar symptoms can be
seen by
a)Toxic levels of nutrients
b)Pathogenic organisms
c)Air pollutants
d)Pesticides
Symptoms may vary with plant species or variety.
8. Important Point to remember for
Stress diagnosis
Yield or growth rate is usually impaired before other
symptoms of deficiency occur so that some other
indications of at least the potential for a deficiency must
be considered in any diagnosis procedure.
9. Important Definitions
An element is a pure chemical that contains
only one type of atom.
A compound is a molecule that contains one
or more elements.
10. Essential Elements
The 16 elements required by plants are obtained
from the soil, water and air.
Thirteen of these elements must be supplied by the
soil.
Six of the soil elements required by plants are
needed in relatively large amounts and are usually
added to the soil through fertilizer or lime. These
are called macronutrients.
The remaining 7 elements supplied by soil are
required in very small amounts and are termed
micronutrients.
11. Macronutrients
Macronutrients are
needed in
relatively large
amounts by plants.
Element Symbol Source
Oxygen O Air/Water
Hydrogen H Air/Water
Carbon C Air/Water
Nitrogen N Soil
Phosphorus P Soil
Sulfur S Soil
Potassium K Soil
Calcium Ca Soil
Magnesium Mg Soil
12. Macronutrients – Form Used By Plants
Nutrient Form Used
Carbon CO2
Oxygen H2O
Hydrogen H2O
Nitrogen NO3
-
, NH4
+
Phosphorus H2PO4 & HPO4
2-
Potassium K+
Calcium Ca2
+
Magnesium Mg2+
Sulfur SO4
-
Charge on the
molecule
13. Micronutrients
Micronutrients are
needed in
relatively small
amounts by plants.
They are usually
supplied by
fertilizers such as
iron chelate fertilizer
Element Symbol Source
Iron Fe Soil
Manganese Mn Soil
Boron B Soil
Molybdenum Mo Soil
Copper Cu Soil
Zinc Zn Soil
Chlorine Cl Soil
14. Micronutrients – Form Used By
PlantsNutrient Form Used
Iron Fe2+
Manganese Mn2+
Boron H2B03
-
Molybdenum MoO4
2-
Copper Cu2+
Zinc Zn2+
Chlorine Cl-
Charge on the
molecule
15. Nutrient Deficiency Symptoms
Nutrient deficiency symptoms usually appear on the
plant when one or more nutrients are in short supply.
In many cases, deficiency may occur because an added
nutrient is not in the form the plant can use.
Deficiency symptoms for specific elements are included
on the "Key to Nutrient Disorders".
16. Nutrient Deficiencies
Every soil is not susceptible to the same nutrient
deficiencies.
For example, coarse-textured soils low in organic matter
are susceptible to sulfur deficiencies whereas sulfur is
usually in adequate supply in clayey soils or soils high in
organic matter.
17. Macronutrient Deficiencies & Soils
Element Soil Factor Causing Deficiency
N & K Excessive leaching on coarse-textured low organic matter soils
P Acid low organic matter soils
Cold wet soils such as occurs during early spring
Newly cleared soils
S Excessive leaching on coarse-textured low organic matter soils
in areas where air pollution is low (minimal levels of SO2 in the
air)
Ca & Mg Excessive leaching on coarse-textured low organic matter soils
Soils where large amounts of K have been applied
18. Micronutrient Deficiencies & Soils
Element Soil Factor Causing Deficiency
Fe Poorly drained soils, Low organic matter soils, pH>7.0, Soils
high in P
Zn Cold wet soils low in organic matter and highly leached, High
pH soils (pH>7.0), Soils high in P, Exposed subsoils
Cu Peat and muck soils, High pH, sandy soils, Soils heavily
fertilized with N
B Excessive leaching on coarse-textured low organic matter soils,
Soils with pH>7.0
Mn Excessive leaching on coarse-textured low organic matter soils,
Soil with pH>6.5
Mo Soils high in Fe oxides, high adsorption of molybdenum, Soil
cropped for a long time
19. Deficiency Symptoms - N
General chlorosis.
Chlorosis progresses
from light green to yellow.
Entire plant becomes
yellow under prolonged
stress.
Growth is immediately
restricted and plants soon
become spindly and drop
older leaves. http://plantsci.sdstate.edu/woodardh/soilfert/Nutrient_Defi
ciency_Pages/soy_def/SOY-N1.JPG
20. Deficiency Symptoms - P
Leaves appear dull, dark
green, blue green, or red-
purple, especially on the
underside, and especially
at the midrib and vein.
Petioles may also exhibit
purpling. Re-striction in
growth may be noticed.
http://www-unix.oit.umass.edu/~psoil120/images/tomatox2.jpg
http://www.ext.vt.edu/news/periodicals/viti
culture/04octobernovember/photo3.jpg
21. Deficiency Symptoms - K
Leaf margins tanned,
scorched, or have necrotic
spots (may be small black
spots which later coalesce).
Margins become brown and
cup downward.
Growth is restricted and die
back may occur.
Mild symptoms appear first
on recently matured leaves.
http://www.ipm.iastate.edu/ipm/icm/files/images/antonio004f.jpg
22. Deficiency Symptoms - Ca
Growing points usually
damaged or dead (die
back).
Margins of leaves
developing from the
growing point are first
to turn brown.
http://hubcap.clemson.edu/~blpprt/acid_photos/Blossom
EndRot.JPG
23. Deficiency Symptoms - Mg
Marginal chlorosis or
chlorotic blotches which
later merge.
Leaves show yellow
chlorotic interveinal tissue
on some species, reddish
purple progressing to
necrosis on others.
Younger leaves affected
with continued stress.
Chlorotic areas may
become necrotic, brittle,
and curl upward.
Symptoms usually occur
late in the growing season.
http://quorumsensing.ifas.ufl.edu/HCS200/images/deficie
ncies/-Mgcq.jpg
24. Deficiency Symptoms - S
Leaves uniformly light green, followed by
yellowing and poor spindly growth.
Uniform chlorosis does not occur
http://www.ag.ndsu.nodak.edu/aginfo/entomol
ogy/ndsucpr/Years/2007/june/7/soils.jpg
http://www.ces.ncsu.edu/plymouth/cropsci/gr
aphics/sulfur2.jpg
25. Deficiency Symptoms - Cu
Leaves wilt, become
chlorotic, then necrotic.
Wilting and necrosis
are not dominant
symptoms.
http://images.google.com/url?
q=http://ipm.ncsu.edu/Scouting_Small_Grains/Grain_ima
ges/fig4.jpg&usg=AFQjCNE2vzRwrqp65VR_xKRlo2LQO
gWI3g
26. Symptoms of Iron Deficiency in plants
Iron Deficiency in plants
Distinct yellow or white
areas appear between
veins, and veins
eventually become
chlorotic.
Symptoms are rare on
mature leaves.
http://bexar-
tx.tamu.edu/HomeHort/F1Column/2003Article
s/Graphics/iron%20chlorosis.jpg
27. Deficiency Symptoms - Mn
Chlorosis is less
marked near veins.
Some mottling occurs
in interveinal areas.
Chlorotic areas
eventually become
brown, transparent, or
necrotic.
Symptoms may appear
later on older leaves.
http://www.ca.uky.edu/HLA/Dunwell/KHC/110-122.JPG
28. Deficiency Symptoms - Zn
Leaves may be abnormally small and necrotic.
Internodes are shortened.
http://plantsci.sdstate.edu/woodardh/soilfert/Nu
trient_Deficiency_Pages/corn_def/CORN-
ZN1.JPG
http://agri.atu.edu/people/Hodgson/FieldCrop
s/Mirror/Nutrient%20Def_files/slide24.jpg
29. Deficiency Symptoms - B
Young, expanding
leaves may be necrotic
or distorted followed by
death of growing points.
Internodes may be
short, especially at
shoot terminals.
Stems may be rough,
cracked, or split along
the vascular bundles. http://www.canr.msu.edu/vanburen/ffc12.jpg
30. Crops Highly Susceptible to Deficiencies
Element Crops
Mn Soybean, Small Grain & Peanuts
Cu Wheat & Corn
Zn Corn
Mo Soybeans & Cauliflower
B Alfalfa, Apples, Peanuts, Tobacco &
Tomatoes
Fe Ornamentals, Fruit Trees, Soybeans, Grain
Sorghum & Some Grasses