Washoe, a chimpanzee, was taught American Sign Language by Beatrix and Allen Gardner. Washoe learned over 100 signs and could combine them into simple sentences. She also understood many more signs than she produced and could engage in basic question-answer conversations.
Other chimpanzees like Sarah and Lana were taught artificial symbolic communication systems involving shapes or keyboard symbols instead of sign language. Sarah could request food by selecting symbols, while Lana could ask for water by pressing four specific symbols in a keyboard message.
Kanzi, unlike other chimpanzees, learned a symbolic system spontaneously by observing his mother communicate and did not require direct teaching. He developed a large symbolic vocabulary and could understand