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HOW TO QUANTIFY HIERARCHY?
Dániel Czégel
Dept. of Biological Physics,
Eötvös Loránd University, Budapest
Dept. of Cognitive Science, TU Budapest
IFISC
UBIQUITY OF HIERARCHICAL STRUCTURES
2
MULTI-LEVEL MODULARITY OF THE BRAIN
3
Meunier, D., Lambiotte, R., Fornito, A., Ersche, K. D., & Bullmore, E. T. (2009).
Hierarchical modularity in human brain functional networks. Frontiers in neuroinformatics, 3.
MENTAL LEXICON
4
De Deyne, S., Verheyen, S., Perfors, A., & Navarro, D. J. Evidence for widespread thematic structure in the mental lexicon. (2015)
GENE REGULATION, FOOD WEB
5
Ma, H. W., Buer, J., & Zeng, A. P. (2004).
Hierarchical structure and modules in the Escherichia coli
transcriptional regulatory network revealed by a new top-down approach.
BMC bioinformatics, 5(1), 199.
LEADER-FOLLOWER & DOMINANCE HIERARCHY
6
https://www.youtube.com/watch?v=FVBiR7pl71U&feature=youtu.be
https://www.youtube.com/watch?v=QMpIvlKPk3k&feature=youtu.be
Nagy, M., Vásárhelyi, G., Pettit, B., Roberts-Mariani, I., Vicsek, T., & Biro, D. (2013).
Context-dependent hierarchies in pigeons. Proceedings of the National Academy of Sciences, 110(32), 13049-13054.
7
1. What is hierarchical
organization exactly?
2. Why is hierarchical
organization so efficient?
WHAT IS HIERARCHY?
8
 framework: directed (weighted/unweighted) networks
 hierarchy measure:
OUTLINE
 1. Four examples of H
 2. Intuitive requirements for H
 3. Random Walk Hierarchy
9
1. KRACKHARDT’S HIERARCHY
 1. Reachability graph R(G)
 2.
Krackhardt, D. (1994). Graph theoretical dimensions of informal organizations.
Computational organization theory, 89(112), 123-140.
10
2. LINK-FLOW HIERARCHY
Luo, J., & Magee, C. L. (2011). Detecting evolving patterns of self‐organizing networks by
flow hierarchy measurement. Complexity, 16(6), 53-61.
11
~ Reynolds Number
GLOBAL REACHING CENTRALITY
 1. Reachability graph R(G)
 2.
 ~Heterogenity in reachability
Mones, E., Vicsek, L., & Vicsek, T. (2012). Hierarchy measure for complex networks.
PloS one, 7(3), e33799.
12
TREENESS, FEEDFORWARDNESS, ORDERABILITY
 3 dim morphospace; each dim: different aspect of hierarchy
Corominas-Murtra, B., Goñi, J., Solé, R. V., & Rodríguez-Caso, C. (2013). On the origins of hierarchy in complex
networks. Proceedings of the National Academy of Sciences, 110(33), 13316-13321.
13
14
WHAT IS A HIERARCHY MEASURE, THEN?
 1. Based on the global topology; statistics of local
properties (e.g., degree distribution) are not enough!
 2. Intensive:
 3. Minimal for networks where the nodes are
indistinguishable (e.g., directed cycle, full graph)
 4. Maximal for acyclic networks; in particular,
15
HOW TO MEASURE HIERARCHY?
 Goal: to define a hierarchy measure , that
 meets all of these requirements
 is easily generalizable to weighted networks
 is computationally efficient
16
RANDOM WALK HIERARCHY
 From every node, track the
information backwards through
every possible path
 ~ Random walk on the nodes
 Stationary distribution ~ how
much the given node is the source
of the overall information
 Then:
17
Czégel, D., & Palla, G. (2015). Random walk hierarchy measure:
What is more hierarchical, a chain, a tree or a star? Scientific Reports. (arXiv preprint arXiv:1508.07732.)
RANDOM WALK HIERARCHY
 Transition probabilities?
 node j: which node the info have
come from?
 node i: limited resource for
communication
 The two effects combined:
18
RANDOM WALK HIERARCHY
 This way: eventually all random
walker would converge to the root
node(s).
 However, we want to measure the
overall structure, not just the
number of root nodes!
 -> Every step: inject the same
amount of random walkers to
every node, then normalize!
19
RANDOM WALK HIERARCHY
 The process:
 1. Do infinitely many times:
 a) Inject f/N random walkers to each node:
 b) Transition:
 c) Normalize, i.e., divide by 1+f. (~decay of random walkers)
 2. From the stationary distribution:
20
RESULTS: ACYCLIC NETWORKS
 Free parameter: characteristic decay length (or time)
of random walkers:
21
RESULTS: REAL NETWORKS
 What kind of structures
are the most (and least)
hierarchical ones?
 -> Compared to their
randomized counterparts:
22
APPLICATION: FOOD WEB ANALYSIS
23
THANK YOU!
24
The research was partially supported by the European Union and the European Social Fund
through project FuturICT.hu (grant no.:TAMOP-4.2.2.C-11/1/KONV-2012-0013)
and by the Hungarian National Science Fund (OTKA K105447).
czegel_d@yahoo.com

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How to quantify hierarchy?

  • 1. HOW TO QUANTIFY HIERARCHY? Dániel Czégel Dept. of Biological Physics, Eötvös Loránd University, Budapest Dept. of Cognitive Science, TU Budapest IFISC
  • 3. MULTI-LEVEL MODULARITY OF THE BRAIN 3 Meunier, D., Lambiotte, R., Fornito, A., Ersche, K. D., & Bullmore, E. T. (2009). Hierarchical modularity in human brain functional networks. Frontiers in neuroinformatics, 3.
  • 4. MENTAL LEXICON 4 De Deyne, S., Verheyen, S., Perfors, A., & Navarro, D. J. Evidence for widespread thematic structure in the mental lexicon. (2015)
  • 5. GENE REGULATION, FOOD WEB 5 Ma, H. W., Buer, J., & Zeng, A. P. (2004). Hierarchical structure and modules in the Escherichia coli transcriptional regulatory network revealed by a new top-down approach. BMC bioinformatics, 5(1), 199.
  • 6. LEADER-FOLLOWER & DOMINANCE HIERARCHY 6 https://www.youtube.com/watch?v=FVBiR7pl71U&feature=youtu.be https://www.youtube.com/watch?v=QMpIvlKPk3k&feature=youtu.be Nagy, M., Vásárhelyi, G., Pettit, B., Roberts-Mariani, I., Vicsek, T., & Biro, D. (2013). Context-dependent hierarchies in pigeons. Proceedings of the National Academy of Sciences, 110(32), 13049-13054.
  • 7. 7 1. What is hierarchical organization exactly? 2. Why is hierarchical organization so efficient?
  • 8. WHAT IS HIERARCHY? 8  framework: directed (weighted/unweighted) networks  hierarchy measure:
  • 9. OUTLINE  1. Four examples of H  2. Intuitive requirements for H  3. Random Walk Hierarchy 9
  • 10. 1. KRACKHARDT’S HIERARCHY  1. Reachability graph R(G)  2. Krackhardt, D. (1994). Graph theoretical dimensions of informal organizations. Computational organization theory, 89(112), 123-140. 10
  • 11. 2. LINK-FLOW HIERARCHY Luo, J., & Magee, C. L. (2011). Detecting evolving patterns of self‐organizing networks by flow hierarchy measurement. Complexity, 16(6), 53-61. 11 ~ Reynolds Number
  • 12. GLOBAL REACHING CENTRALITY  1. Reachability graph R(G)  2.  ~Heterogenity in reachability Mones, E., Vicsek, L., & Vicsek, T. (2012). Hierarchy measure for complex networks. PloS one, 7(3), e33799. 12
  • 13. TREENESS, FEEDFORWARDNESS, ORDERABILITY  3 dim morphospace; each dim: different aspect of hierarchy Corominas-Murtra, B., Goñi, J., Solé, R. V., & Rodríguez-Caso, C. (2013). On the origins of hierarchy in complex networks. Proceedings of the National Academy of Sciences, 110(33), 13316-13321. 13
  • 14. 14
  • 15. WHAT IS A HIERARCHY MEASURE, THEN?  1. Based on the global topology; statistics of local properties (e.g., degree distribution) are not enough!  2. Intensive:  3. Minimal for networks where the nodes are indistinguishable (e.g., directed cycle, full graph)  4. Maximal for acyclic networks; in particular, 15
  • 16. HOW TO MEASURE HIERARCHY?  Goal: to define a hierarchy measure , that  meets all of these requirements  is easily generalizable to weighted networks  is computationally efficient 16
  • 17. RANDOM WALK HIERARCHY  From every node, track the information backwards through every possible path  ~ Random walk on the nodes  Stationary distribution ~ how much the given node is the source of the overall information  Then: 17 Czégel, D., & Palla, G. (2015). Random walk hierarchy measure: What is more hierarchical, a chain, a tree or a star? Scientific Reports. (arXiv preprint arXiv:1508.07732.)
  • 18. RANDOM WALK HIERARCHY  Transition probabilities?  node j: which node the info have come from?  node i: limited resource for communication  The two effects combined: 18
  • 19. RANDOM WALK HIERARCHY  This way: eventually all random walker would converge to the root node(s).  However, we want to measure the overall structure, not just the number of root nodes!  -> Every step: inject the same amount of random walkers to every node, then normalize! 19
  • 20. RANDOM WALK HIERARCHY  The process:  1. Do infinitely many times:  a) Inject f/N random walkers to each node:  b) Transition:  c) Normalize, i.e., divide by 1+f. (~decay of random walkers)  2. From the stationary distribution: 20
  • 21. RESULTS: ACYCLIC NETWORKS  Free parameter: characteristic decay length (or time) of random walkers: 21
  • 22. RESULTS: REAL NETWORKS  What kind of structures are the most (and least) hierarchical ones?  -> Compared to their randomized counterparts: 22
  • 23. APPLICATION: FOOD WEB ANALYSIS 23
  • 24. THANK YOU! 24 The research was partially supported by the European Union and the European Social Fund through project FuturICT.hu (grant no.:TAMOP-4.2.2.C-11/1/KONV-2012-0013) and by the Hungarian National Science Fund (OTKA K105447). czegel_d@yahoo.com