Colony collapse disorder (CCD) describes the phenomenon where the majority of worker bees in a colony disappear, leaving the queen alone with a few nurse bees. CCD was first discovered in 1869 and given different names over time. A collapsed colony from CCD is characterized by the presence of capped brood, food, and the queen bee, but absence of most worker bees. Reports of disappearing honey-producing hives have decreased their numbers in the United States significantly since the 1940s. The exact causes of CCD are unknown but may include pesticides, mites, fungi, beekeeping practices, and commercial bee transport, acting together rather than individually.