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Land Plants
Clubmosses
and friends
Spore-dispersed
Euphyllophytes
Seed Plants
megaphylls
Equisetophytes
We will be looking at 2 orders today:
•Equisetales
•Sphenophyllales
Equisetophyte features:
•Creeping rhizomes and aerial stems
•Leaves (& branches) in whorls at nodes
•Jointed stems with prominent ridges
•Silica impregnated epidermis
•Sporangia arranged in strobili
Equisetales, Equisetum
Equisetum strobilus with
whorls of hexagonal
units called
sporangiophores
• photosynthetic
• spherical in shape
• with elaters -- ribbon-like
extentions of the spore coat
Equisetum
spores
Equisetum gametophytes tend to be
unisexual
Archegonia
Archegoniate
gametophyte
Antheridiate
gametophyte
Equisetum stem anatomy
Anatomy of the Equisetum
internode
An Equisetum vascular bundle
Equisetum strobilus anatomy
Spores
Spores Terminal flange
Equisetales: Calamites, a fossil horsetail from the Carboniferous
We will look at three form genera in the composite genus, Calamites:
Annularia
Asterophyllites
Calamites
Sphenophyllales:
Sphenophyllum

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Equisetopsida Botany - good presentation