There are approximately 10,000 living bird species globally. Modern birds are characterized by feathers, a beak without teeth, laying hard-shelled eggs, a high metabolism, a four-chambered heart, and a lightweight but strong skeleton. While most birds can fly, some are flightless like ratites and penguins. Birds have specialized respiratory and digestive systems adapted for flight and many species undertake long or short migrations. They communicate visually and vocally and live socially, with extended parental care of eggs and offspring.