AJAX allows web pages to be updated asynchronously by exchanging data with a web server behind the scenes, allowing parts of a page to change without reloading the entire page. Tuenti uses AJAX extensively to update parts of their single-page application, caching content on both client and server sides for scalability. They route requests to different server farms based on client location and cache content to improve performance. Tuenti serves billions of images per day using multiple CDNs and pre-fetches content to minimize load times.