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Morphology of Angiosperms - 2
Stem	
	
	
Dr.	A.S.	Naik	
Head,	Dept	.	of	Botany	
S.B.E.S.	COLLEGE	OF	SCIENCE
What	are	stems?
Types	of	stems
Forms	of	stems
Culms	and	Caudex	
						Euphorbia	caudex	
						Bamboo	culms
Modifica;ons	of	stem
Bulbs	and	Tubers	
Bulb	of	Onion	 Tubers	of	Potato
Corms	
Yam-Elephant	foot	corm			
Amorphophallus		 Colocasia	corm-arbi
Rhizome	
e.g.	Ginger,	Turmeric
Aerial	modificaIons	
Thorns:	e.g.	Indian	Plum	
•  Axillary	buds	modified.	
•  Long/short,	sharply	
pointed	structures	
•  Give	protec;on	
Tendrils:	e.g.	Passiflora		
•  Axillary	buds	modified	into	
•  coiled	wiry	tendils.	
•  Found	in	weak	stemmed	
plants.	
•  Help	in	climbing	on	hosts.
Aerial	modifica<ons-contd.	
•  Phylloclads	of	Opun*a	 •  Phylloclades:	
1.  Greek:	Phyllos-leaf,		klados-
branch.	
2.  A	flaQened	stem	resembling	a	
leaf	
3.  Performs	photosynthesis.	
4.  Leaves	modified	into	spines.	
5.  FlaQened	green	stems		also	
							holds	water.
Cladodes	
•  Cladodes:	
1.  La;n-Kladodes=having	many	
shoots.	
2.  A	phylloclade	of	one	or	two	
internode		
3.  a	flaQened	leaf	like	stem	arising	
in	the	axils	of	a	minute,	bract-like,	
true	leaf.		
Cladodes:	e.g.	Asperagus
Cladophylls	
•  Cladophyll	of	Ruscus.	
						“Cladophylls,”	or	highly	
modified	stems	that	have	
been	flaQened	to	
essen;ally	serve	the	
purpose	of	a	leaf.
Bulbils	
•  Bulbils:	e.g.	Dioscoria	
1.  A	speical	mul;cellular	body	
developed	from	floral/axillary	
bud,meant	for	reproduc;on.		
2.  The	axillary	bud	develops	into	a	
bulbil.		
3.  This	bulbil	detaches	from	the	
mother	plant	and	grows	up	into	a	
new	independent	plant.
Sub-aerial	modifica;ons
Stolons	
•  										
•  Lateral	branches	called	stolon	
originate	from	the	
underground	stem.	
	
•  	Grow	horizontally	outwards	
for	a	varying	distance	in	the	
soil.	
	
•  Ul;mately	emerges	out	of	the	
ground	and	develops	into	a	
new	plant.
Runner	
•  The	runner	arises	from	the	
base	of	the	stem	as	a	lateral	
branch		
•  Runs	along	the	surface	of	the	
soil.	Dis;nct	nodes	and	
internodes.	At	each	node	
produces	roots	below	and	
leaves	above.	
•  		Many	runners	are	produced	
by	the	mother	plant	and	they	
spread	out	on	the	ground	on	
all	sides.		
•  Upon	separa;on	of	a	runner,	
they	are	capable	of	growing	
independently.	
						E.g.,	Oxalis,	Fragaria,		
						Centella	asta<ca,	Mentha
Offset	
•  An	offset	is	a	short	
thick	runner	like	
branch	which	produces	
a	new	plant	at	its	;p.		
•  The	offsets	grow	in	all	
direc;ons	from	the	
main	stem	of	the	
parent	plant.		
•  If	any	injury	results,	it	
is	capable	of	leading	an	
independent	
existence.	E.g.,	Pis;a,	
Eichhornia.
Sucker	
•  A	lateral	branch	arising	
close	to	the	ground	level.	
•  	Traveling	underground	
for	some	distance.	
•  	Turning	up	at	its	end	and	
producing	a	new	plant	is	a	
sucker.		
•  Eg.	Chrysanthemum

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Stem