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Article scientifique
Arid rangelands in the Cholistan Desert
(Pakistan)
Rubina Akhter1
Muhammad Arshad2
1
National Herbarium Programme,
National Agricultural Research Council
(NARC),
Islamabad,
Pakistan
<rubinaflora@yahoo.com>
2
Cholistan Institute of Desert Studies,
Islamia University,
Bahawalpur,
Pakistan
<arshadbwp@yahoo.com>
Abstract
The Cholistan Desert, once a prosperous, lively, and thriving jungle is now by and
large a desolated piece of land. Its productivity potential is on the decline despite the
fact that the number of animals in this desert is on the increase. This sandy desert is
situated in the southern part of Punjab (Pakistan) with highly saline soils and a
brackish subsoil aquifer. It supports a human population of 110,000 pastoral
nomads depending exclusively upon livestock for their livelihood. Life sustainability in
this desert revolves round annual precipitation. The summer in the desert is extremely
harsh and punishing. Some xeric plant species do survive during severe droughts but
undergo tremendous grazing pressure leading to partial eradication; as result, the
flora and fauna have been thinning out gradually with the increasing severity of
desertization. The paper highlights the range livestock production in the Cholistan
Desert.
Key words: desert, desertization, pastoralism, rangelands, Pakistan.
Résumé
Les parcours arides et désertiques du Cholistan (Pakistan)
Le désert du Cholistan qui fut autrefois une jungle vivante et prospère n’est plus qu’un
territoire désolé. Son potentiel de productivité est en déclin bien que le nombre
d’animaux s’accroisse dans ce désert. Il s’agit d’un désert sableux situé dans le sud
du Punjab, avec des sols très salés et un aquifère phréatique d’eau saumâtre dans le
sous-sol. Il comprend une population de 110 000 pasteurs nomades dont la survie
dépend entièrement de leur bétail. La durabilité de la vie dans ce désert tourne
toujours autour du problème des précipitations annuelles. Certaines espèces végéta-
les arrivent à survivre pendant les sécheresses sévères mais sont alors soumises à des
pressions pastorales considérables conduisant à leur éradication partielle. Il s’ensuit
que la flore et la faune se sont progressivement éclaircies et que la désertification
s’est aggravée. Cet article souligne le déclin de la production pastorale et animale
dans le désert du Cholistan.
Mots clés : désert, désertification, pastoralisme, parcours, Pakistan.
Introduction
Location
The Cholistan desert is located in southern
Punjab extending through the Nara and
Thar deserts of Sindh (Pakistan) between
latitudes 27o
42′ and 29o
45′ N and
longitudes 69o
52′ and 75o
24′ E
(figure 1), covering about 2.6 million
hectares [1-4]. Soils are classified as
either saline or saline-sodic, with pH
ranging from 8.2 to 8.4 and from 8.8 to
9.6, respectively. Based on topography,
parent material, soil and vegetation, the
Cholistan Desert can be divided into two
geomorphic regions; the northern region
is called Lesser Cholistan bordering canal-
irrigated areas covering about 7,770 km2
and the southern region is called Greater
Cholistan and covers about 18,130 km2
SĂ©cheresse 2006 ; 17 (1-2) : 210-7
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[1-3]. The old Hakra riverbed (dried about
600 years ago) is the dividing line
between the two regions. Lesser Cholistan
comprises the desert margin and includes
all the area north of the Hakra, while
Greater Cholistan is essentially the area
south of the old Hakra riverbed. The north-
ern part of Lesser Cholistan includes an
irrigation zone of 280,000 hectares serv-
ed by a canal network, where only
130,000 hectares are commandable but
only a small part is actually irrigated.
Climate and soils
Cholistan is a hot hyperarid sandy desert.
The mean annual rainfall varies from less
than 100mm in the west to 200mm in the
east, chiefly falling during monsoon (July
through September). Rainfall is very incon-
sistentinquantityanddurationandprolong-
ed droughts are common once every
10 years. Temperatures are high in summer
and mild in winter with no frost. The mean
summer temperature (May-July) is
34-38o
C with the highest reaching over
51.6o
C [5,6] (figure 2). The soils are
generally saline, alkaline and gypsifer-
ous composed of granites, schists,
gneiss, and slates. The dunes reach an
average height of about 100m in
Greater Cholistan and about 30m in
Lesser Cholistan [2-4, 7-9].
Lesser Cholistan consists of large saline allu-
vial flats (locally called dahars) alternating
with low sandy ridges/dunes. The clayey
flat areas in Lesser Cholistan are generally
homogenous to a depth ranging from 30
to 90cm. These soils are classified as
either saline or saline-sodic, with pH rang-
ing from 8.2 to 8.4 and from 8.8 to 9.6,
respectively. Greater Cholistan is a wind-
resorted sandy desert and comprised of
old river terraces, large sand dunes and
less interdunal flat areas.
There are no permanent, natural bodies of
surface water in Cholistan. Factors like low
rainfall, high rate of water infiltration, and
high evaporation rate prevent the natural
accumulation of surface water [1].
Rainwater is collected in man-made dug-
Nawan Kot
Derawar Fort
Din Garh
Yazman
Mouj Garh
River
Hakra
O
ld
Bed Marot Fort Abbas
Khan Garh
Bijnot
Islam Garh
SCALE
0 20 40 60 80 100 Km.
Lahore
Cholistan
JAMMU &
KASHMIR
(Disputed Territory)
Quetta
Arabian Sea
I
N
D
I
A
A
F
G
H
A
N
IS
T
A
N
BALUCHISTAN
SINDH
IRAN
Karachi
CHOLISTAN DESERT
B A H
A W
A L P U
R
(District)
R
A
H
I
M
Y
A
R
K
H
A
N
(
D
i
s
t
r
i
c
t
)
S
I
N
D
H
B
A
H
A
W
A
L
N
A
G
A
R
(
D
i
s
t
r
i
c
t
t
)
N
C H O L I S T A N
I
I
N D
A
PAKISTAN
PUN JAB
N.W.F.P.
Peshawar
Figure 1. Location map of Cholistan.
0
10
20
30
40
50
60
Rainfall
0
5
10
15
20
25
30
35
40
45
Temperature
Rainfall (mm) Temperature (°C)
J
a
n
u
a
r
y
F
e
b
r
u
a
r
y
M
a
r
c
h
A
p
r
i
l
M
a
y
J
u
n
e
J
u
l
y
A
u
g
u
s
t
S
e
p
t
e
m
b
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r
O
c
t
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N
o
v
e
m
b
e
r
D
e
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e
m
b
e
r
Figure 2. Ombrothermal diagram of Cholistan Desert.
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out water ponds called tobas. Tobas are
made in clayey flats locally called dahars
with a large catchment area to avoid the
loss of runoff and water percolation.
Underground water is at a depth of
30-50m, generally brackish, containing
salts 9,000-24,000 mg/L [3, 7].
Socioeconomic aspects
of pastoralism
The total human population of this desert is
around 110,000 nomadic pastoralists.
The majority of the people live on the
periphery of the desert and the interior of
the desert is thinly populated. The econ-
omy of the region is predominantly pastor-
al and people have been practising a
nomadic life style for centuries. The
nomads own smaller to large herds of
camels, cattle, sheep and goats. The inte-
rior desert area is not connected by a
modern communication system and sandy
desert tracks are used for travel by camels
or jeeps. Local people use camels as a
mode of transportation. Habitations are
small and extremely scattered [3, 10].
The pastoral system is characterized by a
mass movement of animals and people
throughout the year in search of water and
forage within the desert. The onset and
distribution of monsoon rain mainly dictate
the pattern of movement of nomadic herd-
ers (figure 3). Around the months of
March or April, depleted feed and
water resources in the interior of the
desert force nomadic households and
their herds to move towards nearby irrig-
ated areas. Supplementary income is
earned through unskilled labour in
towns or irrigated farms. In irrigated
agricultural fields the pastoralist nomads
have free grazing of livestock on wheat
stubbles, drinking water for humans and
livestock and markets for livestock and
their by-products. In return, farmers in
the irrigated agricultural lands obtain
sufficientlabourforcropharvesting,farm-
ing operations and animal manure to
enhance soil fertility through camping of
livestock on fallow fields.
The nomads and their herds return to the
desert around July or August with the onset
of monsoon showers. Distances travelled
during this movement vary from 10 to
100 km. While in the desert, the natural
vegetation is the main source of fodder
and tobas serve as a source of drinking
water for both nomads and their livestock
[10]. Tobas belonging to one clan are
generally located close to each other
(often within a 2-4-km radius). During
October and November, when water
resources dry up, each clan moves its
herds to semi-permanent settlements where
primitive unlined wells and kunds (usually
lined) are available [1, 3]. Upon exhaust-
ion of these water resources, the desert is
vacated and nomads move to peripheral
areas of the desert where water and
fodder are available.
The economy of these nomads entirely
depends on fragile and meagre natural
resources associated with inconsistent rain
pattern. Job opportunities are confined to
labour in agricultural fields or other minor
activities due to lack of education or skilled
training. Most of the nomads live below
poverty line in the absence of basic human
needs like clean drinking water or suffi-
cient food, health and education for their
children. Livestock breeding, improvement
of performance or range management is
not practised scientifically.
Few nomads manage to shift milk-
producing animals near the roadside to
earn some living. Livestock is used for
exchange of gifts in communal ceremonies
like weddings, tribal celebrations, child-
birthorfunerals,andanimalsareslaughter-
ed for feast for the guests. A nomad’s
status in the desert life-style is chiefly deter-
mined by the size of herd. All livestock are
indigenous breeds well adapted to climat-
ic conditions. Herd reproductive perform-
ance is generally poor with low birth rates
and high mortality due to poor nutrition,
lack of healthcare and climatic stresses [1,
3, 11]. Veterinary health care centres are
not available to majority of livestock.
All the nomads have an unwritten code of
ethics for their range territories and use of
water points: tobas, kunds, or wells. Each
clan has access to traditionally-defined ter-
ritories to use for grazing irrespective of
the condition of the range vegetation. The
exhaustive use of range vegetation has
resulted in the decline of palatable species
especially grasses, overall vegetation
cover, malnutrition of livestock and degrad-
ing wildlife habitats.
Range vegetation
Phytogeographically, the vegetation of the
Cholistan desert belongs to the Nubo-
Sindhian Province of the Sudanian Region
and is typical of arid regions [1].
ThesparsevegetationofCholistanisdomin-
ated by bush forming perennial shrubs
with scattered small trees. A large number
of annual and ephemeral species appear
after rains, complete their life cycle in a
short span and dry up after dispersing
seeds. Most of the shrubs and trees are
leafless or have reduced and modified
leaflike structures. Several members of
Chenopodiaceae like Haloxylon, Suaeda,
Salsola form the dominant species along
with the hardy grasses of rangelands.
Most of the species have a remarkable
capacity to regenerate even in minimum
rainfall. The seeds are also hardy enough
to withstand severe high temperature, arid-
ity and multiple years of drought. Rainfall
is generally confined to few showers in the
months of July to September. The rainfall
ranging from 100-300 makes the desert
green with vegetation cover up to 60% in
some areas.
The vegetation of Lesser Cholistan is domin-
ated by several species of shrubs and
perennial grasses dotted by few species of
small trees. However, in Greater Cholistan
trees are generally lacking, the shrubs are
sparser and species diversity is poor.
Vegetation cover is poor on sand dunes
and unstable sand dunes are devoid of
any vegetation. Some interdunal areas
Stay at settlements
in irrigated areas
Stay at semi-permanent
settlements
Stay in the desert
at the tobas
W
i
n
t
e
r
P
o
s
t
-
M
o
n
s
o
o
n
S
u
m
m
e
r
Spring
D
E
C
Figure 3. Yearly movement pattern of pastoralists in Cholistan.
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have a better vegetation depending on the
water retention capacity of the soil.
Ground water table is very low and in
most of the cases not fit for drinking.
Main vegetation types
It is difficult to define distinct community
types associated with habitats like sand
dunes, interdunal flat plains, saline plains
or dry water-courses flooded periodically
during rain. Some species are typically
associated with a distinct habitat type.
Some main vegetation types based on
floristically dominant and structurally
important species are described below.
• Haloxylon-Calligonum Community
This is a widespread community type
dominated by leafless shrubs of Haloxylon
stocksii and Calligonum polygonoides.
Other associated species include Lasiurus
scindicus, Cymbopogon, jwarancusa,
Suaeda fruticosa, Dipterygium glaucum,
Crotalariaburhia,etc.Thistypeofcommun-
ities generally covers flat plains with com-
pact soil and saline conditions. Several
species of annuals and ephemerals
emerge after rains, covering the ground
but contributing very little biomass.
The shrubs have strong root systems and
hold the soil together protecting the soil
from wind erosion. Sometimes small
mounds of soil are developed around the
bushes while the surrounding soil is blown
away with strong winds.
• Acacia-Tamarix Community
The Acacia nilotica and Tamarix aphylla
community is found in peripheral areas
of Cholistan. Acacia nilotica is locally
considered as poor man’s teak wood for
its valuable timber used for cheap furniture
and agriculture implements. Leaves
provide forage for livestock. Tamarix bran-
ches are used for thatching rooftops and
fences around livestock enclosures. Trees
also provide shade for resting livestock
seeking shelter from strong sun. Several
species of grasses and herbs are associat-
ed with these communities like Cenchrus,
Pennisetum, Eragrostis, Fagonia, Diptery-
gium, Indigofera, etc. Large hemispherical
clumps of Heliotropium crispum are com-
mon in the peripheral regions of the desert
which hold soil from erosion and conserve
water.
• Prosopis-Salvadora Community
The Prosopis cineraria and Salvadora
oleoides community is generally found in
Lesser Cholistan regions where rainfall is
not less than 100mm. Prosopis cineraria is
considered as an excellent forage species
with a very deep root system, very well
adapted to the desert conditions. The
branches are extensively lopped for feed-
ing young lambs and goats not able to
roam around for forage. Generally the top
shoots are left for future growth and lower
branches are cut for livestock. It also forms
excellent firewood for cooking as well as
charcoal making. Salvadora oleoides is
an evergreen tree with deformed trunk and
a deep root system heavily grazed by
camels and goats. It is a very slow-
growing tree with poor regeneration
through seeds but it profusely produces
suckers forming large clumps. Sometimes
only clump-forming large bushy structures
are seen due to heavy browsing; tree-like
shape is rare; ripe fruits are also edible.
Other associated species include Acacia
nilotica, Zizyphus nummularia, Tamarix
aphylla, grass species like Lasiurus, Cym-
bopogon, Sporobolus, Panicum, Cen-
chrus,Aristida,etc.Someannualsassociat-
ed with this community include Zygophyllum,
Dipterygium, Indigofera, Glinus, Fagonia,
Heliotropium, etc.
• Lasiurus-Cymbopogon Community
Perennial species of clump forming grass
species like Lasiurus scindicus and Cymbo-
pogon jwarancusa are developed on poo-
rer soils on flat plains with compact soil.
This community type is generally poor in
species diversity, having mainly annual
short-lived species appearing after rains.
These grass species are palatable in early
stage of growth; coarse hard old leaves
are not generally grazed.
Lasiurus scindicus and Cymbopogon jwa-
rancusa are abundantly distributed throu-
ghout the Cholistan Desert. Scattered
shrubs of Haloxylon, Suaeda, Crotalaria,
Aerua are also sparsely found.
• Acacia-Pennisetum Community
Acacia jacquemontii and Pennisetum divi-
sum are generally associated with dry
water courses with better soil and moisture
retention capacity. Acacia jacquemontii is
atallshrubbyspecieswithextensivebranch-
ing from the base. It is a good forage
species and a preferred firewood shrub in
the desert. Pennisetum divisum is a tall
large clump-forming grass with a good
soil-binding ability preventing soil erosion
from strong winds and flash floods. This is
an excellent fodder grass relished by live-
stock producing green tender leaves in
minimum moisture conditions.
Other associated species include species
of Convolvulus, Launnaea, Indigofera, Tri-
bulus, Trianthema, Neurada, etc. Several
grass species include Ochthochloa, Pani-
cum, Aeluropus, Stipagrostis, Cenchrus,
etc.
• Aerua-Crotalaria Community
This community is common on poor saline
soils. Usually, large shrubs and trees are
absent. Grasses like Sporobolus, Ochtho-
chloa, Desmostachya species are found
having stiff leaves, grazed when young.
They are usually found in interdunal saline
sandy soils.
• Capparis-Suaeda Community
Capparis aphylla and Suaeda fruticosa
are also widespread in poor soils with
inadequate moisture contents. Capparis
aphylla, a leafless shrub, sometimes
attains tree-like form; its branches are used
for thatching roof tops. Suaeda fruticosa
forms evergreen large-sized bushy dome-
like clumps; leaves are reduced to scales.
Vegetation is very sparse, poor in species
diversity; short-lived annual plants appear
after rains for a short period.
• Leptadenia-Calotropis Community
The Leptadenia pyrotechnica and Calotro-
pis procera community develops in poor
sandy soils; not palatable for livestock, it
spreads in areas with heavy grazing pres-
sure. Leptadenia pyrotechnica is a much
branched leafless shrub not grazed by
livestock but the branches are used for
thatching roofs and fencing. Cyperus
conglomerates is a compact clump-
forming sedge with roots having good
sand-binding ability, found frequently on
sand dunes under poor moisture condi-
tions, grazed when young.
Grazing potential of ranges
The vegetation in Cholistan is typical of
arid and semiarid climate consisting of
xerophytic species adapted to high season-
al temperatures, low humidity, moisture
fluctuations and wide variety of edaphic
conditions [12]. Compared to the hyper-
arid southern region, the vegetation cover
is relatively better in Eastern Cholistan
(200mm rainfall zone). A wide range of
nutritious and drought-tolerant plant spe-
cies (128 species belonging to 33 fami-
lies) of grasses, herbs, shrubs and trees
occupy this desert [9]. Hardy species
adapted to high temperature and low soil
moisture include Lasiurus scindicus, Sporo-
bolus iocladus, Aeluropus lagopoides and
Cyperus conglomerates.
Major forage plants and corresponding
characteristics, use and grazing animals
in the Cholistan Desert are listed in
table 1.
Pastoralism in Cholistan
Pastoral
The livestock depend entirely on self-
regenerating and self-maintaining range
resources throughout the year or resort to
canal and riverain areas for grazing with
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Table I. Major forage plants of the Cholistan Desert and corresponding characteristics, use, and grazing animals.
Plant Species Parts used Palatability Season Frequency Grazing animals
Perennial Grasses
Stipagrostis plumosa Lv, Sh HP Monsoon, Spring V. Common Cattle, sheep, goat
Cenchrus ciliaris Lv, Sh HP Monsoon, Spring Rare Cattle, Sheep Goat
Cymbopogon jwarancusa Lv, Sh LP Monsoon, Spring V. Common Sheep, Cattle
Ochthochloa compressa Lv, Sh HP Monsoon, Spring V. Common Cattle, Sheep Goat
Lasiurus scindicus Lv, Sh HP Monsoon, Spring V. Common Cattle, Sheep Goat
Panicum antidotale Lv MP Monsoon, Spring Rare Cattle, Sheep Goat, Camel
Panicum turgidum Lv HP Monsoon, Spring Common Cattle, Sheep Goat, Camel
Sporobolus iocladus Lv MP Monsoon, Spring Common Cattle, Sheep Goat, Camel
Aeluropus lagopoides Lv, Sh H.P Monsoon, Spring Common Cattle, Sheep Goat, Camel
Sedge
Cyperus conglomeratus Sh, Lv HP Monsoon, Spring, Winter Common Cattle, Sheep Goat, Camel
Annual Grasses
Aristida adscensionis Sh, Lv MP Monsoon Common Sheep, Goat
Aristida mutabilis Sh, Lv MP Monsoon Common Sheep, Goat
Aristida funiculata Sh, Lv MP Monsoon Common Sheep, Goat
Aristida hystricula Sh LP Monsoon Common Sheep, Goat
Cenchrus biflorus Sh, Lv MP Monsoon Common Cattle, Sheep, Goat
Cenchrus prieurii Sh, Lv HP Monsoon Common Cattle, Sheep, Goat
Enneapogon desvauxii Lv, Sh HP Monsoon Rare Sheep, Goat
Eragrostis barrelieri Lv, Sh HP Monsoon Common Sheep, Goat
Leptothrium senegalense Lv, Sh MP Monsoon Common Cattle, Sheep, Goat
Tragus racemosus Lv, Sh MP Monsoon Common Cattle, Sheep, Goat
Forbs
Tribulus longipetalus Sh, Lv, Fl HP Monsoon Common Cattle, Sheep Goat, Camel
Tribulus terrestris Sh, Lv, Fl HP Monsoon, Winter, Spring Rare Cattle, Sheep Goat, Camel
Fagonia cretica Lv LP Monsoon, Winter, Spring Common Sheep
Anticharis linearis Lv, Sh LP Monsoon Common Sheep, Goat
Indigofera sessiliflora Lv, Sh, Fl MP Monsoon Rare Sheep, Goat
Indigofera argentea Lv, Sh, Fl MP Monsoon Common Sheep, Goat
Indigofera hochstetri Lv, Sh, Fl MP Monsoon Common Sheep, Goat
Neurada procumbense Lv, Fl MP Winter Rare Sheep, Goat
Glinus lotoides Lv, Sh, Fl LP Monsoon, Spring Common Cattle, Sheep Goat, Camel
Euphorbia prostrata Lv LP Monsoon, Spring Common Sheep, Goat
Mukia madraspatana Lv HP Monsoon Common Sheep, Goat
Cressa cretica Lv, Sh MP Monsoon, Spring Common Cattle, Sheep, Goat
Convolvulus microphyllus Lv, Sh MP Monsoon, Spring Common Sheep, Goat, Camel
Launea residifolia Lv, Sh LP Winter Common Sheep, Goat, Camel
Launea nudicaulis Lv, Sh LP Winter Common Sheep, Goat, Camel
Dipterygium glaucum Sh LP Monsoon, Spring V. Common Camel
Cleome brachycarpa Sh LP Monsoon, Spring Common Camel
Farsetia hamiltonii Sh LP Monsoon, Spring V. Common Sheep, Goat
Heliotropium strigosum Lv, Sh MP Monsoon, Spring Common Sheep, Goat, Camel
Heliotropium crispum Lv, Sh LP Throughout the year Common Camel, Sheep
Trianthema portulacastrum Sh MP Monsoon Common Camel, Sheep
Zaleya pentendra Sh MP Monsoon Common Camel, Sheep
Sesuvium sesuvioides Sh MP Monsoon Common Camel, Sheep
Limeum indicum Sh MP Monsoon V. Common Camel, Sheep
Shrubs
Crotalaria burhia Lv LP Throughout the year Common Camel
Aerva persica Lv LP Throughout the year Common Camel
Calotropis procera Lv LP Throughout the year Common Cattle, Sheep, Goat
Pulicaria rajputanae Lv LP Throughout the year Common Camel
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or without purchased fodder for sup-
plementation in the pre-monsoon period.
Agropastoral
Along with range resources, the livestock
also utilize fodder crops from agricultural
farms owned by the pastoralists or by
tenant farmers in the irrigated tract. The
croplands provide food security for both
the livestock and their owners. The impor-
tance of crops varies according to the
status of small owner-pastoralists depend-
ing mainly on free grazing along the
canals or roadsides or on additional pur-
chased fodder to support livestock in the
pre-monsoon period. After the monsoon,
when sufficient forage is available, the
livestock move to desert rangelands for
grazing.
Sedentary pastoralists
Livestock owners living in permanent settle-
ments working as tenant farmers or small
farmers generally use nearby grazing
areas. The livestock comprising mainly
buffaloes, cattle, some sheep and goats
confined to the irrigated fields are mainly
stall-fed or use surrounding areas for graz-
ing and come back in the evening. The
feed resources comprise a variable pro-
portion of crop residues, fodders and peri-
irrigation grazing areas. Due to the com-
petition from the transhumant livestock,
feed supply during the pre-monsoon
period may be constrained.
Transhumant
The majority of livestock and people
seasonally move to the desert and sub-
sequently again towards irrigated mar-
gins. The point of origin is irrigated areas,
and in the desert they have fairly well-
defined range territory for various clans;
however, variation in the route and the
sequence of use of tobas are governed by
the water and forage quantity available in
the range.
The livestock start moving to the desert at
the onset of monsoons in July/August
when the tobas are filled with water. They
stay around the tobas as long as water is
available, and move to the wells or kunds
when the tobas can no longer support the
livestock or people. The salinity in the wells
rises with the advancement of dry season.
The accessible forage is exhausted till
March/April and the salinity in the wells
renders the water unsuitable for cattle.
Livestock start to return towards the irrigat-
ed margins. Only camels and goats are
left to survive on evergreen browse.
The precise times of movements depend
on the rains and the resulting availability
of water and forage in the desert; in good
years water may be available in tobas
until January or February, and in normal
years it may dry out before that.
Desert nomads
The livestock remain all the year in the
desert and are dependent upon the range
resources for fodder and water. Almost all
the camels, about 50% goats and a small
number of cattle are represented in the
system. The animals move with the trans-
humant livestock from one toba to the next
and subsequently to the wells and/or
kunds. However, they remain at the wells
or kunds when the transhumant livestock
retreat to the irrigated margins. Generally,
each clan/group tends to remain at a
particular well or kund until the tobas are
filled again.
Livestock
Breeds
Camels, cattle, sheep and goats are the
predominant types of livestock. Cattle are
the most precious livestock species of
pastoral lifestyle followed by sheep, goats
and camels. The productivity (birth rates)
among cattle is the lowest. Lambing occurs
mainly during the months of January/
ebruary. Goats are kept in smaller num-
bers due to: i) lack of browse when ani-
mals are taken to irrigated areas during
droughts; ii) to avoid predation by jackals;
and iii) because of difficulties encountered
in controlled grazing in irrigated areas.
Camels are also owned mainly for trans-
portation purposes. Notable breeds of
livestock in Cholistan are as follows:
Cattle: Cholistani and Hasari
Goat: Jattal (Cholistan goat)
Sheep: Buchi, khadali, sipli
Camel: Marecha and Brella
Generally, split-herding prevails in cattle
and small ruminants. Small calves in the
case of cows and lambs and kids in case
of sheep and goats are kept at pens near
the tobas, while the adults are allowed to
roam around for grazing in the open
range areas of the desert. Most cattle and
camel are not herded. In the case of the
cattle, the animals are driven out of the
house to graze freely and they return home
by themselves in the evening. The camels,
however, remain in the desert unherded
Table I (cont’d). Major forage plants of the Cholistan Desert and corresponding characteristics, use, and grazing animals.
Plant Species Parts used Palatability Season Frequency Grazing animals
Calligonum polygonoides Sh, Fl MP Throughout the year V. Common Camel, Cattle, Goat
Acacia jacquemontii Lv, Sh, Fl HP Throughout the year Common Sheep, Goat, Camel
Haloxylon recurvum Sh MP Throughout the, year V. Common Camel
Haloxylon salicornicum Sh LP Throughout the year V. Common Camel
Salsola baryosma Sh MP Throughout the year V. Common Camel
Suaeda fruticosa Sh, Lv MP Throughout the Year V. Common Camel
Capparis decidua Sh MP Throughout the Year V. Common Camel, Cow
Zizyphus nummularia Lv HP Throughout the Year Common Sheep, Goat, Cow, Camel
Leptadenia pyrotechnica Sh NP Throughout the Year Common Camel
Trees
Prosopis cineraria Lv, Sh, Fl, Fr HP Throughout the year V. Common Sheep, Goat, Cow, Camel
Acacia nilotica Lv, Sh, Fl, Fr HP Throughout the year Common Sheep, Goat, Cow, Camel
Prosopis juliflora Lv, Fr MP Throughout the year Rare Sheep, Goat, Camel
Zizyphus spina christi Lv HP Throughout the year Rare Sheep, Goat, Cow, Camel
Tamarix aphylla Sh MP Throughout the year Common Camel
Salvadora oleoides Lv, Sh HP Throughout the year Rare Sheep, Goat, Cow, Camel
Lv = leaves; Sh = shoot/stem; Fl = flower; Fr. = fruit; HP = highly palatable; MP = moderately palatable; LP = less palatable; NP = non palatable.
Sécheresse vol. 17, n° 1-2, janvier-juin 2006 215
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for months. Small ruminants (sheep and
goats) are attended by herdsman mainly
to avoid predation by jackals.
Mortality rate is high among cattle alone
ranges from 5 to 60 percent [1]. The main
causes of mortality are related to droughts,
poor nutrition due to shortage of feed and
water and diseases caused by nutrition
stress. No livestock health facilities are
available in the desert, only limited health
services are available in peripheral small
towns and vaccination of livestock is not
practised regularly.
Livestock production
and availability of feed
The rangelands in Cholistan desert are
monsoonal and forage production in these
rangelands depends heavily on monsoon
rains (amount, time, and frequency). The
health of the animals, their production,
birth and mortality rate relates directly to
the availability of vegetation. During norm-
al rainfall years (100–200mm) green her-
baceous production remains critically low,
particularly for cattle and sheep and ani-
mals remain unnourished. Once or twice
in ten years forage production is sufficient
for year-round grazing of the livestock.
During good years the production of live-
stock and by-products is sufficient enough
when owners have some earnings. After
successive drought years the forage pro-
duction declines drastically and vegeta-
tion growth is practically nil. The grazing
period starts from August till February
during good rainy years.
Both the composition and production of
browse and herbaceous forage vary with
the range types given the same amount of
rainfall. In the best ranges, herbaceous
forage yields 60-70% whereas browse
yields 30-40% of the total forage product-
ion. In moderate ranges, herbage product-
ion varies from 35 to 45% and browse
from 55 to 65%. In poor ranges, herbage
yields less than 20% and browse more
than 80% [1]. Almost all the herbaceous
forage is utilized during monsoon and
post-monsoon season, while green browse
is available in the area throughout the
year. The herbaceous vegetation is very
short-lived and it dries up when the soil
moisture is no longer available. The feed
requirements of camels and goats, in the
winter and pre-monsoon are provided by
green browse (Calligonum and Haloxy-
lon), while cattle and sheep suffer severe
nutritional deficiencies. The winter rains
are very scarce so spring vegetation is
generally poor, represented by few annual
species of grasses and forbs providing
very little biomass for grazing.
At present, the total livestock population in
Cholistan is around 134,798 animal units
(AU) (table 2). The animal unit is consider-
ed as an adult cow weighing 350 kg
(400 kg at international level) and
consuming 7 kg dry matter forage per
day in Cholistani conditions. The total
annual dry matter forage demand in
Cholistan is .34 million tons while the
available dry matter forage is .12 mil-
lion tons. Thus there is an annual short-
age of 0.22 million tons dry matter
forage (table 3). It has been noticed that
at the moment, some 88,655 AU are in
excess of the present carrying capacity
[13]. The rangelands of Cholistan are
under severe threat of degradation
because of overgrazing and harsh climat-
ic conditions. Due to continuous grazing,
the desirable palatable species are
vanishing at an alarming rate and relative-
ly unpalatable species are spreading and
dominating the landscape. The severity of
the problem can be seen from the fact that
such highly desirable grass species
as Pennisetum divisum and Cenchrus
setigerus have disappeared from most
part of the Cholistan desert rangelands.
Livestock production
and health management
Three inter-related aspects of animal
health i.e., feed, water and disease have
been encountered in Cholistan desert.
Deficiencies in the availability of forage
(quantity and quality), drinking water
(saline or polluted) and free mixing of
diseased animals with healthy ones during
grazing expose livestock to various types
of diseases. Veterinary health centres or
hospitals are not available towards the
interior of the desert and very few poorly-
equipped small units are available in peri-
pheral cities. Livestock owners often
become distressed and helpless when their
livestock, particularly cattle fall seriously
ill. Some traditional homemade herbal
preparations are used to treat sick ani-
mals. The mortality rate is very high and
drought conditions increase stress due to
malnutrition and lack of water causing
various diseases. The most common
diseases occurring in various livestock
species are as follows:
– large ruminants: haemorrhagic septicae-
mia, black quarter, foot and mouth
disease, anthrax, mange, surra, camel
pox, endo- and ectoparasites, etc.;
– small ruminants: enterotoxemia, pleuro-
pneumonia, sheep and goat pox, anthrax,
liver fluke, endo- and ectoparasites, etc.
The amount of livestock population that
have died and suffered from various
diseases in Cholistan desert is given
in table 4. Traditionally, reasonable
disease diagnostic know-how is avail-
able at the toba level. However, disease
treatment capabilities of the herders are
very limited and medicines are avail-
able at places that are far away from
their tobas. Therefore, a large number of
Table II. Livestock population and forage demand in the Cholistan Desert (census 1996).
Animal type Population Animal Unit (AU) Dry matter intake per day
(kg)
Annual forage demand (tons)
Cattle 73,048 7,3048 7 186,638
Sheep 14,7429 21,061 1 53,812
Goats 53,680 7,669 1 19,593
Camels 16,510 33,020 14 84,366
Total 290,667 134,798 10 344,409
Table III. Annual demand and supply of dry matter forage in Lesser and Greater Cholistan.
Livestock population (AU) Annual forage demand (tons) Forage available tons Deficit (tons)
Lesser (1,237,000
ha)
Greater (1,369,000 ha) Total (tons) (2,606,000 ha)
134,798 344,409 55,962 61,934 117,896 226,513
216 Sécheresse vol. 17, n° 1-2, janvier-juin 2006
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the animals infected with disease were
either not provided any treatment or
were given indigenous treatment.
Livestock marketing
The pastoral nomads in Cholistan desert
are living very much below the poverty line
due to the lack of education or of skilled
labour and to their entire dependence on
meagre natural resources. The quantity
and quality of livestock is not good enough
to support a family. Marketing is generally
practised through a middleman (locally
called Beopari) who collects animals from
various nomads camping sites near tobas
on the outskirts of small towns. Seasonal,
monthly, and weekly livestock markets
operate at various levels depending on the
availability of animals for sale. Livestock
by-products including homemade mats
prepared from raw wool are also brought
to the markets. Camel bone and camel
skin products prepared with primitive skills
are also prepared on a small scale. But
due to the lack of training or of refined
skills, the market is very limited. The money
earned through sale of livestock is not
enough to meet the requirements for food
or for animal feed and fodder.
Constraints
The major constraint in livestock produc-
tion in Cholistan Desert is the shortage of
sweet water. This is compounded by the
prolonged droughts of many years when
toba water dried out completely.
In Greater Cholistan, feed for livestock is
still available, but the toba water is deplet-
ed and the thirsty herds are forced to
migrate towards semi-permanent settle-
ments where well water is adequate but of
poor and saline quality not fit for drinking.
The wells are unlined and must be re-dug
each season. On the other hand in the
western part (Lesser Cholistan) the
quantities of both water and feed are
inadequate.
Landless pastoralists suffer due to the scarc-
ity of rangelands for grazing in the irrigat-
ed fringes where they work as poorly-paid
labour or as tenant farmers on farmlands
generally used for agricultural crops.
The combination of long distances travel-
led by the livestock in search of forage,
harsh temperature rising above 50o
C, in-
adequacy of feed, undernourishment and
highly saline drinking water from wells, all
contribute to high mortality rates. â– 
References
1. Food and Agriculture Organization (FAO).
Pakistan – Cholistan Area Development Project.
Report No. 59/53 ADB-PAK 58 (Final version).
Rome: FAO, 1993.
2. Akbar G, Arshad M. Developing sustainable
strategies for Cholistan desert: opportunities and
perspectives. Science Vision 2000; 5: 77-85.
3. Akbar G, Khan TN, Arshad M. Cholistan
desert, Pakistan. Rangelands 1996; 18: 124-8.
4. Chaudhry AA, Hussain A, Hameed M,
Ahmad R. Biodivesity in Cholistan desert, Pun-
jab, Pakistan. In: Shazad AM, Charles AW,
Usman AS, eds. Biodiversity of Pakistan. Islama-
bad (Pakistan): Pakistan Museum of Natural His-
tory, 1997.
5. Mughal MR. Ancient Cholistan – archaeology
and architecture. Lahore (Pakistan): Ferozsons
(Pvt.) Ltd., 1997.
6. Arshad M, Akbar G, Rashid S. Wealth of
medicinal plants of Cholistan desert, Pakistan:
conservational strategies. Hamdarad Medicus
2003; XLV: 25-34.
7. Baig MS, Akram M, Hassan MA. Possibilities
for range development in Cholistan desert as
reflected by its physiography and soils. Pak J For
1980; 30: 61-71.
8. Rao AR, Arshad M. Perennial grasses of Cho-
listan desert and their distribution. Proceedings of
a seminar on “Peoples participation in the mana-
gement of resources in arid lands. Bahawalpur
(Pakistan): Cholistan Institute of Desert Studies,
Islamia University, 1991.
9. Arshad M. Rao AR. Flora of Cholistan desert
(Systematic list of trees, shrubs and herbs). J Econ
Tax Bot 1994; 18: 615-25.
10. Arshad M, Rao AR, Akbar G. Masters of
disaster in Cholistan desert, Pakistan: pattern of
nomadic migration. UNEP Desertification Control
Bulletin 1999; 35: 33-9.
11. Mumtaz KK. Habitat and desert: The case of
Cholistan. In: Taylor BB, ed. The changing rural
habitat; Volume 1: Case Studies. Singapore: The
Concept Media/Agha Khan Award for Architec-
ture, 1982.
12. Rao AR, Arshad M, Shafiq M. Perennial
grass germplasm of Cholistan desert and their
phytosociology. Bahawalpur (Pakistan): Cholis-
tan Institute of Desert Studies, Islamia University,
1989.
13. Akbar G, Arshad M. Feasibility of the resto-
ration of vegetation in the Cholistan desert as a
natural habitat of Houbara bustard and other
wild life species. Final Project Report. Lahore
(Pakistan): World Wild Fund (WWF)-Pakistan,
1999.
14. Iqbal M, Farooq U, Basir A, Khan NA,
Malik SZ. In: A baseline survey for the develop-
ment of livestock sector in Cholistan. GmdII, GTZ
P.N.91.2123.7. Lahore: Pak-German Technical
Cooperation (Deutsche Gesellschaft fur Technis-
che Zusammenarbeit, GTZ), Livestock and Dairy
Development Department, 2000.
Table IV. Death and disease incidence among livestock population.
Items Cattle Camel Sheep Goat
Farms reported disease incidence 49.69 5.45 59.02 34.12
Population suffered 5.64 1.65 12.19 19.37
Mortality 2.72 0.55 9.31 7.78
Source: [14].
Sécheresse vol. 17, n° 1-2, janvier-juin 2006 217
Copyright
©
2017
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Libbey
Eurotext.
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Arid rangelands in the cholistan desert

  • 1. Article scientifique Arid rangelands in the Cholistan Desert (Pakistan) Rubina Akhter1 Muhammad Arshad2 1 National Herbarium Programme, National Agricultural Research Council (NARC), Islamabad, Pakistan <rubinaflora@yahoo.com> 2 Cholistan Institute of Desert Studies, Islamia University, Bahawalpur, Pakistan <arshadbwp@yahoo.com> Abstract The Cholistan Desert, once a prosperous, lively, and thriving jungle is now by and large a desolated piece of land. Its productivity potential is on the decline despite the fact that the number of animals in this desert is on the increase. This sandy desert is situated in the southern part of Punjab (Pakistan) with highly saline soils and a brackish subsoil aquifer. It supports a human population of 110,000 pastoral nomads depending exclusively upon livestock for their livelihood. Life sustainability in this desert revolves round annual precipitation. The summer in the desert is extremely harsh and punishing. Some xeric plant species do survive during severe droughts but undergo tremendous grazing pressure leading to partial eradication; as result, the flora and fauna have been thinning out gradually with the increasing severity of desertization. The paper highlights the range livestock production in the Cholistan Desert. Key words: desert, desertization, pastoralism, rangelands, Pakistan. RĂ©sumĂ© Les parcours arides et dĂ©sertiques du Cholistan (Pakistan) Le dĂ©sert du Cholistan qui fut autrefois une jungle vivante et prospère n’est plus qu’un territoire dĂ©solĂ©. Son potentiel de productivitĂ© est en dĂ©clin bien que le nombre d’animaux s’accroisse dans ce dĂ©sert. Il s’agit d’un dĂ©sert sableux situĂ© dans le sud du Punjab, avec des sols très salĂ©s et un aquifère phrĂ©atique d’eau saumâtre dans le sous-sol. Il comprend une population de 110 000 pasteurs nomades dont la survie dĂ©pend entièrement de leur bĂ©tail. La durabilitĂ© de la vie dans ce dĂ©sert tourne toujours autour du problème des prĂ©cipitations annuelles. Certaines espèces vĂ©gĂ©ta- les arrivent Ă  survivre pendant les sĂ©cheresses sĂ©vères mais sont alors soumises Ă  des pressions pastorales considĂ©rables conduisant Ă  leur Ă©radication partielle. Il s’ensuit que la flore et la faune se sont progressivement Ă©claircies et que la dĂ©sertification s’est aggravĂ©e. Cet article souligne le dĂ©clin de la production pastorale et animale dans le dĂ©sert du Cholistan. Mots clĂ©s : dĂ©sert, dĂ©sertification, pastoralisme, parcours, Pakistan. Introduction Location The Cholistan desert is located in southern Punjab extending through the Nara and Thar deserts of Sindh (Pakistan) between latitudes 27o 42′ and 29o 45′ N and longitudes 69o 52′ and 75o 24′ E (figure 1), covering about 2.6 million hectares [1-4]. Soils are classified as either saline or saline-sodic, with pH ranging from 8.2 to 8.4 and from 8.8 to 9.6, respectively. Based on topography, parent material, soil and vegetation, the Cholistan Desert can be divided into two geomorphic regions; the northern region is called Lesser Cholistan bordering canal- irrigated areas covering about 7,770 km2 and the southern region is called Greater Cholistan and covers about 18,130 km2 SĂ©cheresse 2006 ; 17 (1-2) : 210-7 210 SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 2. [1-3]. The old Hakra riverbed (dried about 600 years ago) is the dividing line between the two regions. Lesser Cholistan comprises the desert margin and includes all the area north of the Hakra, while Greater Cholistan is essentially the area south of the old Hakra riverbed. The north- ern part of Lesser Cholistan includes an irrigation zone of 280,000 hectares serv- ed by a canal network, where only 130,000 hectares are commandable but only a small part is actually irrigated. Climate and soils Cholistan is a hot hyperarid sandy desert. The mean annual rainfall varies from less than 100mm in the west to 200mm in the east, chiefly falling during monsoon (July through September). Rainfall is very incon- sistentinquantityanddurationandprolong- ed droughts are common once every 10 years. Temperatures are high in summer and mild in winter with no frost. The mean summer temperature (May-July) is 34-38o C with the highest reaching over 51.6o C [5,6] (figure 2). The soils are generally saline, alkaline and gypsifer- ous composed of granites, schists, gneiss, and slates. The dunes reach an average height of about 100m in Greater Cholistan and about 30m in Lesser Cholistan [2-4, 7-9]. Lesser Cholistan consists of large saline allu- vial flats (locally called dahars) alternating with low sandy ridges/dunes. The clayey flat areas in Lesser Cholistan are generally homogenous to a depth ranging from 30 to 90cm. These soils are classified as either saline or saline-sodic, with pH rang- ing from 8.2 to 8.4 and from 8.8 to 9.6, respectively. Greater Cholistan is a wind- resorted sandy desert and comprised of old river terraces, large sand dunes and less interdunal flat areas. There are no permanent, natural bodies of surface water in Cholistan. Factors like low rainfall, high rate of water infiltration, and high evaporation rate prevent the natural accumulation of surface water [1]. Rainwater is collected in man-made dug- Nawan Kot Derawar Fort Din Garh Yazman Mouj Garh River Hakra O ld Bed Marot Fort Abbas Khan Garh Bijnot Islam Garh SCALE 0 20 40 60 80 100 Km. Lahore Cholistan JAMMU & KASHMIR (Disputed Territory) Quetta Arabian Sea I N D I A A F G H A N IS T A N BALUCHISTAN SINDH IRAN Karachi CHOLISTAN DESERT B A H A W A L P U R (District) R A H I M Y A R K H A N ( D i s t r i c t ) S I N D H B A H A W A L N A G A R ( D i s t r i c t t ) N C H O L I S T A N I I N D A PAKISTAN PUN JAB N.W.F.P. Peshawar Figure 1. Location map of Cholistan. 0 10 20 30 40 50 60 Rainfall 0 5 10 15 20 25 30 35 40 45 Temperature Rainfall (mm) Temperature (°C) J a n u a r y F e b r u a r y M a r c h A p r i l M a y J u n e J u l y A u g u s t S e p t e m b e r O c t o b e r N o v e m b e r D e c e m b e r Figure 2. Ombrothermal diagram of Cholistan Desert. SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 211 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 3. out water ponds called tobas. Tobas are made in clayey flats locally called dahars with a large catchment area to avoid the loss of runoff and water percolation. Underground water is at a depth of 30-50m, generally brackish, containing salts 9,000-24,000 mg/L [3, 7]. Socioeconomic aspects of pastoralism The total human population of this desert is around 110,000 nomadic pastoralists. The majority of the people live on the periphery of the desert and the interior of the desert is thinly populated. The econ- omy of the region is predominantly pastor- al and people have been practising a nomadic life style for centuries. The nomads own smaller to large herds of camels, cattle, sheep and goats. The inte- rior desert area is not connected by a modern communication system and sandy desert tracks are used for travel by camels or jeeps. Local people use camels as a mode of transportation. Habitations are small and extremely scattered [3, 10]. The pastoral system is characterized by a mass movement of animals and people throughout the year in search of water and forage within the desert. The onset and distribution of monsoon rain mainly dictate the pattern of movement of nomadic herd- ers (figure 3). Around the months of March or April, depleted feed and water resources in the interior of the desert force nomadic households and their herds to move towards nearby irrig- ated areas. Supplementary income is earned through unskilled labour in towns or irrigated farms. In irrigated agricultural fields the pastoralist nomads have free grazing of livestock on wheat stubbles, drinking water for humans and livestock and markets for livestock and their by-products. In return, farmers in the irrigated agricultural lands obtain sufficientlabourforcropharvesting,farm- ing operations and animal manure to enhance soil fertility through camping of livestock on fallow fields. The nomads and their herds return to the desert around July or August with the onset of monsoon showers. Distances travelled during this movement vary from 10 to 100 km. While in the desert, the natural vegetation is the main source of fodder and tobas serve as a source of drinking water for both nomads and their livestock [10]. Tobas belonging to one clan are generally located close to each other (often within a 2-4-km radius). During October and November, when water resources dry up, each clan moves its herds to semi-permanent settlements where primitive unlined wells and kunds (usually lined) are available [1, 3]. Upon exhaust- ion of these water resources, the desert is vacated and nomads move to peripheral areas of the desert where water and fodder are available. The economy of these nomads entirely depends on fragile and meagre natural resources associated with inconsistent rain pattern. Job opportunities are confined to labour in agricultural fields or other minor activities due to lack of education or skilled training. Most of the nomads live below poverty line in the absence of basic human needs like clean drinking water or suffi- cient food, health and education for their children. Livestock breeding, improvement of performance or range management is not practised scientifically. Few nomads manage to shift milk- producing animals near the roadside to earn some living. Livestock is used for exchange of gifts in communal ceremonies like weddings, tribal celebrations, child- birthorfunerals,andanimalsareslaughter- ed for feast for the guests. A nomad’s status in the desert life-style is chiefly deter- mined by the size of herd. All livestock are indigenous breeds well adapted to climat- ic conditions. Herd reproductive perform- ance is generally poor with low birth rates and high mortality due to poor nutrition, lack of healthcare and climatic stresses [1, 3, 11]. Veterinary health care centres are not available to majority of livestock. All the nomads have an unwritten code of ethics for their range territories and use of water points: tobas, kunds, or wells. Each clan has access to traditionally-defined ter- ritories to use for grazing irrespective of the condition of the range vegetation. The exhaustive use of range vegetation has resulted in the decline of palatable species especially grasses, overall vegetation cover, malnutrition of livestock and degrad- ing wildlife habitats. Range vegetation Phytogeographically, the vegetation of the Cholistan desert belongs to the Nubo- Sindhian Province of the Sudanian Region and is typical of arid regions [1]. ThesparsevegetationofCholistanisdomin- ated by bush forming perennial shrubs with scattered small trees. A large number of annual and ephemeral species appear after rains, complete their life cycle in a short span and dry up after dispersing seeds. Most of the shrubs and trees are leafless or have reduced and modified leaflike structures. Several members of Chenopodiaceae like Haloxylon, Suaeda, Salsola form the dominant species along with the hardy grasses of rangelands. Most of the species have a remarkable capacity to regenerate even in minimum rainfall. The seeds are also hardy enough to withstand severe high temperature, arid- ity and multiple years of drought. Rainfall is generally confined to few showers in the months of July to September. The rainfall ranging from 100-300 makes the desert green with vegetation cover up to 60% in some areas. The vegetation of Lesser Cholistan is domin- ated by several species of shrubs and perennial grasses dotted by few species of small trees. However, in Greater Cholistan trees are generally lacking, the shrubs are sparser and species diversity is poor. Vegetation cover is poor on sand dunes and unstable sand dunes are devoid of any vegetation. Some interdunal areas Stay at settlements in irrigated areas Stay at semi-permanent settlements Stay in the desert at the tobas W i n t e r P o s t - M o n s o o n S u m m e r Spring D E C Figure 3. Yearly movement pattern of pastoralists in Cholistan. 212 SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 4. have a better vegetation depending on the water retention capacity of the soil. Ground water table is very low and in most of the cases not fit for drinking. Main vegetation types It is difficult to define distinct community types associated with habitats like sand dunes, interdunal flat plains, saline plains or dry water-courses flooded periodically during rain. Some species are typically associated with a distinct habitat type. Some main vegetation types based on floristically dominant and structurally important species are described below. • Haloxylon-Calligonum Community This is a widespread community type dominated by leafless shrubs of Haloxylon stocksii and Calligonum polygonoides. Other associated species include Lasiurus scindicus, Cymbopogon, jwarancusa, Suaeda fruticosa, Dipterygium glaucum, Crotalariaburhia,etc.Thistypeofcommun- ities generally covers flat plains with com- pact soil and saline conditions. Several species of annuals and ephemerals emerge after rains, covering the ground but contributing very little biomass. The shrubs have strong root systems and hold the soil together protecting the soil from wind erosion. Sometimes small mounds of soil are developed around the bushes while the surrounding soil is blown away with strong winds. • Acacia-Tamarix Community The Acacia nilotica and Tamarix aphylla community is found in peripheral areas of Cholistan. Acacia nilotica is locally considered as poor man’s teak wood for its valuable timber used for cheap furniture and agriculture implements. Leaves provide forage for livestock. Tamarix bran- ches are used for thatching rooftops and fences around livestock enclosures. Trees also provide shade for resting livestock seeking shelter from strong sun. Several species of grasses and herbs are associat- ed with these communities like Cenchrus, Pennisetum, Eragrostis, Fagonia, Diptery- gium, Indigofera, etc. Large hemispherical clumps of Heliotropium crispum are com- mon in the peripheral regions of the desert which hold soil from erosion and conserve water. • Prosopis-Salvadora Community The Prosopis cineraria and Salvadora oleoides community is generally found in Lesser Cholistan regions where rainfall is not less than 100mm. Prosopis cineraria is considered as an excellent forage species with a very deep root system, very well adapted to the desert conditions. The branches are extensively lopped for feed- ing young lambs and goats not able to roam around for forage. Generally the top shoots are left for future growth and lower branches are cut for livestock. It also forms excellent firewood for cooking as well as charcoal making. Salvadora oleoides is an evergreen tree with deformed trunk and a deep root system heavily grazed by camels and goats. It is a very slow- growing tree with poor regeneration through seeds but it profusely produces suckers forming large clumps. Sometimes only clump-forming large bushy structures are seen due to heavy browsing; tree-like shape is rare; ripe fruits are also edible. Other associated species include Acacia nilotica, Zizyphus nummularia, Tamarix aphylla, grass species like Lasiurus, Cym- bopogon, Sporobolus, Panicum, Cen- chrus,Aristida,etc.Someannualsassociat- ed with this community include Zygophyllum, Dipterygium, Indigofera, Glinus, Fagonia, Heliotropium, etc. • Lasiurus-Cymbopogon Community Perennial species of clump forming grass species like Lasiurus scindicus and Cymbo- pogon jwarancusa are developed on poo- rer soils on flat plains with compact soil. This community type is generally poor in species diversity, having mainly annual short-lived species appearing after rains. These grass species are palatable in early stage of growth; coarse hard old leaves are not generally grazed. Lasiurus scindicus and Cymbopogon jwa- rancusa are abundantly distributed throu- ghout the Cholistan Desert. Scattered shrubs of Haloxylon, Suaeda, Crotalaria, Aerua are also sparsely found. • Acacia-Pennisetum Community Acacia jacquemontii and Pennisetum divi- sum are generally associated with dry water courses with better soil and moisture retention capacity. Acacia jacquemontii is atallshrubbyspecieswithextensivebranch- ing from the base. It is a good forage species and a preferred firewood shrub in the desert. Pennisetum divisum is a tall large clump-forming grass with a good soil-binding ability preventing soil erosion from strong winds and flash floods. This is an excellent fodder grass relished by live- stock producing green tender leaves in minimum moisture conditions. Other associated species include species of Convolvulus, Launnaea, Indigofera, Tri- bulus, Trianthema, Neurada, etc. Several grass species include Ochthochloa, Pani- cum, Aeluropus, Stipagrostis, Cenchrus, etc. • Aerua-Crotalaria Community This community is common on poor saline soils. Usually, large shrubs and trees are absent. Grasses like Sporobolus, Ochtho- chloa, Desmostachya species are found having stiff leaves, grazed when young. They are usually found in interdunal saline sandy soils. • Capparis-Suaeda Community Capparis aphylla and Suaeda fruticosa are also widespread in poor soils with inadequate moisture contents. Capparis aphylla, a leafless shrub, sometimes attains tree-like form; its branches are used for thatching roof tops. Suaeda fruticosa forms evergreen large-sized bushy dome- like clumps; leaves are reduced to scales. Vegetation is very sparse, poor in species diversity; short-lived annual plants appear after rains for a short period. • Leptadenia-Calotropis Community The Leptadenia pyrotechnica and Calotro- pis procera community develops in poor sandy soils; not palatable for livestock, it spreads in areas with heavy grazing pres- sure. Leptadenia pyrotechnica is a much branched leafless shrub not grazed by livestock but the branches are used for thatching roofs and fencing. Cyperus conglomerates is a compact clump- forming sedge with roots having good sand-binding ability, found frequently on sand dunes under poor moisture condi- tions, grazed when young. Grazing potential of ranges The vegetation in Cholistan is typical of arid and semiarid climate consisting of xerophytic species adapted to high season- al temperatures, low humidity, moisture fluctuations and wide variety of edaphic conditions [12]. Compared to the hyper- arid southern region, the vegetation cover is relatively better in Eastern Cholistan (200mm rainfall zone). A wide range of nutritious and drought-tolerant plant spe- cies (128 species belonging to 33 fami- lies) of grasses, herbs, shrubs and trees occupy this desert [9]. Hardy species adapted to high temperature and low soil moisture include Lasiurus scindicus, Sporo- bolus iocladus, Aeluropus lagopoides and Cyperus conglomerates. Major forage plants and corresponding characteristics, use and grazing animals in the Cholistan Desert are listed in table 1. Pastoralism in Cholistan Pastoral The livestock depend entirely on self- regenerating and self-maintaining range resources throughout the year or resort to canal and riverain areas for grazing with SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 213 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 5. Table I. Major forage plants of the Cholistan Desert and corresponding characteristics, use, and grazing animals. Plant Species Parts used Palatability Season Frequency Grazing animals Perennial Grasses Stipagrostis plumosa Lv, Sh HP Monsoon, Spring V. Common Cattle, sheep, goat Cenchrus ciliaris Lv, Sh HP Monsoon, Spring Rare Cattle, Sheep Goat Cymbopogon jwarancusa Lv, Sh LP Monsoon, Spring V. Common Sheep, Cattle Ochthochloa compressa Lv, Sh HP Monsoon, Spring V. Common Cattle, Sheep Goat Lasiurus scindicus Lv, Sh HP Monsoon, Spring V. Common Cattle, Sheep Goat Panicum antidotale Lv MP Monsoon, Spring Rare Cattle, Sheep Goat, Camel Panicum turgidum Lv HP Monsoon, Spring Common Cattle, Sheep Goat, Camel Sporobolus iocladus Lv MP Monsoon, Spring Common Cattle, Sheep Goat, Camel Aeluropus lagopoides Lv, Sh H.P Monsoon, Spring Common Cattle, Sheep Goat, Camel Sedge Cyperus conglomeratus Sh, Lv HP Monsoon, Spring, Winter Common Cattle, Sheep Goat, Camel Annual Grasses Aristida adscensionis Sh, Lv MP Monsoon Common Sheep, Goat Aristida mutabilis Sh, Lv MP Monsoon Common Sheep, Goat Aristida funiculata Sh, Lv MP Monsoon Common Sheep, Goat Aristida hystricula Sh LP Monsoon Common Sheep, Goat Cenchrus biflorus Sh, Lv MP Monsoon Common Cattle, Sheep, Goat Cenchrus prieurii Sh, Lv HP Monsoon Common Cattle, Sheep, Goat Enneapogon desvauxii Lv, Sh HP Monsoon Rare Sheep, Goat Eragrostis barrelieri Lv, Sh HP Monsoon Common Sheep, Goat Leptothrium senegalense Lv, Sh MP Monsoon Common Cattle, Sheep, Goat Tragus racemosus Lv, Sh MP Monsoon Common Cattle, Sheep, Goat Forbs Tribulus longipetalus Sh, Lv, Fl HP Monsoon Common Cattle, Sheep Goat, Camel Tribulus terrestris Sh, Lv, Fl HP Monsoon, Winter, Spring Rare Cattle, Sheep Goat, Camel Fagonia cretica Lv LP Monsoon, Winter, Spring Common Sheep Anticharis linearis Lv, Sh LP Monsoon Common Sheep, Goat Indigofera sessiliflora Lv, Sh, Fl MP Monsoon Rare Sheep, Goat Indigofera argentea Lv, Sh, Fl MP Monsoon Common Sheep, Goat Indigofera hochstetri Lv, Sh, Fl MP Monsoon Common Sheep, Goat Neurada procumbense Lv, Fl MP Winter Rare Sheep, Goat Glinus lotoides Lv, Sh, Fl LP Monsoon, Spring Common Cattle, Sheep Goat, Camel Euphorbia prostrata Lv LP Monsoon, Spring Common Sheep, Goat Mukia madraspatana Lv HP Monsoon Common Sheep, Goat Cressa cretica Lv, Sh MP Monsoon, Spring Common Cattle, Sheep, Goat Convolvulus microphyllus Lv, Sh MP Monsoon, Spring Common Sheep, Goat, Camel Launea residifolia Lv, Sh LP Winter Common Sheep, Goat, Camel Launea nudicaulis Lv, Sh LP Winter Common Sheep, Goat, Camel Dipterygium glaucum Sh LP Monsoon, Spring V. Common Camel Cleome brachycarpa Sh LP Monsoon, Spring Common Camel Farsetia hamiltonii Sh LP Monsoon, Spring V. Common Sheep, Goat Heliotropium strigosum Lv, Sh MP Monsoon, Spring Common Sheep, Goat, Camel Heliotropium crispum Lv, Sh LP Throughout the year Common Camel, Sheep Trianthema portulacastrum Sh MP Monsoon Common Camel, Sheep Zaleya pentendra Sh MP Monsoon Common Camel, Sheep Sesuvium sesuvioides Sh MP Monsoon Common Camel, Sheep Limeum indicum Sh MP Monsoon V. Common Camel, Sheep Shrubs Crotalaria burhia Lv LP Throughout the year Common Camel Aerva persica Lv LP Throughout the year Common Camel Calotropis procera Lv LP Throughout the year Common Cattle, Sheep, Goat Pulicaria rajputanae Lv LP Throughout the year Common Camel 214 SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 6. or without purchased fodder for sup- plementation in the pre-monsoon period. Agropastoral Along with range resources, the livestock also utilize fodder crops from agricultural farms owned by the pastoralists or by tenant farmers in the irrigated tract. The croplands provide food security for both the livestock and their owners. The impor- tance of crops varies according to the status of small owner-pastoralists depend- ing mainly on free grazing along the canals or roadsides or on additional pur- chased fodder to support livestock in the pre-monsoon period. After the monsoon, when sufficient forage is available, the livestock move to desert rangelands for grazing. Sedentary pastoralists Livestock owners living in permanent settle- ments working as tenant farmers or small farmers generally use nearby grazing areas. The livestock comprising mainly buffaloes, cattle, some sheep and goats confined to the irrigated fields are mainly stall-fed or use surrounding areas for graz- ing and come back in the evening. The feed resources comprise a variable pro- portion of crop residues, fodders and peri- irrigation grazing areas. Due to the com- petition from the transhumant livestock, feed supply during the pre-monsoon period may be constrained. Transhumant The majority of livestock and people seasonally move to the desert and sub- sequently again towards irrigated mar- gins. The point of origin is irrigated areas, and in the desert they have fairly well- defined range territory for various clans; however, variation in the route and the sequence of use of tobas are governed by the water and forage quantity available in the range. The livestock start moving to the desert at the onset of monsoons in July/August when the tobas are filled with water. They stay around the tobas as long as water is available, and move to the wells or kunds when the tobas can no longer support the livestock or people. The salinity in the wells rises with the advancement of dry season. The accessible forage is exhausted till March/April and the salinity in the wells renders the water unsuitable for cattle. Livestock start to return towards the irrigat- ed margins. Only camels and goats are left to survive on evergreen browse. The precise times of movements depend on the rains and the resulting availability of water and forage in the desert; in good years water may be available in tobas until January or February, and in normal years it may dry out before that. Desert nomads The livestock remain all the year in the desert and are dependent upon the range resources for fodder and water. Almost all the camels, about 50% goats and a small number of cattle are represented in the system. The animals move with the trans- humant livestock from one toba to the next and subsequently to the wells and/or kunds. However, they remain at the wells or kunds when the transhumant livestock retreat to the irrigated margins. Generally, each clan/group tends to remain at a particular well or kund until the tobas are filled again. Livestock Breeds Camels, cattle, sheep and goats are the predominant types of livestock. Cattle are the most precious livestock species of pastoral lifestyle followed by sheep, goats and camels. The productivity (birth rates) among cattle is the lowest. Lambing occurs mainly during the months of January/ ebruary. Goats are kept in smaller num- bers due to: i) lack of browse when ani- mals are taken to irrigated areas during droughts; ii) to avoid predation by jackals; and iii) because of difficulties encountered in controlled grazing in irrigated areas. Camels are also owned mainly for trans- portation purposes. Notable breeds of livestock in Cholistan are as follows: Cattle: Cholistani and Hasari Goat: Jattal (Cholistan goat) Sheep: Buchi, khadali, sipli Camel: Marecha and Brella Generally, split-herding prevails in cattle and small ruminants. Small calves in the case of cows and lambs and kids in case of sheep and goats are kept at pens near the tobas, while the adults are allowed to roam around for grazing in the open range areas of the desert. Most cattle and camel are not herded. In the case of the cattle, the animals are driven out of the house to graze freely and they return home by themselves in the evening. The camels, however, remain in the desert unherded Table I (cont’d). Major forage plants of the Cholistan Desert and corresponding characteristics, use, and grazing animals. Plant Species Parts used Palatability Season Frequency Grazing animals Calligonum polygonoides Sh, Fl MP Throughout the year V. Common Camel, Cattle, Goat Acacia jacquemontii Lv, Sh, Fl HP Throughout the year Common Sheep, Goat, Camel Haloxylon recurvum Sh MP Throughout the, year V. Common Camel Haloxylon salicornicum Sh LP Throughout the year V. Common Camel Salsola baryosma Sh MP Throughout the year V. Common Camel Suaeda fruticosa Sh, Lv MP Throughout the Year V. Common Camel Capparis decidua Sh MP Throughout the Year V. Common Camel, Cow Zizyphus nummularia Lv HP Throughout the Year Common Sheep, Goat, Cow, Camel Leptadenia pyrotechnica Sh NP Throughout the Year Common Camel Trees Prosopis cineraria Lv, Sh, Fl, Fr HP Throughout the year V. Common Sheep, Goat, Cow, Camel Acacia nilotica Lv, Sh, Fl, Fr HP Throughout the year Common Sheep, Goat, Cow, Camel Prosopis juliflora Lv, Fr MP Throughout the year Rare Sheep, Goat, Camel Zizyphus spina christi Lv HP Throughout the year Rare Sheep, Goat, Cow, Camel Tamarix aphylla Sh MP Throughout the year Common Camel Salvadora oleoides Lv, Sh HP Throughout the year Rare Sheep, Goat, Cow, Camel Lv = leaves; Sh = shoot/stem; Fl = flower; Fr. = fruit; HP = highly palatable; MP = moderately palatable; LP = less palatable; NP = non palatable. SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 215 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 7. for months. Small ruminants (sheep and goats) are attended by herdsman mainly to avoid predation by jackals. Mortality rate is high among cattle alone ranges from 5 to 60 percent [1]. The main causes of mortality are related to droughts, poor nutrition due to shortage of feed and water and diseases caused by nutrition stress. No livestock health facilities are available in the desert, only limited health services are available in peripheral small towns and vaccination of livestock is not practised regularly. Livestock production and availability of feed The rangelands in Cholistan desert are monsoonal and forage production in these rangelands depends heavily on monsoon rains (amount, time, and frequency). The health of the animals, their production, birth and mortality rate relates directly to the availability of vegetation. During norm- al rainfall years (100–200mm) green her- baceous production remains critically low, particularly for cattle and sheep and ani- mals remain unnourished. Once or twice in ten years forage production is sufficient for year-round grazing of the livestock. During good years the production of live- stock and by-products is sufficient enough when owners have some earnings. After successive drought years the forage pro- duction declines drastically and vegeta- tion growth is practically nil. The grazing period starts from August till February during good rainy years. Both the composition and production of browse and herbaceous forage vary with the range types given the same amount of rainfall. In the best ranges, herbaceous forage yields 60-70% whereas browse yields 30-40% of the total forage product- ion. In moderate ranges, herbage product- ion varies from 35 to 45% and browse from 55 to 65%. In poor ranges, herbage yields less than 20% and browse more than 80% [1]. Almost all the herbaceous forage is utilized during monsoon and post-monsoon season, while green browse is available in the area throughout the year. The herbaceous vegetation is very short-lived and it dries up when the soil moisture is no longer available. The feed requirements of camels and goats, in the winter and pre-monsoon are provided by green browse (Calligonum and Haloxy- lon), while cattle and sheep suffer severe nutritional deficiencies. The winter rains are very scarce so spring vegetation is generally poor, represented by few annual species of grasses and forbs providing very little biomass for grazing. At present, the total livestock population in Cholistan is around 134,798 animal units (AU) (table 2). The animal unit is consider- ed as an adult cow weighing 350 kg (400 kg at international level) and consuming 7 kg dry matter forage per day in Cholistani conditions. The total annual dry matter forage demand in Cholistan is .34 million tons while the available dry matter forage is .12 mil- lion tons. Thus there is an annual short- age of 0.22 million tons dry matter forage (table 3). It has been noticed that at the moment, some 88,655 AU are in excess of the present carrying capacity [13]. The rangelands of Cholistan are under severe threat of degradation because of overgrazing and harsh climat- ic conditions. Due to continuous grazing, the desirable palatable species are vanishing at an alarming rate and relative- ly unpalatable species are spreading and dominating the landscape. The severity of the problem can be seen from the fact that such highly desirable grass species as Pennisetum divisum and Cenchrus setigerus have disappeared from most part of the Cholistan desert rangelands. Livestock production and health management Three inter-related aspects of animal health i.e., feed, water and disease have been encountered in Cholistan desert. Deficiencies in the availability of forage (quantity and quality), drinking water (saline or polluted) and free mixing of diseased animals with healthy ones during grazing expose livestock to various types of diseases. Veterinary health centres or hospitals are not available towards the interior of the desert and very few poorly- equipped small units are available in peri- pheral cities. Livestock owners often become distressed and helpless when their livestock, particularly cattle fall seriously ill. Some traditional homemade herbal preparations are used to treat sick ani- mals. The mortality rate is very high and drought conditions increase stress due to malnutrition and lack of water causing various diseases. The most common diseases occurring in various livestock species are as follows: – large ruminants: haemorrhagic septicae- mia, black quarter, foot and mouth disease, anthrax, mange, surra, camel pox, endo- and ectoparasites, etc.; – small ruminants: enterotoxemia, pleuro- pneumonia, sheep and goat pox, anthrax, liver fluke, endo- and ectoparasites, etc. The amount of livestock population that have died and suffered from various diseases in Cholistan desert is given in table 4. Traditionally, reasonable disease diagnostic know-how is avail- able at the toba level. However, disease treatment capabilities of the herders are very limited and medicines are avail- able at places that are far away from their tobas. Therefore, a large number of Table II. Livestock population and forage demand in the Cholistan Desert (census 1996). Animal type Population Animal Unit (AU) Dry matter intake per day (kg) Annual forage demand (tons) Cattle 73,048 7,3048 7 186,638 Sheep 14,7429 21,061 1 53,812 Goats 53,680 7,669 1 19,593 Camels 16,510 33,020 14 84,366 Total 290,667 134,798 10 344,409 Table III. Annual demand and supply of dry matter forage in Lesser and Greater Cholistan. Livestock population (AU) Annual forage demand (tons) Forage available tons Deficit (tons) Lesser (1,237,000 ha) Greater (1,369,000 ha) Total (tons) (2,606,000 ha) 134,798 344,409 55,962 61,934 117,896 226,513 216 SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.
  • 8. the animals infected with disease were either not provided any treatment or were given indigenous treatment. Livestock marketing The pastoral nomads in Cholistan desert are living very much below the poverty line due to the lack of education or of skilled labour and to their entire dependence on meagre natural resources. The quantity and quality of livestock is not good enough to support a family. Marketing is generally practised through a middleman (locally called Beopari) who collects animals from various nomads camping sites near tobas on the outskirts of small towns. Seasonal, monthly, and weekly livestock markets operate at various levels depending on the availability of animals for sale. Livestock by-products including homemade mats prepared from raw wool are also brought to the markets. Camel bone and camel skin products prepared with primitive skills are also prepared on a small scale. But due to the lack of training or of refined skills, the market is very limited. The money earned through sale of livestock is not enough to meet the requirements for food or for animal feed and fodder. Constraints The major constraint in livestock produc- tion in Cholistan Desert is the shortage of sweet water. This is compounded by the prolonged droughts of many years when toba water dried out completely. In Greater Cholistan, feed for livestock is still available, but the toba water is deplet- ed and the thirsty herds are forced to migrate towards semi-permanent settle- ments where well water is adequate but of poor and saline quality not fit for drinking. The wells are unlined and must be re-dug each season. On the other hand in the western part (Lesser Cholistan) the quantities of both water and feed are inadequate. Landless pastoralists suffer due to the scarc- ity of rangelands for grazing in the irrigat- ed fringes where they work as poorly-paid labour or as tenant farmers on farmlands generally used for agricultural crops. The combination of long distances travel- led by the livestock in search of forage, harsh temperature rising above 50o C, in- adequacy of feed, undernourishment and highly saline drinking water from wells, all contribute to high mortality rates. â–  References 1. Food and Agriculture Organization (FAO). Pakistan – Cholistan Area Development Project. Report No. 59/53 ADB-PAK 58 (Final version). Rome: FAO, 1993. 2. Akbar G, Arshad M. Developing sustainable strategies for Cholistan desert: opportunities and perspectives. Science Vision 2000; 5: 77-85. 3. Akbar G, Khan TN, Arshad M. Cholistan desert, Pakistan. Rangelands 1996; 18: 124-8. 4. Chaudhry AA, Hussain A, Hameed M, Ahmad R. Biodivesity in Cholistan desert, Pun- jab, Pakistan. In: Shazad AM, Charles AW, Usman AS, eds. Biodiversity of Pakistan. Islama- bad (Pakistan): Pakistan Museum of Natural His- tory, 1997. 5. Mughal MR. Ancient Cholistan – archaeology and architecture. Lahore (Pakistan): Ferozsons (Pvt.) Ltd., 1997. 6. Arshad M, Akbar G, Rashid S. Wealth of medicinal plants of Cholistan desert, Pakistan: conservational strategies. Hamdarad Medicus 2003; XLV: 25-34. 7. Baig MS, Akram M, Hassan MA. Possibilities for range development in Cholistan desert as reflected by its physiography and soils. Pak J For 1980; 30: 61-71. 8. Rao AR, Arshad M. Perennial grasses of Cho- listan desert and their distribution. Proceedings of a seminar on “Peoples participation in the mana- gement of resources in arid lands. Bahawalpur (Pakistan): Cholistan Institute of Desert Studies, Islamia University, 1991. 9. Arshad M. Rao AR. Flora of Cholistan desert (Systematic list of trees, shrubs and herbs). J Econ Tax Bot 1994; 18: 615-25. 10. Arshad M, Rao AR, Akbar G. Masters of disaster in Cholistan desert, Pakistan: pattern of nomadic migration. UNEP Desertification Control Bulletin 1999; 35: 33-9. 11. Mumtaz KK. Habitat and desert: The case of Cholistan. In: Taylor BB, ed. The changing rural habitat; Volume 1: Case Studies. Singapore: The Concept Media/Agha Khan Award for Architec- ture, 1982. 12. Rao AR, Arshad M, Shafiq M. Perennial grass germplasm of Cholistan desert and their phytosociology. Bahawalpur (Pakistan): Cholis- tan Institute of Desert Studies, Islamia University, 1989. 13. Akbar G, Arshad M. Feasibility of the resto- ration of vegetation in the Cholistan desert as a natural habitat of Houbara bustard and other wild life species. Final Project Report. Lahore (Pakistan): World Wild Fund (WWF)-Pakistan, 1999. 14. Iqbal M, Farooq U, Basir A, Khan NA, Malik SZ. In: A baseline survey for the develop- ment of livestock sector in Cholistan. GmdII, GTZ P.N.91.2123.7. Lahore: Pak-German Technical Cooperation (Deutsche Gesellschaft fur Technis- che Zusammenarbeit, GTZ), Livestock and Dairy Development Department, 2000. Table IV. Death and disease incidence among livestock population. Items Cattle Camel Sheep Goat Farms reported disease incidence 49.69 5.45 59.02 34.12 Population suffered 5.64 1.65 12.19 19.37 Mortality 2.72 0.55 9.31 7.78 Source: [14]. SĂ©cheresse vol. 17, n° 1-2, janvier-juin 2006 217 Copyright © 2017 John Libbey Eurotext. TĂ©lĂ©chargĂ© par un utilisateur anonyme le 22/02/2017.