Engler and Prantl system of classification in plant taxonomy
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General features and structure of cyanobacteria
1. General Characters and Structure of
Cyanobacteria
Dr. V. RAMESH
Assistant Professor & Head In-Charge,
Department of Botany,
Vivekananda College,
Tiruvedakam West – 625 234
Madurai
2. Cyanobacteria
Definition: Cyanobacteria represents the major group
of photosynthetic bacteria that carry out oxygenic
photosynthesis. It releases oxygen and uses water as
an electron-donating substrate, i.e. splits water
molecule to release oxygen. Other than this, they are
also capable of nitrogen-fixing.
Classification of Cyanobacteria
Kingdom: Monera
Division: Eubacteria
Class: Cyanobacteria
Examples: Nostoc sp., Oscillotoria sp., Spirulina sp.,
Lyngbya sp., Azolla sp., Anaebina sp., Gleotrichia sp.
3. Characteristics of Cyanobacteria
Cyanobacteria are prokaryotic, aerobic and photoautotrophic
organisms. Its size ranges from 1-10µm. It contains a primitive
nucleus. Membrane-bound organelles are absent in
cyanobacteria. Chlorophyll a, phycocyanin and phycoerythrin
are the photosynthetic pigments
Chlorophyll a: Imparts green colour
Phycocyanin: Imparts blue colour
Phycoerythrin: Imparts red colour
As cyanobacteria also called blue-green algae, is due to the
presence of chlorophyll a and phycocyanin. Chlorophyll
involves in photosystem I whereas phycocyanin involves in
photosystem II. Its photosynthesis is “oxygenic”.
Cyanobacteria have the ability to fix atmospheric N2. The
reserve food material is “Cyanophycean starch”.
4. Habitat
These are the most genetically diverse organisms that
show a broad range of habitat. They can live in
freshwater, marine water, damp places, moist soil, hot
springs etc.
Types: Cyanobacteria can be of three kinds: Unicellular,
colonial and filamentous.
Unicellular: These are single-celled, having mucilage sheath
covering. Examples: Chroococcus sp.
Colonial: These are multicellular. It forms when single cells
colonise themselves. Examples: Gleocapsa sp.
Filamentous: These founds in the form of a chain, covered by
a sheath. It consists of: Hetero-cysts: Helps in N2 fixation ,
Akinete: Collects the food material. Examples: Nostoc sp.,
Oscillotoria sp. etc.
6. The structural components of cyanobacteria are:
Sheath: envelope of cyanobacteria, mucilaginous, hygroscopic in nature. It contains
“pectic substances”. Its appearance is fibrillar under the microscope.
Function: sheath protects the cell from unfavourable conditions. It keeps the cell
moist.
Cell Wall
It is present between the sheath and plasma membrane. The pores that are present on
the cell wall helps in secretion of mucilage. It consists of four layers represented by L-I,
L-II, L-III and L-IV and interconnects with each other by “Plasmodesmata“.
Plasma Membrane
It is a bilayer membrane, also called “Plasmalemma“. It is 70Å thick, selectively
permeable, maintains the structural integrity of the cell. There is a structure called
“Lamellosomes” forms when plasma membrane invaginates and fuses with lamellae. It
encloses cytoplasm and other cell inclusions.
Cytoplasm
It consists of two regions:
Chromoplasm: It comprises of a vesicular structure called thylakoids, having a capacity
of “Photophosphorylation”.
Centroplasm: It is colourless and regarded as the primitive nucleus (devoid of nuclear
membrane and nucleolus). Some granular structures present inside this.
7. Cytoplasmic inclusions: It includes:
Cyanophycean granules: It is the nitrogenous reserve material. It
constitutes about 8% of the total cell weight.
Gas vacuoles: These are hollow, tough and elongated in shape. Its
diameter is 75nm and length is 200-1000nm.
Carboxysomes: These are polyhedral bodies, contains 1,5-ribulose
biphosphate carboxylase.
Phosphate bodies: These are the volutin granules.
Phycobilisomes: Phycobiliproteins when aggregates into a ball like
structure form phycobilisomes. It is attached to the outer surface of
the thylakoid.
DNA matrix: Naked DNA fibrils dispersed in the cytoplasm, lacks
Nucleoplasm. The base composition of DNA differs. Cyanobacteria
contain covalently closed, non-functional, circular plasmid DNA.