This document analyzes the duration of enamel formation in anterior teeth of Neanderthals compared to three modern human groups. It finds that while Neanderthal formation times were likely faster than Inuit, they were not faster than those of an English sample and were clearly slower than a southern African sample. This suggests Neanderthal tooth growth, and by extension somatic growth, falls within the range of modern human variation.
Periféricos de comunicación y de almacenamientoNASH Salas
Este documento describe diferentes periféricos de comunicación y almacenamiento para computadoras. Entre los periféricos de comunicación se encuentran fax-módems, tarjetas de red, tarjetas inalámbricas y controladores de puertos como puertos paralelos, serie e infrarrojo. Los periféricos de almacenamiento mencionados incluyen discos duros, CDs, DVDs, discos flexibles y sus características como duración, densidad, reutilización y tipo de acceso.
Human chorionic gonadotropin (hCG) is best known as the hormone detected in pregnancy tests, but it has many other roles and functions beyond early pregnancy. While hCG was long thought to only function in supporting the corpus luteum during pregnancy, research over the past 20 years has shown that hCG regulates numerous non-gonadal tissues through its receptors. These regulatory roles include supporting fetal growth and development, safeguarding the fetus from rejection, and allowing delivery when the fetus is mature. The broad regulatory roles of hCG are not entirely unexpected, as many other hormones also have multiple sites of action beyond their main targets. The evolution of these multiple hormone functions is thought to provide functional redundancy and efficiency in response
1) The document discusses how it was previously thought that Alzheimer's disease (AD) development in elderly women was due to decreasing estrogen levels after menopause. However, recent evidence suggests it may instead be due to a progressive increase in luteinizing hormone (LH) levels.
2) LH has been shown to promote the amyloidogenic pathway which leads to amyloid beta plaque formation in the hippocampus, a brain region involved in AD. Blocking LH levels has shown therapeutic benefits.
3) The document provides background on AD symptoms and pathology, such as amyloid plaques and neurofibrillary tangles. It also discusses the shift in understanding of LH's actions in the brain through its receptors, such as
El documento describe la Web 2.0, incluyendo sus características principales como ser más interactiva y dinámica, permitiendo una mayor participación de los usuarios a través de herramientas como blogs, wikis y redes sociales. También discute los tipos de aplicaciones Web 2.0 como redes sociales, generación de contenido y herramientas, y concluye resumiendo 10 cosas importantes que debemos saber sobre la Web 2.0 como ser una plataforma social, flexible y participativa.
SHREE SIDDHI SOFT SOLUTIONS is one of the best ieee project centers in Trichy, SHREE SIDDHI SOFT SOLUTIONS offers IEEE 2016 Final Year projects for Engineering students in JAVA, Dot Net, Android, Oracle,matlab,embedded system and PHP technologies, SHREE SIDDHI SOFT SOLUTIONS is a one of the leading research and Development in TRICHY
Contact US:
SHREE SIDDHI SOFT SOLUTIONS
#3, 4th FLOOR, LAKSHMI ARCADE,
11TH CROSS, THILLAI NAGAR,
TRICHY-620018.
Phone NO : +91 90032-44446,900323336,0431-4040672
Email : shreesiddhisoftsolutionstrichy@gmail.com
Website : www.shreesiddhisoftsolutions.com
www.siddhisoftsolutions.com
SHREE SIDDHI SOFT SOLUTIONS is one of the best ieee project centers in Trichy, SHREE SIDDHI SOFT SOLUTIONS offers IEEE 2016 Final Year projects for Engineering students in JAVA, Dot Net, Android, Oracle,matlab,embedded system and PHP technologies, SHREE SIDDHI SOFT SOLUTIONS is a one of the leading research and Development in TRICHY
Contact US:
SHREE SIDDHI SOFT SOLUTIONS
#3, 4th FLOOR, LAKSHMI ARCADE,
11TH CROSS, THILLAI NAGAR,
TRICHY-620018.
Phone NO : +91 90032-44446,900323336,0431-4040672
Email : shreesiddhisoftsolutionstrichy@gmail.com
Website :
www.shreesiddhisoftsolutions.com
www.siddhisoftsolutions.com
Periféricos de comunicación y de almacenamientoNASH Salas
Este documento describe diferentes periféricos de comunicación y almacenamiento para computadoras. Entre los periféricos de comunicación se encuentran fax-módems, tarjetas de red, tarjetas inalámbricas y controladores de puertos como puertos paralelos, serie e infrarrojo. Los periféricos de almacenamiento mencionados incluyen discos duros, CDs, DVDs, discos flexibles y sus características como duración, densidad, reutilización y tipo de acceso.
Human chorionic gonadotropin (hCG) is best known as the hormone detected in pregnancy tests, but it has many other roles and functions beyond early pregnancy. While hCG was long thought to only function in supporting the corpus luteum during pregnancy, research over the past 20 years has shown that hCG regulates numerous non-gonadal tissues through its receptors. These regulatory roles include supporting fetal growth and development, safeguarding the fetus from rejection, and allowing delivery when the fetus is mature. The broad regulatory roles of hCG are not entirely unexpected, as many other hormones also have multiple sites of action beyond their main targets. The evolution of these multiple hormone functions is thought to provide functional redundancy and efficiency in response
1) The document discusses how it was previously thought that Alzheimer's disease (AD) development in elderly women was due to decreasing estrogen levels after menopause. However, recent evidence suggests it may instead be due to a progressive increase in luteinizing hormone (LH) levels.
2) LH has been shown to promote the amyloidogenic pathway which leads to amyloid beta plaque formation in the hippocampus, a brain region involved in AD. Blocking LH levels has shown therapeutic benefits.
3) The document provides background on AD symptoms and pathology, such as amyloid plaques and neurofibrillary tangles. It also discusses the shift in understanding of LH's actions in the brain through its receptors, such as
El documento describe la Web 2.0, incluyendo sus características principales como ser más interactiva y dinámica, permitiendo una mayor participación de los usuarios a través de herramientas como blogs, wikis y redes sociales. También discute los tipos de aplicaciones Web 2.0 como redes sociales, generación de contenido y herramientas, y concluye resumiendo 10 cosas importantes que debemos saber sobre la Web 2.0 como ser una plataforma social, flexible y participativa.
SHREE SIDDHI SOFT SOLUTIONS is one of the best ieee project centers in Trichy, SHREE SIDDHI SOFT SOLUTIONS offers IEEE 2016 Final Year projects for Engineering students in JAVA, Dot Net, Android, Oracle,matlab,embedded system and PHP technologies, SHREE SIDDHI SOFT SOLUTIONS is a one of the leading research and Development in TRICHY
Contact US:
SHREE SIDDHI SOFT SOLUTIONS
#3, 4th FLOOR, LAKSHMI ARCADE,
11TH CROSS, THILLAI NAGAR,
TRICHY-620018.
Phone NO : +91 90032-44446,900323336,0431-4040672
Email : shreesiddhisoftsolutionstrichy@gmail.com
Website : www.shreesiddhisoftsolutions.com
www.siddhisoftsolutions.com
SHREE SIDDHI SOFT SOLUTIONS is one of the best ieee project centers in Trichy, SHREE SIDDHI SOFT SOLUTIONS offers IEEE 2016 Final Year projects for Engineering students in JAVA, Dot Net, Android, Oracle,matlab,embedded system and PHP technologies, SHREE SIDDHI SOFT SOLUTIONS is a one of the leading research and Development in TRICHY
Contact US:
SHREE SIDDHI SOFT SOLUTIONS
#3, 4th FLOOR, LAKSHMI ARCADE,
11TH CROSS, THILLAI NAGAR,
TRICHY-620018.
Phone NO : +91 90032-44446,900323336,0431-4040672
Email : shreesiddhisoftsolutionstrichy@gmail.com
Website :
www.shreesiddhisoftsolutions.com
www.siddhisoftsolutions.com
El documento proporciona información sobre el uso histórico de drogas en varias culturas y civilizaciones antiguas. Resalta que las drogas se usaban con fines medicinales, religiosos y recreativos en Mesopotamia, Egipto, Grecia y Roma. También examina el uso de drogas en la prehistoria europea y en las culturas precolombinas de América. Por último, analiza brevemente algunos usos médicos y no médicos de drogas a través de la historia.
Teresa Clotilde Ojeda Sánchez: En el marco del proceso de preparación, respuesta y rehabilitación de la gestión del riesgo de desastres del Sistema Educativo Peruano le presentamos el Módulo intermedio de capacitación para la brigada de seguridad frente al riesgo social ante situaciones de emergencia y desastres. Este documento de formación destaca la importancia de la Brigada de seguridad frente a riesgo social ante situaciones de emergencia y/o desastres, como un proceso utilizado poner a buen recaudo la salud y vida de estudiantes, docentes, personal administrativo u otros miembros de la comunidad educativa.
La tabla muestra tres grupos de palabras clave relacionadas con la historia de Roma antigua - grupo A: destronar, consulado, imperio. Grupo B: trono, república, dinastía. Grupo C: monarquía, cónsul, césar - que los estudiantes usarán para crear ideografías combinando cada palabra con una imagen que represente su significado.
Teresa Clotilde Ojeda Sánchez: El Sistema Nacional de Evaluación, Acreditación y Certificación de la Calidad de la Educación (SINEACE) ha publicado la Matriz de evaluación de las Instituciones Educativas de Educación Básica Regular Es el documento que presenta factores, estándares e indicadores de calidad de la gestión educativa construidos participativamente, que orientan a todas las instituciones educativas del país: públicas y privadas, rurales y urbanas; para que puedan identificar cuan cerca o lejos se encuentran de realizar una gestión que favorece la formación integral de todos los estudiantes y a partir de ello, implementar las mejoras que sean necesarias.
La Matriz de evaluación permite identificar las fortalezas en la gestión de la IE; es decir, aquellos aspectos que se están trabajando de manera adecuada. Esto se aprecia cuando se alcanza el nivel "Logrado" de los estándares.
Asimismo, ayudará a reconocer los aspectos por mejorar en la gestión de la IE, así determinar aquellos aspectos que están dejándose de lado, que no se están pudiendo controlar o que no están siendo trabajados suficientemente y que por lo tanto ponen en riesgo el aprendizaje de los estudiantes. Esto se evidencia cuando los estándares se encuentran en el nivel "En Inicio", "Poco avance" o "Avance significativo", según sea el caso
This document provides a summary of Yashu.hul@hotmail.com's professional experience and qualifications. It summarizes his career as a senior management professional with over 9 years of experience in sales, business development, and channel management roles. Currently, he works as a Senior Manager at ANI Technologies Pvt Ltd in Bangalore, focusing on business development and channel management. He has expertise in strategic planning, sales strategies, key account management, and client relationship management.
A modern human pattern of dental development in lower pleistocene hominids fr...Kristian Pedersen
This document summarizes a study of dental development patterns in hominid fossils from the Gran Dolina site in Spain. The fossils are dated to around 0.8 million years ago. Three individuals provided information about their dental development:
1) Hominid 1 showed signs of stress during childhood in dental enamel and dentine.
2) Hominid 2 and 3 died during early childhood before their third molars erupted.
3) Analysis of tooth mineralization stages in the fossils showed more similarity to patterns in modern humans than chimpanzees. This supports the view that by 0.8 million years ago, Homo species had a prolonged developmental pattern like humans.
Evolution & development of periodontium 22.08.14drcherry24
This document discusses the evolution and development of the periodontium. It begins by defining evolution and discussing Darwin's theory of evolution by natural selection and descent with modification. It then discusses the evolution of different parts of the periodontium, including how cementum evolved from tooth ankylosis in early humans to ligamentous suspension in mammals to allow tooth movement. It also discusses the evolution of the alveolar bone, periodontal ligament, and gingiva. The document then shifts to discussing the development of the periodontium, defining development and discussing the development of cementum, alveolar bone, periodontal ligament, and gingiva from embryonic origins. It provides details on the cellular processes involved in
Home Read Sign inEXPLORATIONSCONTENTS Search in boSusanaFurman449
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EXPLORATIONS
CONTENTS
Search in book …
Private: Main Body
9. Early Hominins
Kerryn Warren, Ph.D., University of Cape Town
Lindsay Hunter, Ph.D., University of Witwatersrand
Navashni Naidoo, M.Sc., University of Cape Town
Silindokuhle Mavuso, M.Sc., University of Witwatersrand
Kimberleigh Tommy, M.Sc., University of Witwatersrand
Rosa Moll, M.Sc., University of Witwatersrand
Nomawethu Hlazo, M.Sc., University of Cape Town
Learning Objectives
De�ne what is meant by “hominin”.
Understand what is meant by “derived” and “primitive” traits and why this is relevant for understanding early
hominin evolution.
Understand changing paleoclimates and paleoenvironments during early human evolution, and contextualize
them as potential factors in�uencing adaptations during this time.
Describe the anatomical changes associated with bipedalism in early hominins and the implications for
changes in locomotion.
Describe the anatomical changes associated with dentition in early hominins and their implication for diet in
the Plio-Pleistocene.
Describe early hominin genera and species, including their currently understood dates and geographic ex-
panses and what we know about them. Previous: Primate Evolution
Next: Early Members of the Genus Homo
https://pressbooks-dev.oer.hawaii.edu/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/front-matter/__unknown__/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/wp-login.php?redirect_to=https%3A%2F%2Fpressbooks-dev.oer.hawaii.edu%2Fexplorationsbioanth%2Fchapter%2Fchapter-9-early-hominins-2%2F
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/chapter/__unknown__-10/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/chapter/__unknown__-15/
Describe the earliest stone tool techno-complex and what it implies about the transition from early ho-
minins to our genus.
DEFINING HOMININS
It is through our study of our hominin ancestors and relatives that we are exposed to a world of “might have beens”: of
other paths not taken by our species, other ways of being human. But in order to better understand these different evolu-
tionary trajectories, we must �rst de�ne the terms we are using. If an imaginary line were drawn between ourselves and
our closest relatives, the great apes, bipedalism (or habitually walking upright on two feet) is where that line would be.
Hominin, then, means everyone on “our” side of the line: humans and all of our extinct bipedal ancestors and relatives
since our divergence from the last common ancestor (LCA) with chimpanzees.
Historic interpretations of our evolution, prior to our �nding of early hominin fossils, varied. Debates in the mid-1800s re-
garding hominin origins focused on two key issues:
���Where did we evolve?
���Which traits evolved �rst?
Charles Darwin hypothesized that we evolved ...
MSCIII_Forensic anthropology_age estimation using dentition.pptxSuchita Rawat
This document discusses dental anatomy and forensic dentistry. It begins by describing the basic anatomy of teeth, including the crown, gumline, neck, root, dentin, pulp, cementum, periodontal ligament, and alveolar bone. It then compares the features of primary and permanent teeth. The document outlines methods for estimating age based on teeth, including examining the development and eruption times of primary and secondary teeth. It discusses estimating age in the prenatal, neonatal, and postnatal periods based on tooth weight and formation. Methods for estimating age in children, adolescents, and adults are described, including the Atlas, Schour-Massler, Demirjian, and Gustafson techniques. Amino acid race
Anthropology ortho /certified fixed orthodontic courses by Indian dental acad...Indian dental academy
Welcome to Indian Dental Academy
The Indian Dental Academy is the Leader in continuing dental education , training dentists in all aspects of dentistry and offering a wide range of dental certified courses in different formats.
Indian dental academy has a unique training program & curriculum that provides students with exceptional clinical skills and enabling them to return to their office with high level confidence and start treating patients
State of the art comprehensive training-Faculty of world wide repute &Very affordable.
Human Evolution of the Teeth & Jaws: A Mouthful of History- International Jou...hindol1996
The morphology & function of the teeth & jaws of Homo sapiens has changed with hominid evolution to their present form, when compared to those who lived 25,000 years ago, of the last common ancestor of humans & living apes. The field of human evolution is not normally considered within the scope of dentistry, yet the very same jaws & teeth upon which dentistry depends provide much of the essential evidence of human evolution. This review article tells a story of the patterns of dental morphological evolution in the course of time in relation to human origins. It provides a compelling account of how the interaction of diet, speech & environment has shaped human evolution of the jaws & teeth.
Forensic dentistry is the complete evaluation and examination of dental evidence to aid in administration of criminal as well as civil justice.
Medicolegal significance of teeth, bite marks, dental development ( from neonates to adults and changes at old age )
Used for identification, racial significance in mass disasters, mutilated bodies, DNA analysis, toxicological analysis in cases of poisoning.
Age estimation in civil cases, solving discrepancy of age for athletes playing sports for national and international academy.
various others importance too.
The middle ear bones in mammals are derived from bones in the dentar.pdfaquacareser
The middle ear bones in mammals are derived from bones in the dentary bone (mandible) of
Therapsids. In the Therapsid dentary, there were three bones used in the jaw hinge: the angular,
articular and quadrate. The tympanic, malleus, and incus are homologous to the angular,
articular, and quadrate
Tympanic = Angular
Malleus = Articular
Incus = Quadrate
In early Therapsids, the quadrate, angular and articular were directly connected to the dentary
bone in the jaw hinge. These bones were essential for jaw movement. These bones were also
utilized for crude hearing capabilities. The next stage of evolution came with the separation of
these bones and the dentary. After the bones separated from the jaw, they formed the postdentary
rod. It is believed that these the mammalian middle ear bones evolved at least twice. It is
believed that monotreme and therian ancestors evolved very similar middle ear bones, but in two
separate paths. In monotreme ancestors, there is a mandibular trough, that is believed to have
accommodated the postdentary rod. This rod is believed to have been connected to the trough by
ligaments. This rod was used for hearing and the quadrate, angular, and articular were no longer
needed in the jaw hinge. In therians, the mandibular trough no longer held the quadrate, angular,
and articular. These bones were held to the jaw by ossified Meckel’s cartilage. This bridge of
cartilage and bone extended away from the dentary and approached the area that would make up
the modern ear cavity.
The evolution of the mammalian middle ear and jaw joint were pivotal steps in the evolution of
mammals. It is also a great example of how classical comparative anatomy, paleontology and
developmental biology have come together to piece together how this remarkable transformation
of jaw joint to ear ossicles was able to come about. The homologies of the malleus, incus and
stapes to the articular, quadrate and columella, and tympanic ring and gonial to the angular and
prearticular suggested by comparative anatomy 175 years ago have been recently confirmed by
molecular and developmental biology. The recent discovery of new mammali form fossils has
allowed careful documentation of the shift from primary to secondary jaw articulation, creating
an opportunity to follow the transformation of the post-dentary skeletal elements. This fossil data
has been complemented by the study of marsupial development, providing insight into the
changing role of the malleus and incus, and the relationship of the primary and secondary jaw
joints. We now have a number of unanswered questions. What are the signalling molecules
involved in the interactions between the condylar process and the glenoid fossa that create the
novel mammalian jaw joint? What are the mechanisms that lead to transformation of Meckel’s
cartilage into a ligament allowing isolation of the ear from the jaw? What controls the
distribution of secondary cartilages? What controls the timing of differentiation an.
Growth pattern of mandible /certified fixed orthodontic courses by Indian d...Indian dental academy
The Indian Dental Academy is the Leader in continuing dental education , training dentists in all aspects of dentistry and offering a wide range of dental certified courses in different formats.
Indian dental academy provides dental crown & Bridge,rotary endodontics,fixed orthodontics,
Dental implants courses.for details pls visit www.indiandentalacademy.com ,or call
00919248678078
This document discusses various methods for assessing the growth pattern of bones, with a focus on the mandible. It states that serial cephalometric radiography combined with radiopaque implants is currently the most accurate method for determining mandibular growth patterns, as it allows superimposition of serial images based on fixed implant markers. However, digital subtraction radiography, computed tomography, and magnetic resonance imaging may provide even more detailed information. The document then reviews several methods that have been used historically and in other studies to examine mandibular growth, including anthropometry, vital staining, histology, radiography, implants and impressions.
Teeth of Royalty from a burial in Jera Lompo’E, South Sulawesi, IndonesiaUniversitasGadjahMada
Teeth are one of the key to understand disease and cultural influence in the past. This research describes abnormalities and cultural influences found in teeth belongs to Addatuang Sidenreng which is a royalty buried in Jera Lompo'E, South Sulawesi, Indonesia (17th century). The material consists of 13 teeth which currently curated in Laboratory of Bioanthropology and Paleoanthropology, UGM, Indonesia. This research uses bioarchaeological method and macroscopical analysis. The results are the individual age estimation, health status and cultural influences, such as betel chewing and dental filing which still practiced even after Islam enters Sulawesi, Indonesia.
1. This study examines the tooth roots of the Sima del Elefante mandible (ATE9-1) using microtomography.
2. Microtomography allows visualization of internal dental structures like root canals, revealing untapped biological information.
3. The study reclassifies two teeth, the lower left canine and lower left first premolar, based on analysis of their internal root anatomy, which does not always correspond to external anatomy.
The document describes the skull of Homo naledi, a recently discovered hominin species from South Africa. It provides detailed descriptions and measurements of cranial and mandibular fossils attributed to H. naledi individuals. Six individuals (DH1-DH6) are recognized based on skull fragments. The cranial features of H. naledi are described and compared quantitatively and qualitatively to other hominin species. While the cranial capacity is small, within the range of australopiths and early Homo, some cranial characters of H. naledi are shared with species across the genus Homo. However, the skull is readily distinguishable from existing Homo species. The evolutionary
The document discusses the evolution of the human masticatory system, including jaws and temporomandibular joint (TMJ). It describes how Darwin and Wallace developed the theory of natural selection to explain evolution. The size of human jaws and maxilla have decreased compared to great apes due to dietary changes. The protruding chin is an evolutionary feature that separates humans from ancestors. Early humans experienced a reduction in jaw size that has persisted in modern humans. Diet influences the size and shape of jaws, maxilla, and dental arches.
Essay on The History of Human Evolution
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Human Evolution
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Human Evolution Essay
Homo Sapien Human Evolution
The Importance Of Human Evolution
The Future Of Human Evolution Essay
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Did Human Evolution: How Did Humans Come To Be?
Evolution Of Human Evolution
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Evolution And Its Impact On Human Evolution
El documento proporciona información sobre el uso histórico de drogas en varias culturas y civilizaciones antiguas. Resalta que las drogas se usaban con fines medicinales, religiosos y recreativos en Mesopotamia, Egipto, Grecia y Roma. También examina el uso de drogas en la prehistoria europea y en las culturas precolombinas de América. Por último, analiza brevemente algunos usos médicos y no médicos de drogas a través de la historia.
Teresa Clotilde Ojeda Sánchez: En el marco del proceso de preparación, respuesta y rehabilitación de la gestión del riesgo de desastres del Sistema Educativo Peruano le presentamos el Módulo intermedio de capacitación para la brigada de seguridad frente al riesgo social ante situaciones de emergencia y desastres. Este documento de formación destaca la importancia de la Brigada de seguridad frente a riesgo social ante situaciones de emergencia y/o desastres, como un proceso utilizado poner a buen recaudo la salud y vida de estudiantes, docentes, personal administrativo u otros miembros de la comunidad educativa.
La tabla muestra tres grupos de palabras clave relacionadas con la historia de Roma antigua - grupo A: destronar, consulado, imperio. Grupo B: trono, república, dinastía. Grupo C: monarquía, cónsul, césar - que los estudiantes usarán para crear ideografías combinando cada palabra con una imagen que represente su significado.
Teresa Clotilde Ojeda Sánchez: El Sistema Nacional de Evaluación, Acreditación y Certificación de la Calidad de la Educación (SINEACE) ha publicado la Matriz de evaluación de las Instituciones Educativas de Educación Básica Regular Es el documento que presenta factores, estándares e indicadores de calidad de la gestión educativa construidos participativamente, que orientan a todas las instituciones educativas del país: públicas y privadas, rurales y urbanas; para que puedan identificar cuan cerca o lejos se encuentran de realizar una gestión que favorece la formación integral de todos los estudiantes y a partir de ello, implementar las mejoras que sean necesarias.
La Matriz de evaluación permite identificar las fortalezas en la gestión de la IE; es decir, aquellos aspectos que se están trabajando de manera adecuada. Esto se aprecia cuando se alcanza el nivel "Logrado" de los estándares.
Asimismo, ayudará a reconocer los aspectos por mejorar en la gestión de la IE, así determinar aquellos aspectos que están dejándose de lado, que no se están pudiendo controlar o que no están siendo trabajados suficientemente y que por lo tanto ponen en riesgo el aprendizaje de los estudiantes. Esto se evidencia cuando los estándares se encuentran en el nivel "En Inicio", "Poco avance" o "Avance significativo", según sea el caso
This document provides a summary of Yashu.hul@hotmail.com's professional experience and qualifications. It summarizes his career as a senior management professional with over 9 years of experience in sales, business development, and channel management roles. Currently, he works as a Senior Manager at ANI Technologies Pvt Ltd in Bangalore, focusing on business development and channel management. He has expertise in strategic planning, sales strategies, key account management, and client relationship management.
A modern human pattern of dental development in lower pleistocene hominids fr...Kristian Pedersen
This document summarizes a study of dental development patterns in hominid fossils from the Gran Dolina site in Spain. The fossils are dated to around 0.8 million years ago. Three individuals provided information about their dental development:
1) Hominid 1 showed signs of stress during childhood in dental enamel and dentine.
2) Hominid 2 and 3 died during early childhood before their third molars erupted.
3) Analysis of tooth mineralization stages in the fossils showed more similarity to patterns in modern humans than chimpanzees. This supports the view that by 0.8 million years ago, Homo species had a prolonged developmental pattern like humans.
Evolution & development of periodontium 22.08.14drcherry24
This document discusses the evolution and development of the periodontium. It begins by defining evolution and discussing Darwin's theory of evolution by natural selection and descent with modification. It then discusses the evolution of different parts of the periodontium, including how cementum evolved from tooth ankylosis in early humans to ligamentous suspension in mammals to allow tooth movement. It also discusses the evolution of the alveolar bone, periodontal ligament, and gingiva. The document then shifts to discussing the development of the periodontium, defining development and discussing the development of cementum, alveolar bone, periodontal ligament, and gingiva from embryonic origins. It provides details on the cellular processes involved in
Home Read Sign inEXPLORATIONSCONTENTS Search in boSusanaFurman449
Home Read Sign in
EXPLORATIONS
CONTENTS
Search in book …
Private: Main Body
9. Early Hominins
Kerryn Warren, Ph.D., University of Cape Town
Lindsay Hunter, Ph.D., University of Witwatersrand
Navashni Naidoo, M.Sc., University of Cape Town
Silindokuhle Mavuso, M.Sc., University of Witwatersrand
Kimberleigh Tommy, M.Sc., University of Witwatersrand
Rosa Moll, M.Sc., University of Witwatersrand
Nomawethu Hlazo, M.Sc., University of Cape Town
Learning Objectives
De�ne what is meant by “hominin”.
Understand what is meant by “derived” and “primitive” traits and why this is relevant for understanding early
hominin evolution.
Understand changing paleoclimates and paleoenvironments during early human evolution, and contextualize
them as potential factors in�uencing adaptations during this time.
Describe the anatomical changes associated with bipedalism in early hominins and the implications for
changes in locomotion.
Describe the anatomical changes associated with dentition in early hominins and their implication for diet in
the Plio-Pleistocene.
Describe early hominin genera and species, including their currently understood dates and geographic ex-
panses and what we know about them. Previous: Primate Evolution
Next: Early Members of the Genus Homo
https://pressbooks-dev.oer.hawaii.edu/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/front-matter/__unknown__/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/wp-login.php?redirect_to=https%3A%2F%2Fpressbooks-dev.oer.hawaii.edu%2Fexplorationsbioanth%2Fchapter%2Fchapter-9-early-hominins-2%2F
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/chapter/__unknown__-10/
https://pressbooks-dev.oer.hawaii.edu/explorationsbioanth/chapter/__unknown__-15/
Describe the earliest stone tool techno-complex and what it implies about the transition from early ho-
minins to our genus.
DEFINING HOMININS
It is through our study of our hominin ancestors and relatives that we are exposed to a world of “might have beens”: of
other paths not taken by our species, other ways of being human. But in order to better understand these different evolu-
tionary trajectories, we must �rst de�ne the terms we are using. If an imaginary line were drawn between ourselves and
our closest relatives, the great apes, bipedalism (or habitually walking upright on two feet) is where that line would be.
Hominin, then, means everyone on “our” side of the line: humans and all of our extinct bipedal ancestors and relatives
since our divergence from the last common ancestor (LCA) with chimpanzees.
Historic interpretations of our evolution, prior to our �nding of early hominin fossils, varied. Debates in the mid-1800s re-
garding hominin origins focused on two key issues:
���Where did we evolve?
���Which traits evolved �rst?
Charles Darwin hypothesized that we evolved ...
MSCIII_Forensic anthropology_age estimation using dentition.pptxSuchita Rawat
This document discusses dental anatomy and forensic dentistry. It begins by describing the basic anatomy of teeth, including the crown, gumline, neck, root, dentin, pulp, cementum, periodontal ligament, and alveolar bone. It then compares the features of primary and permanent teeth. The document outlines methods for estimating age based on teeth, including examining the development and eruption times of primary and secondary teeth. It discusses estimating age in the prenatal, neonatal, and postnatal periods based on tooth weight and formation. Methods for estimating age in children, adolescents, and adults are described, including the Atlas, Schour-Massler, Demirjian, and Gustafson techniques. Amino acid race
Anthropology ortho /certified fixed orthodontic courses by Indian dental acad...Indian dental academy
Welcome to Indian Dental Academy
The Indian Dental Academy is the Leader in continuing dental education , training dentists in all aspects of dentistry and offering a wide range of dental certified courses in different formats.
Indian dental academy has a unique training program & curriculum that provides students with exceptional clinical skills and enabling them to return to their office with high level confidence and start treating patients
State of the art comprehensive training-Faculty of world wide repute &Very affordable.
Human Evolution of the Teeth & Jaws: A Mouthful of History- International Jou...hindol1996
The morphology & function of the teeth & jaws of Homo sapiens has changed with hominid evolution to their present form, when compared to those who lived 25,000 years ago, of the last common ancestor of humans & living apes. The field of human evolution is not normally considered within the scope of dentistry, yet the very same jaws & teeth upon which dentistry depends provide much of the essential evidence of human evolution. This review article tells a story of the patterns of dental morphological evolution in the course of time in relation to human origins. It provides a compelling account of how the interaction of diet, speech & environment has shaped human evolution of the jaws & teeth.
Forensic dentistry is the complete evaluation and examination of dental evidence to aid in administration of criminal as well as civil justice.
Medicolegal significance of teeth, bite marks, dental development ( from neonates to adults and changes at old age )
Used for identification, racial significance in mass disasters, mutilated bodies, DNA analysis, toxicological analysis in cases of poisoning.
Age estimation in civil cases, solving discrepancy of age for athletes playing sports for national and international academy.
various others importance too.
The middle ear bones in mammals are derived from bones in the dentar.pdfaquacareser
The middle ear bones in mammals are derived from bones in the dentary bone (mandible) of
Therapsids. In the Therapsid dentary, there were three bones used in the jaw hinge: the angular,
articular and quadrate. The tympanic, malleus, and incus are homologous to the angular,
articular, and quadrate
Tympanic = Angular
Malleus = Articular
Incus = Quadrate
In early Therapsids, the quadrate, angular and articular were directly connected to the dentary
bone in the jaw hinge. These bones were essential for jaw movement. These bones were also
utilized for crude hearing capabilities. The next stage of evolution came with the separation of
these bones and the dentary. After the bones separated from the jaw, they formed the postdentary
rod. It is believed that these the mammalian middle ear bones evolved at least twice. It is
believed that monotreme and therian ancestors evolved very similar middle ear bones, but in two
separate paths. In monotreme ancestors, there is a mandibular trough, that is believed to have
accommodated the postdentary rod. This rod is believed to have been connected to the trough by
ligaments. This rod was used for hearing and the quadrate, angular, and articular were no longer
needed in the jaw hinge. In therians, the mandibular trough no longer held the quadrate, angular,
and articular. These bones were held to the jaw by ossified Meckel’s cartilage. This bridge of
cartilage and bone extended away from the dentary and approached the area that would make up
the modern ear cavity.
The evolution of the mammalian middle ear and jaw joint were pivotal steps in the evolution of
mammals. It is also a great example of how classical comparative anatomy, paleontology and
developmental biology have come together to piece together how this remarkable transformation
of jaw joint to ear ossicles was able to come about. The homologies of the malleus, incus and
stapes to the articular, quadrate and columella, and tympanic ring and gonial to the angular and
prearticular suggested by comparative anatomy 175 years ago have been recently confirmed by
molecular and developmental biology. The recent discovery of new mammali form fossils has
allowed careful documentation of the shift from primary to secondary jaw articulation, creating
an opportunity to follow the transformation of the post-dentary skeletal elements. This fossil data
has been complemented by the study of marsupial development, providing insight into the
changing role of the malleus and incus, and the relationship of the primary and secondary jaw
joints. We now have a number of unanswered questions. What are the signalling molecules
involved in the interactions between the condylar process and the glenoid fossa that create the
novel mammalian jaw joint? What are the mechanisms that lead to transformation of Meckel’s
cartilage into a ligament allowing isolation of the ear from the jaw? What controls the
distribution of secondary cartilages? What controls the timing of differentiation an.
Growth pattern of mandible /certified fixed orthodontic courses by Indian d...Indian dental academy
The Indian Dental Academy is the Leader in continuing dental education , training dentists in all aspects of dentistry and offering a wide range of dental certified courses in different formats.
Indian dental academy provides dental crown & Bridge,rotary endodontics,fixed orthodontics,
Dental implants courses.for details pls visit www.indiandentalacademy.com ,or call
00919248678078
This document discusses various methods for assessing the growth pattern of bones, with a focus on the mandible. It states that serial cephalometric radiography combined with radiopaque implants is currently the most accurate method for determining mandibular growth patterns, as it allows superimposition of serial images based on fixed implant markers. However, digital subtraction radiography, computed tomography, and magnetic resonance imaging may provide even more detailed information. The document then reviews several methods that have been used historically and in other studies to examine mandibular growth, including anthropometry, vital staining, histology, radiography, implants and impressions.
Teeth of Royalty from a burial in Jera Lompo’E, South Sulawesi, IndonesiaUniversitasGadjahMada
Teeth are one of the key to understand disease and cultural influence in the past. This research describes abnormalities and cultural influences found in teeth belongs to Addatuang Sidenreng which is a royalty buried in Jera Lompo'E, South Sulawesi, Indonesia (17th century). The material consists of 13 teeth which currently curated in Laboratory of Bioanthropology and Paleoanthropology, UGM, Indonesia. This research uses bioarchaeological method and macroscopical analysis. The results are the individual age estimation, health status and cultural influences, such as betel chewing and dental filing which still practiced even after Islam enters Sulawesi, Indonesia.
1. This study examines the tooth roots of the Sima del Elefante mandible (ATE9-1) using microtomography.
2. Microtomography allows visualization of internal dental structures like root canals, revealing untapped biological information.
3. The study reclassifies two teeth, the lower left canine and lower left first premolar, based on analysis of their internal root anatomy, which does not always correspond to external anatomy.
The document describes the skull of Homo naledi, a recently discovered hominin species from South Africa. It provides detailed descriptions and measurements of cranial and mandibular fossils attributed to H. naledi individuals. Six individuals (DH1-DH6) are recognized based on skull fragments. The cranial features of H. naledi are described and compared quantitatively and qualitatively to other hominin species. While the cranial capacity is small, within the range of australopiths and early Homo, some cranial characters of H. naledi are shared with species across the genus Homo. However, the skull is readily distinguishable from existing Homo species. The evolutionary
The document discusses the evolution of the human masticatory system, including jaws and temporomandibular joint (TMJ). It describes how Darwin and Wallace developed the theory of natural selection to explain evolution. The size of human jaws and maxilla have decreased compared to great apes due to dietary changes. The protruding chin is an evolutionary feature that separates humans from ancestors. Early humans experienced a reduction in jaw size that has persisted in modern humans. Diet influences the size and shape of jaws, maxilla, and dental arches.
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Evolution And Its Impact On Human Evolution
Anatomical proof of evolution and fossilsDiegoRmej
This document discusses how scientists have supported the theory of evolution through anatomical evidence and fossil discoveries. It explains that early scientists showed anatomical similarities between organisms to demonstrate evolution. Additionally, fossil discoveries like shark teeth and "tongue stones" provided early evidence that fossils record past life. The document also gives examples of how fossils can provide clues about species interactions and growth patterns to increase understanding of evolutionary history. Finally, it describes how analogous, homologous, and vestigial structures in living things also support evolution by demonstrating common ancestry and evolutionary changes over time.
The document analyzes three specimens of Paranthropus robustus fossils from the Pleistocene period in South Africa that had not been previously examined. It compares the enamel characteristics that have evolved over time in southern Africa during the early Pleistocene. The study uses dental micro wear texture analysis to examine how the enamel of Paranthropus robustus had evolved by looking at extant primate taxa. The main purpose was to understand how and why the enamel, molars, and premolars have changed over time.
"Certainly going back to Sherlock Holmes we have a tradition of forensic science featured in detective stories.”- Jeffrey Deaver.
With time, forensic dentistry have come along, shaking hands with forensic science and gradually taking an important position in the field of forensic science.
Forensic odontology is the branch of dentistry which, in the interest of justice, deals with the proper handling and examination of dental evidence, and with the proper evaluation and presentation of dental findings. Forensic odontologists delve into: identifying unknown human remains, victim’s identification in mass disaster, electing the picture of life style and diet of skeletal remains at forensic and archaeological sites, assessing sex of skeletal remains, age estimation of both living and deceased, analysis and identification of bite marks at crime scenes.
It is the branch of science that applies dental knowledge in civil and criminal investigations. Along with other healthcare providers, dentists encounter cases of injuries which could be non-accidental. Detection, interpretation and management are important from a legal and humanitarian point of view. Dentists should be aware of the legal impact those cases have, and should refer them to the appropriate authorities for suitable action.
Today we consider forensic odontology to be a specialised and reliable method of identification of the deceased, particularly in multiple fatality incidents. Forensic Odontology has established itself as an important, often indispensable, in medicolegal cases, in particular for identification of the dead.
MUSEUM OF NATURAL HISTORY PUBLICATION FOR EDUCATORSV.docxroushhsiu
1) The document discusses human evolution from a behavioral perspective, focusing on when key human behaviors like symbolic thinking, abstract thinking, and theory of mind first emerged based on the fossil and archaeological records.
2) Early humans between 2.6-2.3 mya began using stone tools and scavenging meat, suggesting problem-solving skills, but little evidence of abstract thinking, symbolic thinking, or imagined communities.
3) By around 1 mya, control of fire, new technologies, and nighttime social activities provided evidence of more advanced behaviors, though innovation remained slow.
Similar to Anterior tooth growth periods in neandertals were comparable to those of modern human (guatelli steinberg et al.) (20)
The document discusses the religious beliefs and practices in Scandinavia from 600-900 AD. It describes the principal deities belonging to the Æsir and Vanir groups, which were sometimes in conflict but later reconciled. The Æsir such as Odin and Thor were more popular in western Scandinavia, while the Vanir like Njord and Freyr were predominant in the east. Lesser spiritual entities also played roles in rituals and beliefs. The text examines the distribution of cults across regions and outlines some of the major deities and myths from Old Norse sources.
The document discusses the social structure and political institutions of Scandinavian communities during the Viking Age from 700-900 AD. It analyzes the social hierarchy and divisions based on sex, age, and status using archaeological evidence, textual sources, mythical tales, and literary accounts. Women generally had high social status and equal rights to men. The sources describe distinct social classes of slaves, freemen, and rulers. Archaeological evidence like the rich Oseberg ship burial shows some women held elite aristocratic roles. Overall, the document aims to understand Viking society by examining evidence from multiple genres of sources.
The document discusses the origins of Viking culture in Scandinavia during the Bronze Age and Iron Age. It describes how aspects of culture developed over this period, including ship building, domestic architecture like longhouses, religious practices, and economic activities like agriculture, fishing, and trade. Society was stratified with evidence of wealthier individuals. Overall, the foundations for the expansion of the Viking Age were established through cultural developments spanning many centuries prior.
This document provides an overview of a course on the Vikings between AD 700-1100. It discusses the sources used to study the Vikings, including historical accounts, sagas, runic inscriptions and place names. It also outlines advances in archaeological techniques that have improved understanding of Viking sites, settlements, trade and environment. Interpretive paradigms have shifted from seeing Vikings as pirates to appreciating their social structure and role in state formation, trade and assimilation into other cultures.
Dna sequence of the mitochondrial hypervariable region ii (krings et al.)Kristian Pedersen
1) The authors determined the DNA sequence of the second hypervariable region (HVRII) of mitochondrial DNA from the Neandertal type specimen.
2) When combined with the previously published HVR1 sequence, phylogenetic analysis found the Neandertal mtDNA to fall outside the variation of contemporary human mtDNA sequences.
3) The estimated date of divergence between Neandertal and modern human mtDNA sequences was 465,000 years before present, with a confidence interval of 317,000 to 741,000 years. This supports the Neandertals having a separate evolutionary history from modern humans.
Detecting ancient admixture in humans using sequence polymorphism data (wall)Kristian Pedersen
This document summarizes a study that examines how much genetic data would be needed to determine if archaic human populations like Neanderthals interbred with and contributed to the modern human gene pool. The study introduces a model of ancient population structure involving mixing between groups. It then determines how many unlinked nuclear loci from sequencing data would be required to distinguish between scenarios of single origin vs. multiregional origin of modern humans. The study finds that 50-100 loci would be needed if plausible parameter estimates are used, and that the amount of available data is not yet sufficient to support one model over the other. However, with more sequencing in coming years, the required data should become available to help resolve the debate.
Close correspondence between quantitative and molecular-genetic divergence t...Kristian Pedersen
This document presents a new method for estimating divergence times between populations based on quantitative traits like cranial measurements. The authors develop a phenotypic divergence time (PTD) estimator modeled after genetic divergence time estimators. They apply their PTD estimator to cranial measurements from over 2,500 modern humans and 20 Neandertals. Their analysis estimates the Neandertal-modern human divergence at either 311,000 years ago or 435,000 years ago, depending on assumptions. These dates are similar to estimates from ancient DNA, implying cranial and genetic divergence largely track population history through neutral evolution.
This document reanalyzes ancient mitochondrial DNA sequences recovered from Neandertal bones. Previous studies placed Neandertals at the base of the modern human phylogenetic tree, suggesting they did not contribute to the modern human gene pool. However, these analyses did not account for high substitution rate variation among sites in the human mitochondrial D-loop region or estimate nucleotide substitution model parameters. The authors reanalyze the Neandertal sequences using maximum likelihood methods that account for these factors to provide a more accurate phylogenetic reconstruction.
Ancient dna. would the real neandertal please stand up (cooper et al.)Kristian Pedersen
This document summarizes a study that analyzed mitochondrial DNA sequences recovered from Neandertal specimens and early modern human fossils. The study found:
1) Neandertal mtDNA sequences were not present in early modern humans or contemporary humans, suggesting a limited genetic contribution from Neandertals to modern humans.
2) However, the results depend on assumptions like the fossils actually containing ancient DNA, and more data is needed to definitively resolve the genetic relationship between Neandertals and modern humans.
3) Future studies using more specimens and nuclear DNA may provide more powerful tests of the extent of genetic exchange between Neandertals and early modern humans.
A re appraisal of ceprano calvaria affinities (ascenzi et al.)Kristian Pedersen
The document discusses new modifications made to the reconstruction of the Ceprano calvaria based on additional examination. It provides a reassessment of the morphology and measurements of the calvaria and evaluates its affinities to Homo erectus. New geological data from the Priverno Basin suggests an age of 800-900 thousand years for the calvaria. While showing some similarities, the calvaria exhibits significant morphological variation compared to typical H. erectus specimens.
A cranium for the earliest europeans: phylogenetic position of the hominid fr...Kristian Pedersen
The document summarizes a study analyzing the cranium of a hominid fossil discovered near Ceprano, Italy in 1994. Researchers estimated its age to be between 800-900 thousand years old based on dating of surrounding rock layers. The cranium was reconstructed and compared morphologically to other early European and Asian hominin fossils. Results found it displayed a unique blend of traits linking early Homo erectus to later specimens referred to Homo heidelbergensis. The researchers concluded the Ceprano cranium represents an important transitional form between early hominin groups, filling a gap in the human fossil record in Europe between 1-0.5 million years ago.
A calvarium of late homo erectus from ceprano, italy (ascenzi et al.)Kristian Pedersen
The document summarizes the discovery and context of a late Homo erectus calvarium found in Ceprano, Italy. Key points:
- The calvarium was discovered in 1994 during highway construction in a clay layer dated to around 700,000 years ago.
- The shape and size of the calvarium (capacity of 1185 ml) show it possesses some, but not all, features of Homo erectus.
- A geological survey found the remains in a colluvial-alluvial paleosol originating from an ancient slope, indicating the site was a marshy pool within the Ceprano basin.
The feeding biomechanics and dietary ecology of australopithecus africanus (s...Kristian Pedersen
1) The feeding biomechanics and craniofacial morphology of Australopithecus africanus were analyzed using finite element analysis informed by comparative data.
2) The results suggest that the facial skeleton of A. africanus was well-adapted to withstand loads applied during premolar biting, including stresses along the nasal margins and zygomatic root.
3) However, neither the mastication of small hard objects nor large volumes of food alone can fully explain the evolution of the robust facial features in A. africanus. Rather, key aspects are more likely related to ingesting and processing large mechanically protected foods like nuts and seeds.
The carbon isotope ecology and diet of australopithecus africanus at sterkfon...Kristian Pedersen
The document summarizes a study that analyzed the carbon isotope ratios in tooth enamel from 10 specimens of Australopithecus africanus from Sterkfontein in South Africa, dating to between 2.5-2.0 million years ago. The results show that A. africanus had a varied diet incorporating both C3 forest foods and C4 savanna foods to a greater degree than other early hominins. This suggests A. africanus was a highly adaptable and opportunistic feeder. The isotope data also indicates the individuals exhibited more dietary variation than other early hominin species, arguing against suggestions that multiple species are represented in the A. africanus taxon.
Ontogeny of australopithecines and early homo: evidence from carnial capacity...Kristian Pedersen
The document discusses the benefits of exercise for mental health. Regular physical activity can help reduce anxiety and depression and improve mood and cognitive function. Exercise causes chemical changes in the brain that may help protect against mental illness and improve symptoms.
Cosmogenic Nuclide Dating Of Sahelanthropus Tchadensis And Australopiethcus B...Kristian Pedersen
This study used cosmogenic nuclide dating to determine the ages of sedimentary units containing hominid remains at two localities in Chad - KT 12 locality in Koro Toro and TM 266 locality in Toros-Menalla. At KT 12, which contained Australopithecus bahrelghazali, authigenic 10Be/9Be dating of the sedimentary level yielded an age of 3.58 ± 0.27 Ma, consistent with an early Pliocene age estimation. At TM 266, where Sahelanthropus tchadensis was found, 28 10Be/9Be ages from the anthracotheriid unit containing the hominid remains bracketed its age between 6.8
The document discusses the emergence and characteristics of Neanderthals. Some key points:
- Neanderthals first emerged in Europe and the Middle East between 150,000-200,000 years ago as a distinct species from Homo heidelbergensis.
- They were restricted to Europe and Western Asia, with no evidence found in Africa or East Asia.
- Anatomically, Neanderthals had stockier builds than modern humans and more robust skeletons, especially in the cranium.
- Genetic evidence shows Neanderthals diverged from the modern human line well before 300,000 years ago, with a substantial genetic distance between the two groups.
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Homo erectus expanded out of Africa into Eurasia around 1.6 million years ago. Some of the earliest sites outside of Africa are found in East Asia, particularly China and Java, dating back to around 1.9-1.7 million years ago. Homo erectus inhabited sites across Asia utilizing chopper tool industries rather than hand-axes. Some of the most important early Asian sites include Longgupo Cave in China dating to 1.7-1.9 million years ago, and Sangiran in Java dating to around 1.6 million years ago. Homo erectus expanded into Europe around 1 million years ago, with some of the earliest securely dated sites found in Spain and Italy.
Homo ergaster was an early form of Homo that expanded out of Africa between 2-1.6 million years ago. Evidence from the Dmanisi site in Georgia dates Homo ergaster occupation there to 1.88 million years ago, indicating they dispersed earlier in Africa. Homo ergaster displayed increased brain size and cranial capacity compared to Homo habilis, and were taller yet more gracile. They controlled fire, made Acheulean stone tools like handaxes, and were the first hominins to colonize areas outside of Africa, suggesting increased social complexity and technological adaptation.
This document discusses early hominin species including Australopithecus garhi, Homo habilis, and Homo rudolfensis. A. garhi, found in Ethiopia in 1996, was the earliest known tool-producing hominin dating to between 2.6-2.5 million years ago. H. habilis produced basic stone tools and expanded out of Africa between 2.4-1.5 million years ago. H. rudolfensis, found in Kenya, may be a separate species from H. habilis based on cranial differences. The document also examines the physical characteristics, fossil evidence, locomotion, diet, and basic stone tool production of these early hominins.
How to Interpret Trends in the Kalyan Rajdhani Mix Chart.pdfChart Kalyan
A Mix Chart displays historical data of numbers in a graphical or tabular form. The Kalyan Rajdhani Mix Chart specifically shows the results of a sequence of numbers over different periods.
In our second session, we shall learn all about the main features and fundamentals of UiPath Studio that enable us to use the building blocks for any automation project.
📕 Detailed agenda:
Variables and Datatypes
Workflow Layouts
Arguments
Control Flows and Loops
Conditional Statements
💻 Extra training through UiPath Academy:
Variables, Constants, and Arguments in Studio
Control Flow in Studio
This talk will cover ScyllaDB Architecture from the cluster-level view and zoom in on data distribution and internal node architecture. In the process, we will learn the secret sauce used to get ScyllaDB's high availability and superior performance. We will also touch on the upcoming changes to ScyllaDB architecture, moving to strongly consistent metadata and tablets.
Main news related to the CCS TSI 2023 (2023/1695)Jakub Marek
An English 🇬🇧 translation of a presentation to the speech I gave about the main changes brought by CCS TSI 2023 at the biggest Czech conference on Communications and signalling systems on Railways, which was held in Clarion Hotel Olomouc from 7th to 9th November 2023 (konferenceszt.cz). Attended by around 500 participants and 200 on-line followers.
The original Czech 🇨🇿 version of the presentation can be found here: https://www.slideshare.net/slideshow/hlavni-novinky-souvisejici-s-ccs-tsi-2023-2023-1695/269688092 .
The videorecording (in Czech) from the presentation is available here: https://youtu.be/WzjJWm4IyPk?si=SImb06tuXGb30BEH .
Freshworks Rethinks NoSQL for Rapid Scaling & Cost-EfficiencyScyllaDB
Freshworks creates AI-boosted business software that helps employees work more efficiently and effectively. Managing data across multiple RDBMS and NoSQL databases was already a challenge at their current scale. To prepare for 10X growth, they knew it was time to rethink their database strategy. Learn how they architected a solution that would simplify scaling while keeping costs under control.
inQuba Webinar Mastering Customer Journey Management with Dr Graham HillLizaNolte
HERE IS YOUR WEBINAR CONTENT! 'Mastering Customer Journey Management with Dr. Graham Hill'. We hope you find the webinar recording both insightful and enjoyable.
In this webinar, we explored essential aspects of Customer Journey Management and personalization. Here’s a summary of the key insights and topics discussed:
Key Takeaways:
Understanding the Customer Journey: Dr. Hill emphasized the importance of mapping and understanding the complete customer journey to identify touchpoints and opportunities for improvement.
Personalization Strategies: We discussed how to leverage data and insights to create personalized experiences that resonate with customers.
Technology Integration: Insights were shared on how inQuba’s advanced technology can streamline customer interactions and drive operational efficiency.
Northern Engraving | Nameplate Manufacturing Process - 2024Northern Engraving
Manufacturing custom quality metal nameplates and badges involves several standard operations. Processes include sheet prep, lithography, screening, coating, punch press and inspection. All decoration is completed in the flat sheet with adhesive and tooling operations following. The possibilities for creating unique durable nameplates are endless. How will you create your brand identity? We can help!
Connector Corner: Seamlessly power UiPath Apps, GenAI with prebuilt connectorsDianaGray10
Join us to learn how UiPath Apps can directly and easily interact with prebuilt connectors via Integration Service--including Salesforce, ServiceNow, Open GenAI, and more.
The best part is you can achieve this without building a custom workflow! Say goodbye to the hassle of using separate automations to call APIs. By seamlessly integrating within App Studio, you can now easily streamline your workflow, while gaining direct access to our Connector Catalog of popular applications.
We’ll discuss and demo the benefits of UiPath Apps and connectors including:
Creating a compelling user experience for any software, without the limitations of APIs.
Accelerating the app creation process, saving time and effort
Enjoying high-performance CRUD (create, read, update, delete) operations, for
seamless data management.
Speakers:
Russell Alfeche, Technology Leader, RPA at qBotic and UiPath MVP
Charlie Greenberg, host
"Scaling RAG Applications to serve millions of users", Kevin GoedeckeFwdays
How we managed to grow and scale a RAG application from zero to thousands of users in 7 months. Lessons from technical challenges around managing high load for LLMs, RAGs and Vector databases.
"Frontline Battles with DDoS: Best practices and Lessons Learned", Igor IvaniukFwdays
At this talk we will discuss DDoS protection tools and best practices, discuss network architectures and what AWS has to offer. Also, we will look into one of the largest DDoS attacks on Ukrainian infrastructure that happened in February 2022. We'll see, what techniques helped to keep the web resources available for Ukrainians and how AWS improved DDoS protection for all customers based on Ukraine experience
"$10 thousand per minute of downtime: architecture, queues, streaming and fin...Fwdays
Direct losses from downtime in 1 minute = $5-$10 thousand dollars. Reputation is priceless.
As part of the talk, we will consider the architectural strategies necessary for the development of highly loaded fintech solutions. We will focus on using queues and streaming to efficiently work and manage large amounts of data in real-time and to minimize latency.
We will focus special attention on the architectural patterns used in the design of the fintech system, microservices and event-driven architecture, which ensure scalability, fault tolerance, and consistency of the entire system.
QA or the Highway - Component Testing: Bridging the gap between frontend appl...zjhamm304
These are the slides for the presentation, "Component Testing: Bridging the gap between frontend applications" that was presented at QA or the Highway 2024 in Columbus, OH by Zachary Hamm.
LF Energy Webinar: Carbon Data Specifications: Mechanisms to Improve Data Acc...DanBrown980551
This LF Energy webinar took place June 20, 2024. It featured:
-Alex Thornton, LF Energy
-Hallie Cramer, Google
-Daniel Roesler, UtilityAPI
-Henry Richardson, WattTime
In response to the urgency and scale required to effectively address climate change, open source solutions offer significant potential for driving innovation and progress. Currently, there is a growing demand for standardization and interoperability in energy data and modeling. Open source standards and specifications within the energy sector can also alleviate challenges associated with data fragmentation, transparency, and accessibility. At the same time, it is crucial to consider privacy and security concerns throughout the development of open source platforms.
This webinar will delve into the motivations behind establishing LF Energy’s Carbon Data Specification Consortium. It will provide an overview of the draft specifications and the ongoing progress made by the respective working groups.
Three primary specifications will be discussed:
-Discovery and client registration, emphasizing transparent processes and secure and private access
-Customer data, centering around customer tariffs, bills, energy usage, and full consumption disclosure
-Power systems data, focusing on grid data, inclusive of transmission and distribution networks, generation, intergrid power flows, and market settlement data
AppSec PNW: Android and iOS Application Security with MobSFAjin Abraham
Mobile Security Framework - MobSF is a free and open source automated mobile application security testing environment designed to help security engineers, researchers, developers, and penetration testers to identify security vulnerabilities, malicious behaviours and privacy concerns in mobile applications using static and dynamic analysis. It supports all the popular mobile application binaries and source code formats built for Android and iOS devices. In addition to automated security assessment, it also offers an interactive testing environment to build and execute scenario based test/fuzz cases against the application.
This talk covers:
Using MobSF for static analysis of mobile applications.
Interactive dynamic security assessment of Android and iOS applications.
Solving Mobile app CTF challenges.
Reverse engineering and runtime analysis of Mobile malware.
How to shift left and integrate MobSF/mobsfscan SAST and DAST in your build pipeline.
Discover top-tier mobile app development services, offering innovative solutions for iOS and Android. Enhance your business with custom, user-friendly mobile applications.
What is an RPA CoE? Session 2 – CoE RolesDianaGray10
In this session, we will review the players involved in the CoE and how each role impacts opportunities.
Topics covered:
• What roles are essential?
• What place in the automation journey does each role play?
Speaker:
Chris Bolin, Senior Intelligent Automation Architect Anika Systems
Essentials of Automations: Exploring Attributes & Automation ParametersSafe Software
Building automations in FME Flow can save time, money, and help businesses scale by eliminating data silos and providing data to stakeholders in real-time. One essential component to orchestrating complex automations is the use of attributes & automation parameters (both formerly known as “keys”). In fact, it’s unlikely you’ll ever build an Automation without using these components, but what exactly are they?
Attributes & automation parameters enable the automation author to pass data values from one automation component to the next. During this webinar, our FME Flow Specialists will cover leveraging the three types of these output attributes & parameters in FME Flow: Event, Custom, and Automation. As a bonus, they’ll also be making use of the Split-Merge Block functionality.
You’ll leave this webinar with a better understanding of how to maximize the potential of automations by making use of attributes & automation parameters, with the ultimate goal of setting your enterprise integration workflows up on autopilot.
2. upon-Tyne. The southern African sample derives from several
indigenous populations, with a mixture of ethnic backgrounds.
The Alaskan sample is an archaeological one of Point Hope
Inuit, spanning six culture periods: the Near Ipiutak (500–100
B.C.), Ipiutak (100 B.C. to A.D. 500), Birnirk (A.D. 500–900),
Western Thule (A.D. 900–1300), Tigara (A.D. 1300–1700), and
Recent (A.D. 1700–present) (25).
Materials and Methods
In this study, only one tooth (right or left) was used from each
individual for each tooth type. The choice of right or left teeth
was made on the basis of which antimere was most complete.
Only teeth estimated to have 80% or more of their crown heights
intact (i.e., minimally worn teeth) were selected for analysis.
Fig. 1. Relationship between perikymata and striae of Retzius. Cross stria-
Crown heights were measured by using a reticule calibrated to a
tions appear at higher magnifications as varicosities and constrictions along
the enamel prisms. Images courtesy of Jay Kelley and Tanya Smith (35).
magnification of 50. Original crown heights were recon-
structed by following the contour of each side of the tooth cusp
and projecting it until the sides meet. Both measured and
Ferrell (22) found a modal periodicity of 8 and a mean period- reconstructed crown heights were recorded.
icity of 9 in a sample of 49 Danish canines. Continuing histo- For the Neandertal and Inuit samples, high-resolution poly-
logical work on a small sample of fossil human specimens vinyl siloxanes (Coltene’s President Jet and Struer’s RepliSet)
indicates that they also had mean and modal periodicities of 8 or were used to make dental impressions from the buccal surfaces
9 (23). D.G.-S. has observed a histological section from the of anterior teeth. These were cast in high-resolution epoxy
Tabun II Neandertal (courtesy of M. C. Dean and the Natural (Struer’s Epofix) and were coated with a gold–palladium alloy.
History Museum of London), which appears to have a period- Perikymata were counted under a light microscope, and a
icity of 8. The totality of this evidence suggests that Neandertals scanning electron microscope was used to create a micrographic
would have had mean and modal periodicities similar to those of record of tooth surfaces. Each replica was oriented orthogonally
modern and fossil humans. to the microscope’s optical axis.
However, periodicities within modern human populations are The samples from Newcastle and southern Africa are thin
highly variable, exhibiting a range of 6–12 days (present study sections on which striae of Retzius were counted under trans-
and refs. 20–22 and 24). The greatest source of variation in mitted light microscopy. Only striae clearly outcropping onto the
periodicities in two different hominin samples is therefore surface as perikymata were counted. For this reason, we refer to
expected to come from variation introduced by sampling error the striae counts in our histological sample as perikymata counts.
rather than from any fundamental difference in mean periodi- For the Newcastle and southern African samples, it was possible
cities between populations or species. Because of the wide to count cross striations to determine the periodicity for each
variation in periodicities within human populations, any analysis tooth. Hence the number of days it took to form the imbrica-
of crown formation times based solely on perikymata counts with tional enamel in each of these teeth could be calculated directly
unknown periodicities must consider the potential range of by multiplying the tooth’s periodicity by the total number of
periodicities that might exist in the sample. Unlike any previous perikymata on the tooth.
study of crown formation times in hominin fossils, our analysis Each tooth was divided into 10% increments (deciles) of the
considers this range by taking advantage of a recently discovered reconstructed crown height, and perikymata were counted
relationship between total perikymata counts and periodicity within the increments. For teeth missing up to 20% of their
(22, 24), as we describe below. crowns because of wear, estimates of perikymata were made for
the first two deciles. Perikymata counts within the first two
Samples deciles of complete crowns used in this study have very low
Table 1 lists the Neandertal teeth and sample sizes. The Nean- standard deviations, ranging from one to two perikymata for
dertal sample spans 150,000–40,000 years. The sample from each tooth type within each population. Such low variation
England derives from a single living population from Newcastle- within the first two deciles of growth makes it possible to
Table 1. Sample composition
Neandertal, Inuit, Southern African, Newcastle,
Tooth type Neandertal specimens n* n n n
UI1 Krapina 91, 93, 94, 123, 126, 155, 194, 195; Devil’s Tower 1, Le 10 10 20 19
Moustier 1
UI2 Krapina 122, 128, 130, 131, 148, 156, 160, 196; Le Moustier 1 9 10 21 16
UC Krapina Maxilla E, 37, 56, 76, 102, 103, 141, 142, 144, 146, 191; 14 9 26 39
Kulna, Le Moustier 1, La Quina 5
˚
LI1 Krapina Mandible E, 73; Ochoz, Tabun II, Le Moustier 1 5 12 20 15
LI2 Krapina Mandible C, Mandible D, Mandible E, 71, 90; Ochoz, 8 14 23 13
Tabun II, Le Moustier 1
LC Krapina Mandible D, Mandible E, 75, 119, 120, 121, 145; Ochoz, 10 10 24 13
Tabun II, Le Moustier 1
Total teeth 55 65 134 115
Total individuals 30 17 114 115
*Total number of Krapina Neandertals based on the designation of associated teeth as ‘‘Krapina Dental People’’ (34).
14198 www.pnas.org cgi doi 10.1073 pnas.0503108102 Guatelli-Steinberg et al.
3. accurately estimate growth in slightly worn teeth based on the Table 2. Descriptive statistics for periodicities: Range, mode, and
perikymata counts in the first and second deciles of complete mean for two population samples
crowns for each tooth type and each population sample. Teeth Population Range Mode Mean SD
were excluded from the study if more than one decile beyond the
first two deciles contained indistinct perikymata. For teeth in Southern African 6–12 9 9.097 1.207
which a single decile contained indistinct perikymata, counts Newcastle 7–11 9 8.748 1.042
were estimated from adjacent deciles. To eliminate interob-
server error, only counts by D.J.R. were used in the statistical
analysis. Intraobserver error for D.J.R. has been calculated at could further be argued that the Neandertals in our sample
5% for perikymata counts (13). might be forming their teeth in shorter periods than southern
We test the hypothesis that Neandertals grew their teeth in Africans, even 15% shorter, if the Neandertals in our sample had
shorter time periods than the modern human comparative still lower periodicities. Therefore, we use the southern African
samples in two ways. First, using the same method of Ramirez- regression equations for the six anterior teeth relating periodicity
Rozzi and Bermudez de Castro (15), we compare the means of
´ to perikymata counts to determine what the periodicities in our
the total perikymata counts per tooth of the Neandertal sample Neandertal samples would have to be if their imbricational
with each of the comparative samples. Under the assumption enamel formation times were equivalent to or 15% shorter than
that the mean periodicity of the Neandertal sample is equivalent those of southern Africans. We refer to these conditional
to that of the modern human comparative samples, Neandertals periodicities as ‘‘hypothetical periodicities.’’
can be inferred to be forming their imbricational enamel in The key to our analysis is that the known lower limit of periodi-
shorter periods of time only if they have significantly lower mean
cities in modern humans and African apes is 6. In fact, there are only
perikymata counts than the comparative samples. We first
2 individuals of 184 in Dean and Reid’s (20) combined African ape
analyze differences in perikymata count means across popula-
and human sample, exhibiting periodicities of 6. In the combined
tion samples for all anterior teeth combined, so that our results
Newcastle and southern African samples used in our study, there is
can be compared with those of Ramirez-Rozzi and Bermudez de ´
only one case of 249 teeth in which the periodicity was 6. Smith et
Castro (15). On the combined sample, we used a general linear
model and performed an ANOVA in which the factors were al. (21) have found no chimpanzee with periodicities of 6. Thus
tooth type, population, and the interaction of tooth type and we reject the hypothesis that Neandertals grew their teeth in the
population. We conducted an additional ANOVA using recon- same period, or in 15% less time, than southern Africans if the only
structed crown height as a covariate, because we found that, way for these hypotheses to be true is to assume that any of our
within populations, crown heights are generally positively cor- Neandertal specimens had periodicities 6.
related with the total number of perikymata on a tooth. We We obtain hypothetical periodicities for Neandertals under
performed pair-wise contrasts of total perikymata count means the assumption of equivalence to southern African crown for-
between Neandertals and each comparative sample using Dun- mation times by inserting Neandertal perikymata counts into the
nett’s simultaneous t tests for significant differences (26). Then, southern African regression equations. We determine what the
for each tooth type, we conducted one-way ANOVAs of total periodicities for the Neandertals would be if their teeth grew in
perikymata counts and Dunnett’s simultaneous t tests to deter- 15% less time than those of the southern African sample in the
mine whether there were significant differences in mean periky- following way. We first multiply the hypothetical Neandertal
mata counts between Neandertals and each comparative sample. periodicities based on southern African regression equations by
The second way we test the hypothesis of abbreviated growth the number of perikymata. This gives an approximate time in
in Neandertal teeth takes into account the unknown periodicities days for Neandertal enamel formation that is comparable to that
in our Neandertal sample. For each tooth type from a given of a southern African tooth with the same number of perikymata.
population, there is a strong negative correlation (r from 0.90 We then multiply this approximate enamel formation time by
to 0.99) between total perikymata counts on teeth and their 0.85 and divide it by the number of perikymata for each
periodicities (22, 24). Because the time it takes for imbricational Neandertal tooth to obtain a second set of hypothetical peri-
enamel to form is equal to the total number of perikymata on a odicities for Neandertals.
tooth multiplied by its periodicity, the strong negative correla-
tion between periodicity and total perikymata counts means that Results
crown formation times are fairly constant for each tooth type in Table 2 contains basic descriptive statistics on periodicity for the
a population, with only small standard deviations from the mean southern African and Newcastle samples, the two samples for
(22, 24). Regression equations of the following form can be used
to describe this relationship for each tooth type in the Newcastle
and southern African samples: Periodicity (perikymata
count). For each population (Newcastle and southern Africa),
every point on the regression line for a particular tooth type
represents a value for periodicity and total perikymata count,
which, when multiplied together, result in that tooth type’s
average crown formation time. The r2 values for these 12
regression equations (two populations, each with six tooth types)
range from 0.828 to 0.949, all with P values 0.001. They are
therefore highly predictive of the relationship between the total
number of perikymata and periodicity for a given tooth type, a
relationship that is determined by the time it takes for a
ANTHROPOLOGY
particular tooth type to form (22, 24).
As shown in Results, perikymata counts in our Neandertal
sample are higher than those of the southern African sample;
nevertheless, it could be argued that if the Neandertals in our
sample had lower periodicities than southern Africans, they Fig. 2. Plot of the interaction of tooth type by population for total periky-
might be growing their teeth in equivalent periods of time. It mata counts.
Guatelli-Steinberg et al. PNAS October 4, 2005 vol. 102 no. 40 14199
4. Table 3. ANOVA results for total perikymata counts Table 4. Dunnett’s t tests for differences in perikymata count
means across populations, all tooth types combined
Without crown height With crown
as covariate, height as covariate, Comparison,
R2 66.39% R2 70.82% Neandertal Difference
subtracted from of means t value P value
Factors df F P df F P
Inuit 15.24 4.753 0.0000
Population 3 78.74 0.000 3 90.59 0.000
Newcastle 7.94 2.706 0.0175
Tooth type 5 62.19 0.000 5 31.20 0.000
Southern African 19.50 6.945 0.0000
Population by 15 5.14 0.000 15 5.46 0.000
tooth type
Crown height 1 52.08 0.000
differences in perikymata counts between Neandertals and the
modern human samples. The Inuit have statistically significantly
higher counts than Neandertals on UI1, UC, and LC, whereas
which we could directly determine periodicity. Note that when
the two populations do not show a statistically significant
rounded to the nearest whole number, mean, and modal peri-
difference in UI2, LI1, and LI2. The Newcastle sample has
odicities for both samples are nine.
statistically significantly higher counts than Neandertals on UI1
Table 3 contains the results of two ANOVAs of perikymata
and LC but statistically significantly lower counts on LI2, and
counts in the combined sample of anterior teeth, one ANOVA
without and one with reconstructed crown height as a covariate. these two populations do not show statistically significantly
The population factor has four levels: Neandertal, Inuit, New- different counts on UI2, UC, and LI1. Last, the southern African
castle, and southern African. The tooth type factor has six levels: sample shows statistically significantly lower counts than Nean-
upper and lower first incisors, second incisors, and canines. dertals on all incisors, with statistically insignificant differences
Although the inclusion of crown height as a covariate causes the on both canines. In some cases, the comparisons at the level of
F value for the population main effect to increase and the F value tooth type are made with small sample sizes, potentially con-
for the tooth type main effect to decrease, both main effects and tributing to nonsignificant results. However, when the differ-
the interaction between them remain as significant sources of ences are significant, they confirm that Neandertal mean periky-
variation. Fig. 2 shows the graphs of the interaction plots from mata counts are lower than the mean perikymata counts of the
the first ANOVA (without adjustment for crown height). Fig. 2 Inuit, variable with respect to those of the sample from New-
shows that Neandertal mean perikymata counts are generally castle, and higher than those of southern Africans.
lower than the mean perikymata counts of the Inuit, variable Hypothetical periodicities calculated for Neandertal teeth
with respect to those of the sample from Newcastle, and gen- using regression equations relating periodicity to the total
erally higher than those of southern Africans. number of striae in the southern African sample reveal distri-
Fig. 3 gives 95% confidence intervals of the means for all butions for two tooth types (LI1 and LI2) that include periodi-
anterior teeth combined, and Table 4 contains Dunnett’s simul- cities 6, the lower limit of the known range of periodicities in
taneous t test results of mean differences in perikymata counts African apes and humans. Neandertal LI1 and LI2 must there-
between Neandertals and each comparative modern human fore have grown more slowly than the LI1 and LI2 of southern
sample. Both the 95% confidence intervals and Dunnett’s si-
multaneous t tests show that Neandertals have lower means than
Table 5. Dunnett’s t tests for differences in perikymata count
the Inuit, higher means than the southern Africans, and means
means across populations, separated by tooth type
that are not significantly different from those of the sample from
Newcastle. Thus, assuming equivalent mean and modal periodi- Difference
cities across the samples, Neandertals form their teeth more Tooth type of means t value P value
quickly than the Inuit and more slowly than southern Africans UI1: Neandertal subtracted from
and cannot be distinguished in their imbricational enamel for- Inuit 32.30 4.259 0.0002
mation times from the Newcastle sample. Newcastle 27.18 4.103 0.0004
All one-way ANOVAs for difference in mean perikymata Southern African 20.60 3.137 0.0073
counts across populations for each tooth type were statistically UI2: Neandertal subtracted from
significant, with P values 0.001 (results not shown). Table 5 Inuit 7.96 1.148 0.4936
shows Dunnett’s simultaneous t tests by tooth type for mean Newcastle 9.69 1.542 0.2729
Southern African 28.63 4.765 0.0001
UC: Neandertal subtracted from
Inuit 25.738 3.0512 0.0082
Newcastle 10.584 1.7206 0.2007
Southern African 4.121 0.6296 0.8481
LI1: Neandertal subtracted from
Inuit 0.67 0.105 0.9986
Newcastle 2.10 0.344 0.9591
Southern African 25.22 4.286 0.0002
LI2: Neandertal subtracted from
Inuit 12.36 1.763 0.1743
Newcastle 22.60 3.185 0.0061
Southern African 42.23 6.461 0.0000
LC: Neandertal subtracted from
Inuit 38.500 4.2728 0.0002
Newcastle 40.085 4.7299 0.0001
Fig. 3. Ninety-five percent confidence intervals for mean perikymata counts Southern African 3.825 0.5044 0.9128
(all anterior teeth combined).
14200 www.pnas.org cgi doi 10.1073 pnas.0503108102 Guatelli-Steinberg et al.
5. this is true. What is known is that across primate species,
gestation length, age of weaning, age of female sexual maturity,
and life span are highly correlated with the age at which the
permanent first molar erupts (11, 12). In addition, brain weight
correlates with first molar eruption with an r of 0.98 across 21
species of primates (11, 12). Dean et al. (13) have noted that, in
modern humans and Paranthropus, the lower canine completes
enamel formation at about the same time that the first molar
emerges. If this were also the case for Neandertals, whose
calcification patterns are similar to those of modern humans
(27), then our data indicate that the time of Neandertal first
molar emergence was not different from that of modern south-
ern Africans.
Fig. 4. Distribution of hypothetical periodicities (represented by solid circles) It has previously been suggested that the third molar erupted
for Neandertal lower I2 sample under the assumption of 15% shorter growth at 15 years of age in Neandertals (28), possibly because of a
periods than the southern African lower I2 sample. Actual periodicities for the systemically accelerated dental development schedule (15). Yet,
southern African lower I2 sample shown on same histogram, represented by
the possibility that Neandertals erupted their third molars early
triangles.
does not necessarily imply rapid overall dental development.
Advanced third molar eruption in Neandertals may have been
Africans. It is possible that the UI1, UI2, UC, and LC of the result of their large jaws and retromolar spaces that made
Neandertals grew in the same amount of time as in southern early eruption of the third molar possible (29).
Africans, because the hypothetical periodicities fall in the range It has also been suggested that if Neandertals suffered high
of 6–10 days. Under the assumption of growth periods abbre- adult mortality rates, then they might be expected to have had
viated by 15% in Neandertal teeth relative to southern Africans, abbreviated periods of childhood growth (10, 15). Adult mor-
the hypothetical periodicities of all six anterior tooth types tality rates directly select for the timing of maturation across
include periodicities 6. It is therefore not possible for Nean- mammals; a larger risk of dying selects for rapid maturation (9,
dertal imbricational enamel formation times to have been 15% 30, 31). However, Smith (32) notes that if Neandertals had
shorter than those of the southern Africans in our sample. To accelerated life histories, then this would leave them with a
illustrate, Fig. 4 shows the hypothetical periodicities for the ‘‘peculiar’’ relationship between brain size and maturation, ‘‘two
Neandertal lower I2 under the assumption of a growth period variables that are rarely of step.’’ Because large brains require
15% shorter than that of the southern African lower I2. The extended periods of childhood growth (1–7, 33), the presence of
actual periodicities of the southern African lower I2 sample are large brains in Neandertals suggests that their adult mortality
shown on the same histogram. Note that several teeth in the risks were not high enough to have prevented them from
Neandertal sample would be required to have periodicities 6 evolving prolonged growth periods.
for this assumption to be true. Our study demonstrates that Neandertal anterior tooth for-
mation times are encompassed within the wide range of variation
Discussion that we have shown exists within modern humans. Based on
This study demonstrates that Neandertal anterior tooth imbri- comparison to an Upper Paleolithic–Mesolithic modern human
cational enamel formation times are within the range of variation sample, Ramirez-Rozzi and Bermudez de Castro (15) claimed
´
that Neandertal tooth growth was abbreviated relative to mod-
that three modern human populations exhibit. Our results
ern humans, and from this inference, they concluded that
challenge the central conclusion of Ramirez-Rozzi and Bermu- ´
Neandertals grew up more quickly. Our study arrives at a
dez de Castro (15) that Neandertals had significantly abbrevi-
different conclusion for two reasons. First, we did not rely on the
ated anterior tooth growth periods relative to modern humans.
assumption of a constant periodicity; instead, we additionally
Based solely on a modern human sample from the Upper
considered the range of periodicities that are present within a
Paleolithic–Mesolithic, Ramirez-Rozzi and Bermudez de Castro
´
sample. Second, unlike Ramirez-Rozzi and Bermudez de Castro
´
(15) suggested that Neandertals had anterior tooth imbricational
(15), we compared Neandertals to a broad modern human
enamel formation spans 15% shorter than modern humans. Yet
comparative sample.
if we assume, as these authors do (15), a common average
Our data show that anterior tooth perikymata counts cannot
periodicity across all human samples, then the Neandertals in be used to distinguish Neandertal growth rates from the range
our study would have been forming their teeth over longer time of variation in modern humans once the latter is adequately
periods than a sample of modern southern Africans, in about the assessed. If anterior tooth crown formation periods reflect
same amount of time as the sample from Newcastle, and in overall growth periods, then our study suggests that Neandertals
shorter periods than a sample of Inuit. When the relationship did not reach adulthood any more quickly than do modern
between periodicity and total perikymata counts is used to humans.
analyze the unknown periodicities in our Neandertal sample, our
analysis indicates that the lower incisors of Neandertals are This study was supported by Leakey Foundation grants to D.G.-S. and
growing more slowly than those of southern Africans, whereas C.S.L. and by a College of Social and Behavioral Sciences grant to
for canines and upper incisors, Neandertal and southern African D.G.-S. from Ohio State University. We are grateful to the following
imbricational enamel growth may be comparable. Again, using people and institutions for access to their collections: Kevin Kuyken-
the relationship between periodicity and total perikymata dall and Cynthia Reid (Medical School of the University of the
counts, we show that Neandertal teeth could not have been Witwatersrand, Johannesburg, South Africa), Jakov Radovcic (Croat-
ˇ´
ian Natural History Museum, Zagreb, Croatia), Christopher Stringer
ANTHROPOLOGY
growing in 15% less time than those of southern Africans.
and Robert Kruszynski (Natural History Museum, London), Yoel Rak
Do our results therefore imply that Neandertals shared the and Alon Barash (Tel Aviv University, Tel Aviv), Almutt Hoffman
prolonged childhoods of modern humans? This question could (Museum fur Vor- und Fruhgeschichte Archaeologie Europas, Ber-
¨ ¨ ¨
be answered with a definitive ‘‘yes’’ if enamel formation in lin), Ivana Jarasova (Anthropos Institute, Sankt Augustin, Germany),
anterior teeth were directly linked to the length of the childhood Phillipe Mennecier (Musee de l’Homme, Paris), and Ian Tattersall and
´
growth period. Unfortunately, it is currently not known whether Ken Mowbray (American Museum of Natural History, New York). We
Guatelli-Steinberg et al. PNAS October 4, 2005 vol. 102 no. 40 14201
6. acknowledge the skill and excellence of Pamela Walton, who prepared versity of Illinois, Chicago) for use of the images in Fig. 1. Christopher
all of the slides used in this study. We also thank Dale Hutchinson for Dean, Gary Schwartz, Rebecca Ferrell, Bruce Floyd, and Tanya Smith
his advice, Dan Steinberg for extensive support, Cathy Cooke and provided helpful comments on the manuscript. We thank the anony-
James Patrick Bell for making epoxy replicas, and Cameron Begg and mous reviewer for comments that helped improve the clarity of the
Hank Colijn (Center for Electron Optics, Ohio State University) for paper. Finally, the following people gave advice on the collections, and
their SEM assistance. We thank Tanya Smith (Max Planck Institute for their help is also greatly appreciated: Shara Bailey, Jeff Schwartz, and
Evolutionary Anthropology, Leipzig, Germany) and Jay Kelley (Uni- Trent Holliday.
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