Frogs have smooth, moist skin and lack a tail. They breathe through their skin in water and through lungs on land. Their circulatory system includes two-chambered hearts and lungs. They excrete through paired kidneys and a urinary bladder. Male frogs have testes and vocal sacs, while females have ovaries that release eggs into a cloaca for external fertilization. Frogs benefit ecosystems by eating insects and serving as prey.