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          “In short, the animal and vegetable lines,
        diverging widely above, join below in a loop.”
                                                 loop.”
               —Asa Gray (Brainyquote.com)




       Ecosystems and Biomes
• Two organizing principles
  are ecosystem and biome.
• Ecosystem: A Concept for
  All Scales
   – Ecosystem —the totality of
     interactions among organisms
     and the environment in the
     area of consideration.
       • Encompasses both the living
         and nonliving portion and how
         energy flows among them.
       • Weakness—there is an
         almost infinite variety in the
         magnitude of ecosystems that
         can be studied:
   – Range includes whole
     Earth itself to drop of water.




       Ecosystems and Biomes
                                          • Biome: A Scale for All
                                            Biogeographers
                                            – Biome—a large,
                                              recognizable
                                              assemblage of plants
                                              and animals in
                                              functional interaction
                                              with its environment.
                                            – Most appropriate scale
                                              for understanding
                                              world distribution
                                              patterns.




                                                                       1
Ecosystems and Biomes
• Eleven major types
   – Often significant and even
     predictable relationships
     exist between the biota
     (particularly the flora) of a
     biome and the associated
     climate and soil types.
• Ecotone—the transition
  zone between biotic
  communities in which the
  typical species of one
  community intermingle or
  interdigitate with those of
                                       Marsh/mangrove ecotone with fire, freeze,
  another.                             and hurricane damage. Taken September
                                       20, 2001




                  Terrestrial Flora
• Geographers interested in
  natural vegetation of
  landscape for three
  reasons:
   1. Plants are likely to dominate
      a landscape (except where
      terrain is rugged, climate is
      harsh, or humans have
      intervened);
   2. Vegetation is a sensitive
      indicator of other
      environmental attributes;
   3. Vegetation is often
      instrumental to human
      settlement and activities.




        Characteristics of Plants
• Most very hardy.
• High survival potential
  dependent on:
   – Subsurface root system
   – Reproductive mechanism
       • Perennial—plant that can
         live more than a single
         year despite seasonal
         climatic variations.
       • Annual—plant that
         perishes during times of
         climatic stress but leaves
         behind a reservoir of seeds
         to germinate during the
         next favorable period.




                                                                                   2
Characteristics of Plants
• Common
  characteristics:
   – Roots (to gather
     nutrients and moisture
     and to anchor plant);
   – Stems and branches
     (to support and
     transport nutrients);
   – Leaves (to collect solar
     energy, exchange
     gases, and transpire
     water);
   – Reproductive organs.




           Floristic Terminology
• Categorizing by
  reproduction:
   – Through spores
                                            Spores
      • Those that reproduce by
        spores are in two major
        groups:
          – Bryophytes—spore-
            bearing plants such as
            mosses and liverworts;
            never dominated in
            history, but can be very
                                                     Bryophytes
            important in some
            localized situations.
          – Pteridophytes—spore-
            bearing plants such as
            ferns, horsetails, and
            clubmosses; used to
            dominate continental
            vegetation, but no more.
                                                 Pteridophytes




           Floristic Terminology
   – Through seeds                           Seeds
      • Those that reproduce by
        seeds are in two major
        groups:                                      Gymnosperms
          – Gymnosperms—seed-
            reproducing plants that carry
            their seeds in cones; also               Angiosperms
            known as conifers.
               » Used to be more
                   important, in geologic
                   past.
          – Angiosperms—plants that
            have seeds encased in
            some sort of protective
            body, such as a fruit, a nut,
            or a seedpod.
               » Have dominated
                   planet vegetation for
                   last 50 million to 60
                   million years.




                                                                   3
– Other terms
                   Trees

                          Gymnosperms

                             Softwood
                             Coniferous
                             Needleleaf


                           Angiosperms

                             Hardwood
                             Deciduous
                             Broadleaf
                                                – Fig. 11-3




                Floristic Terminology
•   Categorizing by stem or trunk
    composition:
     – Woody plant—plant that has stem
       composed of hard fibrous material;
       refers mostly to trees and shrubs.
     – Herbaceous—refers to plants that
       have soft stems; mostly grasses,
       forbs, and lichens.
•   Categorizing by leaf retention:
     – Deciduous—refers to trees that
       experience an annual period in
       which all leaves die and usually fall
       from the tree, due either to a cold or
       dry season.
     – Evergreen—a tree or shrub that
       sheds its leaves on a sporadic or
       successive basis, but at any given
       time appears to be fully leaved.




                Floristic Terminology
• Categorizing by leaf
  shape:
     – Broadleaf—tree that has
       flat and expansive leaves.
          • Majority are deciduous.
          • In rainy tropics, everything
            is evergreen.
     – Needleleaf—refers to trees
       adorned with thin slivers of
       tough, leathery, waxy
       needles rather than typical
       leaves.
          • Almost all are evergreen.




                                                              4
Floristic Terminology

• Categorizing by
  supposed structure—but
  this is unsatisfactory.
   – Hardwood—angiosperm
     tree that is usually broad-
     leaved and deciduous.
     Wood has a relatively
     complicated structure, but
     is not always hard.
   – Softwood —gymnosperm
     tree; nearly all such trees
     are needle-leaved
     evergreens with wood of
     simple cellular structure but
     not always soft.




     Environmental Adaptations
• Two prominent adaptation strategies of
  plants to protect against environmental
  stress are
   – Xerophytic adaptations
   – Hygrophytic adaptations




     Environmental Adaptations
• Xerophytic—refers to
  plants structurally
  adapting to withstand
  protracted dry
  conditions.
   – Roots, stems, leaves,
     reproductive cycle can
     all adapt in various
     ways.
• Succulent—plant that
  has fleshy stem that
  stores water.




                                            5
Environmental Adaptations
• Hygrophytic—refers to
  plants structurally
  adapting to withstand
  protracted wet
  conditions.
• Hygrophytic
  Adapatation
   – Hygrophyte—plant that
     requires a saturated or
     semi-saturated
     environment (frequent
     soakings with water).




     Environmental Adaptations
• Likely to have extensive root
  system for anchoring in soft
  ground.
• Usually relies on buoyancy of
  water for support rather than
  stem.
• Many have weak, pliable stems
  so can withstand currents.
• Hydrophytes are often grouped in
  with this category.
   – Hydrophyte—a “water-loving” plant
     that is adapted to live in more or less
     permanently immersed in water.




     Environmental Adaptations
• Hygrophytic Adapatation
   – Hygrophyte—plant that requires a
     saturated or semi-saturated
     environment (frequent soakings with
     water).
   – Likely to have extensive root system
     for anchoring in soft ground.
   – Usually relies on buoyancy of water
     for support rather than stem.
   – Many have weak, pliable stems so
     can withstand currents.
• Hydrophytes are often grouped in
  with this category.
   – Hydrophyte—a “water-loving” plant
     that is adapted to live in more or less
     permanently immersed in water.




                                               6
The Critical Role of Competition
• Competition is key
  in which plants grow
  where.
   – Even though all
     conditions (climatic,
     edaphic, etc.) are
     favorable, a plant
                                       Kudzu Vine
     may not take hold in
     one area because of
     competition.




     Spatial Groupings of Plants
• Geographers usually
  more concerned with
  spatial groupings than
  individual plants.
   – Groups based on
     dominant members,
     dominant appearance,
     or both.




     Spatial Groupings of Plants
• Floristic pattern of Earth is
  impermanent.
• Change can be slow and
  orderly, as in lake infilling.
• Change can be abrupt and
  chaotic, as in wildfire.
   – Climax vegetation—a stable
     plant association of relatively
     constant composition that
     develops at the end of a long
     succession of changes.




                                                    7
Spatial Groupings of Plants
• Is an association in
  equilibrium with prevailing
  environmental conditions.
• Should persist until
  environmental
  disturbance/change
  occurs.
• Seral association—various
  stages leading up to
  climax vegetation.




    Spatial Groupings of Plants
• Geographers can face significant difficulties in
  recognizing spatial groupings.
• As one tries to identify patterns and recognize
  relationships, must make generalizations.
   – When associations are portrayed on maps,
     boundaries usually represent approximations.
   – Human interference plays a major role.
   – Because of human impact, climax vegetation is now
     the exception rather than rule.
• Maps often ignore human interference, so are
  actually maps of theoretical natural vegetation.




    Spatial Groupings of Plants
• Many ways to classify
  plant associations.
   – Geographers usually
     place emphasis on
     structure and
     appearance of
     dominant plants.
   – Major associations
     include forests,
     woodlands,
     shrublands,
     grasslands, deserts,
     tundra, and wetlands.




                                                         8
– Fig. 11-7




     Spatial Groupings of Plants
• Forest—an assemblage of trees growing closely together so that
  their individual leaf canopies generally overlap.
    – Likely to become climax association in any area where moisture is
      adequate and growing season isn’t very short.
• Woodland—tree-dominated association in which the trees are
  spaced more widely apart than those of forests and do not have
  interlacing canopies.
• Shrubland —plant association dominated by relatively short
  woody plants.
    – Wide latitudinal range but usually restricted to semiarid or arid areas.




     Spatial Groupings of Plants
• Grassland—plant association dominated by grasses and forbs.
    – Prominent types are savanna, prairie, and steppe.
    – Associated with semiarid and subhumid climates.
• Desert—actually a climate, not an association per se, but is typified
  by plants widely scattered on bare ground.
• Tundra—a complex mix of very low-growing plants, including
  grasses, forbs, dwarf shrubs, mosses, and lichens, but no trees.
    – Only in the perennially cold climates of high latitudes or high altitudes.




                                                                                   9
Spatial Groupings of Plants
• Wetland—landscape
  characterized by
  shallow, standing
  water all or most of the
  year, with vegetation
  rising above the water
  level.
     – Have much more
       limited geographic                        Carson River, NV
       extent than any other
       above associations.




       Spatial Groupings of Plants
•   Various plant associations will exist in
    relatively narrow zones when mountain
    slopes have significant elevational
    changes in short horizontal distances.
     – Vertical zonation—the horizontal layering
       of different plant associations on a
       mountainside or hillside.
         • Elevation changes are counterpart of
           latitude changes.
         • Treeline elevation varies with latitude.
         • Southern and Northern hemispheres
           experience different elevation–latitude
           relationship, with Southern Hemisphere
           having lower treelines.
         • Reason for discrepancy is not understood
           yet.
•   Can have significant local variations
    caused by a variety of local
    environmental conditions.




          – Vertical Zonation
              • Most apparent in mountains due to changes in
                elevations over short distances




                – Fig. 11-10




                                                                    10
Spatial Groupings of Plants
• Exposure to sunlight is often a critical determinant of
  vegetation composition.
   – Adret slope—Sun slope; slope where Sun’s rays arrive at a
     relatively direct angle.
       • Relatively hot and dry, and its vegetation is sparser and smaller
         than that on adjacent slopes with different exposures.
       • Likely to have species composition different from adjacent slopes.
   – Ubac slope—a slope where sunlight strikes at a low angle and
     hence is much less effective in heating and evaporating than on
     the adret slope, thus producing more luxuriant vegetation of a
     richer diversity.
       • Difference between adret and ubac decreases with increasing
         latitude.
       • Valley-bottom locations can have vegetation composition
         significantly different from slopes running to it.




      – Local Variations
           • Exposure to sunlight
                   – Mountainous landscapes




               – Fig. 11-13




       – Local Variation (continued)
           • Valley-bottom location




               – Fig. 11-14




                                                                              11
Spatial Groupings of Plants
• Riparian vegetation—
  streamside growth,
  particularly prominent
  in relatively dry
  regions, where stream
  courses may be lined
  with trees, although no
  other trees are to be
  found in the
  landscape.




                  Terrestrial Fauna

• Animals occur in much greater
  variety than plants over Earth.
• Animals, however, tend to be
  much less prominent than plants
  in the landscape.
• They tend to be secretive and
  inconspicuous.
• Also, environmental relationships
  are much less clearly evidenced
  by animals than plants.
• Their inconspicuousness makes it
  more difficult to study them, and
  their mobility had lead to greater
  environmental adaptability among
  them.




        Characteristics of Animals
•   Variety of animal life
    is so great that it is
    difficult to find many
    unifying
    characteristics.
    –    Two universal traits
         (though these aren’t
         always immediately
         recognizable):
         1. Mobility               The tree of life at Animal Kingdom
         2. Need to eat plants
            and/or other animals




                                                                        12
• Characteristics of Animals
         – Great diversity
         – Two universal features
              • Motile
              • Heterotrophs
                   – Consumers (incapable of manufacturing
                     food from air, water and sunlight like
                     plants do)




                   – Fig. 11-15, 16, 19a, 21, & 27




                    Kinds of Animals
•   Size and habits are not valid indicators of animal’s significance to
    geographic study.
•   Minute and seemingly inconsequential organisms can play important roles.
•   Examples are carriers of disease, providers of scarce nutrients.
•   More than 90% of all animal species are invertebrates (without backbones).
•   Arthropods most prominent (insects, spiders, centipedes, millipedes,
    crustaceans).
•   Five groups of vertebrates, those with backbone:
     – Fish, amphibians, reptiles, birds, and mammals.




     • Kinds of Animals
         – Invertebrates
         – Vertebrates
              •   Fishes
              •   Amphibians
              •   Reptiles
              •   Birds
              •   Mammals




             – Fig. 11-15, 16, 19a, 21, 27 & 11-17




                                                                                 13
Kinds of Animals
• Most mammals are
  placentals, having
  young grow and develop
  in mother’s body.
     – About 135 species are
       marsupials, in which
       mothers carry young, not
       fully developed at birth, in
       pouches.
     – Two species are
       monotremes—lay eggs.
        • Echidna and duckbill
          platypus.




      Environmental Adaptations
•     Three different kinds of evolutionary
      adaptation by animals:
     1. Physiological
     2. Behavioral
     3. Reproductive




    Example of Animal Adaptations to
              Desert Life
• Faunal diversity can be
  astounding in desert areas
  where water is permanent or
  prolonged.
• Even in areas where open
  water is not available, there
  are pockets of localized
  favorable habitat that permit
  remnant populations to
  survive.
• Most animals are completely
  nocturnal.
• Animals are more conspicuous
  when cooler, such as at night
  and winter.
• Some animals follow rains in
  nomadic fashion.
• Most prominently displayed by
  birds.




                                              14
Example of Animal Adaptations to
              Desert Life
•   Some spend significant time
    underground.
•   Some bury selves to survive long dry
    spells, such as freshwater crayfish and
    crabs.
•   Text provides detailed discussion of
    anatomical and physiological
    adaptations.
•   A few species of rodents can exist from
    birth to death without ever taking a
    drink.
•   Get moisture from food.
•   Some species display ability to delay
    reproductive processes over long dry
    periods until more favorable conditions
    occur.
•   Australian desert kangaroos can delay
    implantation of fertilized blastocyst, so it
    remains in inactive state in uterus until
    better weather conditions occur.




     Competition Among Animals
• Competition can be
  both direct and
  indirect.
• Indirect—rivalry for
  space and resources.
• Direct—antagonism of
  predation.
• Many create social
  groups among own
  species.




     Competition Among Animals
• Some across species, such as
  communal relationship among
  zebras, wildebeest and impalas in
  East African savannas.
• Individual animals are concerned
  either largely or entirely with own
  survival.
• Some animal species concerned
  with survival of mates.
• Some concerned with survival of
  young (more common as maternal
  instinct, though some paternal too).
• Still fewer concerned with survival
  for group.




                                                   15
Cooperation among Animals
• Symbiosis—association of two
  dissimilar organisms, in which
  they live together in some
  fashion.
• Mutualism—symbiotic
  relationship in which the
  association is mutually
  beneficial to both organisms.
• Commensalism—symbiotic
  relationship in which the
  association is neither                         Symbiosis
  beneficial nor injurious to
  either.
• Parasitism—symbiotic
  relationship, in which the
  association benefits one, but
  harms the other; that is, one
  lives on or in the other, to     Mutualism   Commensalism   Parasitism
  detriment of the host.




        Zoogeographic Regions
• Animals’ distribution
  patterns more
  complex and irregular
  because of their
  mobility.
• The broad
  distributions of
  animals nevertheless
  do reflect a general
  distribution of energy
  and food diversity.




        Zoogeographic Regions
• Nine zoogeographic regions are generally
  recognized.
• Represent average conditions and cannot
  portray some common pattern in which
  different groups of animals fit precisely.
• Map on page 329.




                                                                           16
 Zoogeographic Regions
   • Reflective of the general distribution of energy
     and richness of food chemistry




               – Fig. 11-23




        Zoogeographic Regions
• Ethiopian Region
   – Has most diverse vertebrate
     fauna and greatest number
     of mammalian families.
• Oriental Region
   – Similar to Ethiopian but with
     less diversity (save for birds
     and reptiles; large number of
     venomous snakes).
• Palearctic Region
   – Poorer fauna than previous
     two;
   – Probably function of higher
     latitudes and more rigorous
     climate.




   • Palearctic Region
       – Separated from rest of Eurasia by mountains
       – Few endemic species, fewer species than in tropics




               – Fig. 11-23




                                                              17
• Nearctic Region
       – Non-tropical portions of North America
       – Similar to Palearctic due to Bering land bridge




                – Fig. 11-23




        Zoogeographic Regions
• Nearctic Region
   – Faunal assemblage relatively poor (save for
     being well-represented with reptiles).
   – Largely a transitional zone between
     Palearctic and Neotropical groups.
   – Great similarity to Palearctic, so that some
     group together into superregion, Holoarctic.
   – Reflects how faunal dispersal occurred via
     Bering land bridge in geologic past.
• Neotropical Region
   – Has rich and distinctive faunal assemblage:
   – Variety of habitats and isolation from other
     regions;
   – Has a larger number of endemic mammal
     families than any other region;
   – Bird fauna is exceedingly diverse and
     conspicuous.




                 Zoogeographic Regions

• Madagascar Region
   – Dominated by relic
     assemblage of unusual
     forms.
   – Primitive primates (lemurs).
• New Zealand region
   – Fauna dominated by birds
     (mostly flightless).
   – Almost no terrestrial
     vertebrates.
   – No mammals and only a
     few reptiles and
     amphibians.




                                                           18
Zoogeographic Regions
• Pacific Islands Region
    – Limited faunal assemblage.
• Australian Region
    –   Has most distinctive fauna of any region.
    –   Lack of variety is made up by animals’ uniqueness.
    –   The Unique Biota of Australia
    –   More than 90% of the native tree species in Australia are of the
        single genus, Eucalyptus.
         • Above should either be “a single genus, Euc” or “the single genus
           Euc” (no comma)
    – These trees, which there are greater than 400 species of, are
      native only to Australia.
    – The shrubs and bushes of Australia are dominated by a single
      genus, Acacia.
    – The dominance of this genus is attributed to isolation.




         – Australia Region
             • Australia and
               adjacent islands
             • Most distinctive
               fauna of any region
               due to the region’s
               lengthy isolation
                 – Few placental
                   mammals
             • Its unique biota are
               also primarily a
               result of isolation      – Fig. 11-17: Kangaroo. Fig. 11-18:
                                          Monotremes (egg-laying mammals)
                                          Echidna and duckbill platypus.




          Zoogeographic Regions
• Australian fauna is also unique
  because of isolation.
   – Australian fauna is dominated by a
     single primitive mammalian order,
     marsupials.
   – The continent is also the only home to
     the primitive monotremes.
   – Marsupials and monotremes were
     able to flourish in relative isolation
     from competitive and predatory
     pressures that influenced animal
     evolution in other parts of the world.
   – As such, indigenous placental
     mammals are lacking on the
     continent.




                                                                               19
The Major Biomes
• Named for dominant vegetation, but
  encompasses fauna as well as
  interrelationships with soil, climate, and
  topography.




 Major Biomes
  • Summary of each
    biome follows…
     – Distribution (map)
     – Climate types
     – Main vegetation
       types
     – Limiting factors to
       flora and fauna



                – Fig. 11-25
                  (left panel, p. 330)




           – Fig. 11-25 (right panel, p. 331)




                                                20
The Major Biomes
•   Tropical Rainforest
     – Selva — tropical rainforest; a
       distinctive assemblage of tropical
       vegetation that is dominated by a
       great variety of tall, high-crowned
       trees.
          • Probably most complex of all
            terrestrial ecosystems.
     – Distribution closely related to
       climate.
     – Consistent rainfall and relatively
       high temperatures.
     – Layered structure, with second
       layer being a branch canopy
       formed by the high trees that crest
       above the canopy.
     – Undergrowth relatively sparse
       because of lack of light.
     – Interior is region of heavy shade,
       high humidity, windless air,
       continuous warmth, aroma of
       mold and decomposition.
     – Fauna is largely arboreal—tree
       dwelling.
     – Canopy serves as principal food
       source.




                    The Major Biomes
• Tropical Deciduous Forest
     – Not closely correlated with
       specific climatic types;
       distribution more irregular and
       fragmented.
     – Compared to rainforest,
       canopy is less dense, trees
       are shorter, and there is less
       diversity of tree species (but
       greater variety of shrubs and
       other lesser plants).
     – Response to either less total
       precipitation or less periodic
       precipitation.
     – Many trees shed leaves at
       same time, so more sunlight
       can penetrate.
     – Produces classic jungle
       conditions.




                    The Major Biomes
• Tropical Scrub
     – Widespread in drier portions of
       A climatic realm (covers
       extensive areas in tropics and
       subtropics).
     – Dominated by low-growing
       scraggly trees and tall bushes,
       usually with extensive
       understory of grasses.
     – Plant species diversity less
       than that in tropical rainforest
       and tropical deciduous.
     – Faunal diversity very different
       from tropical rainforest and
       tropical deciduous.
     – Moderately rich assemblage of
       ground-dwelling mammals and
       reptiles, and of birds and
       insects.




                                             21
The Major Biomes
   •   Tropical Savanna
       – Distribution of biome doesn’t
         exactly correlate with distribution
         of tropical savanna climate.
       – Incomplete correlation most
         noticeable where seasonal rainfall
         contrasts are greatest (which is
         associated with annual shifting of
         the intertropical convergence zone
         [ITCZ]).
       – Dominated by tall grasses.
       – Some regions actually former
         tropical deciduous forest and even
         tropical rainforest, but humans
         converted it through fires and by
         grazing domestic animals.
       – Has a very pronounced seasonal
         rhythm: wet season, dry season,
         and wildfire season.
       – Savanna fauna varies according
         to continent.
       – Africa has most remarkable,
         diverse large wildlife.
       – Latin America has only sparse
         population of large wildlife.




                     The Major Biomes
• Desert
  – Occurs extensively in
    midlatitude locations in Asia,
    North America, and South
    America with a fairly close
    correlation to Bwh and Bwk
    climates.
  – Vegetation surprisingly
    variable.
  – Shrubs are typical, with
    succulents common in drier
    parts.
  – Trees can be found,
    particularly in Australia.
  – Most deserts have
    moderately diverse faunal
    assemblage.
  – Variety of large mammals is
    limited.




                     The Major Biomes
   •   Mediterranean Woodland and
       Shrub
       – Six widely scattered and relatively
         small areas in midlatitudes.
       – Have pronounced dry-summer
         wet-winter precipitation.
       – Dominant vegetation associations
         are physically similar, but
         taxonomically quite varied.
       – Dominated by dense growth of
         woody shrubs, but also have open
         grassy woodlands.
       – Plant species vary from region to
         region, but in all, the trees and
         shrubs are primarily broadleaf
         evergreens.
       – Many plants are adapted to rapid
         recovery after wildfire.
       – Fauna not particularly distinctive.
       – Seed-eating, burrowing rodents
         common.
       – General overlap of animals
         between this biome and adjacent
         ones.




                                               22
The Major Biomes
     •   Midlatitude Grassland
           – Locational coincidence between
             this biome and steppe climatic
             type is very pronounced in
             Northern Hemisphere.
           – Less distinct climatic correlations
             in Southern Hemisphere.
           – Occurs widely in midlatitudes of
             North America and Eurasia.
           – Low precipitation and/or frequency
             of fire prevent growth of tree or
             shrub seedlings.
           – Characteristics of grasses
             depends on moisture: taller in
             wetter area (prairie), shorter in
             dryer (steppe), and sometimes not
             continuous, but grow in discrete
             tufts.
           – Before human encroachment,
             fauna comprised of large numbers
             of relatively few species, with
             migratory larger herbivores.




                            The Major Biomes
     • Midlatitude Deciduous Forest
           – Used to be far more extensive
             in all Northern Hemisphere
             continents and to some extent
             in tracts in Southern
             Hemisphere.
           – Humans have cleared away
             large portions for agriculture.
           – Fairly dense growth of tall
             broadleaf trees with complete
             canopy in summer.
           – Winter very different, with
             seasonal fall of leaves.
           – Tree species vary greatly from
             region to region.
           – Generally has the richest
             assemblage of fauna in
             midlatitudes.
           – Seasonal variation to fauna
             (hibernation and migration).




                            The Major Biomes
•   Boreal Forest
     –   An extensive needleleaf forest in subarctic
         regions of North America and Eurasia; also
         called taiga.
     –   One of most extensive biomes, occupying
         vast expanse of northern North America and
         Eurasia.
     –   Close correlation with subarctic climatic type.
     –   Has perhaps simplest assemblage of plants.
     –   Most trees are conifers, though in some
         places deciduous trees interrupt the
         coniferous cover.
     –   Trees become spindlier, short, and openly
         spaced in north.
     –   Bogs and swamps numerous because of
         permanently frozen subsoil and derangement
         of normal surface drainage from past
         glaciers.
     –   Faunal diversity limited because of limited
         food supply.
     –   Populations of some species can fluctuate
         enormously in space of year or so.
     –   Insects absent in winter but superabundant in
         brief summer.




                                                           23
The Major Biomes
• Tundra
    – Distribution along northern
      edge of Northern
      Hemisphere continents.
    – Essentially a cold desert or
      grassland.
    – No trees, but considerable
      mixture of species
      (grasses, mosses, lichens,
      flowering herbs, and a
      scattering of low shrubs).
    – Dominant animal life
      consists of bird and insects
      during summer.
    – Few species of mammals
      and freshwater fishes and
      almost no reptiles or
      amphibians.




               The Major Biomes
• Alpine Tundra
    – Found in many
      high-elevation
      regions.
    – Above timberline
      there is sparse
      vegetation cover,
      consisting mostly of
      herbaceous plants,
      grasses, and low
      shrubs.




     Human Modification of Natural
        Distribution Patterns
•    Human activities severely alter natural
     distribution patterns of biota.
•    Humans directly influence biotic
     distributions in three ways:
    1. Physical removal of organisms
    2. Habitat modification
    3. Artificial translocation of organisms




                                               24
Physical Removal of Organisms
                                         • Humans severely
                                           modify landscape,
                                           affecting both plants
                                           and animal
                                           inhabitants.
                                         • Cut down, plow up,
                                           pave over, burn out,
            – Fig. 11-41. Central
              America – one of highest
                                           poison, shoot, trap,
              rates of deforestation       otherwise eradicate.
              (due mainly to
              expansion of cattle
              ranching)




             Habitat Modification
                                   • Humans affect native
                                     plants and animals by
                                     changing their habitat.
                                   • Humans change soil
                                     environment through
                                     farming, grazing,
                                     engineering, and
                                     construction.
                                   • Humans degrade
                                     atmospheric environment
                                     through pollution.
                                   • Humans impound, divert,
   • Removal for agriculture often
                                     and pollute waters.
   • results in soil erosion and
   • low crop yields as well as
   • wildlife habitat destruction.




   Tropical Rainforest Removal
• One of Earth’s most serious environmental problems, as
  of last decade or so.
• Rate of deforestation = 51 acres (21 hectares) per
  minute.
• More than half of original African rainforest is now gone,
  about 45% of Asia’s and close to 40% of Latin
  America’s.
• Current situation varies in five major rainforest regions.
• In mid-1980s, extinction rates = about 1 species per day.
• In mid 1990s, extinction rates = about 2 species per
  hour.




                                                                   25
Tropical Rainforest Removal
• Loss of forest also contributes to:
    – Accelerated soil erosion
    – Drought
    – Flooding
    – Water-quality degradation
    – Declining agricultural productivity
    – Greater poverty for rural inhabitants
    – Increasing atmospheric carbon dioxide (greenhouse
      effect).
    – Anticipated economic benefits are usually illusory.




    Tropical Rainforest Removal
• Continuous heavy (and expensive) fertilization necessary for
  sustainable agriculture.
• Losing potential valuable resources—pharmaceutical products, new
  food crops, natural insecticides, industrial material, and crop
  hybridizations (for resisting disease, insects, parasites, and other
  environmental stresses).
• Development of agroforestry (the planting of crops with trees) to
  counteract some of the destruction.
• UNESCO project to set aside reserves to protect biodiversity.
• At present about 300 preserves have been established in more than
  75 countries, encompassing 12 million hectares.
• Artificial Translocation of Organisms




    Tropical Rainforest Removal
• Humans have introduced many
  wild plants and animals into
  “new” habitats.
• Exotic species—organism that
  is introduced into “new” habitats
  in which it did not naturally
  occur.
• Sometimes deliberate,
  sometimes accidental.
• Exotics have had great impacts
  (cats on flightless bird
  populations in New Zealand,
  European flea in all parts of         Walking Catfish, Florida
  world).




                                                                         26
Biotic Rearrangement: The Sad
             Case of Florida
• Human actions in introducing exotic
  (nonnative) species result in usually one of
  two extremes:
• Either exotic dies out in a short time; or
• It flourishes extraordinarily.
• When flourishes, can occasionally have a
  salutary effect, but many cases either
  unsatisfactory or absolutely disastrous.




     Biotic Rearrangement: The Sad
             Case of Florida
• Florida presents perhaps the most frightening case
  study.
• In last decade or so, state has experienced one of
  highest in-migration of people anywhere.
• Has created massive disruptions in ecosystems, and
  exotics are most likely to prosper when ecosystem is
  unstable.
• Artificial drainage provides routes for easy dispersal of
  aquatic organisms.
• Has become the major world center for animal-import
  industry.
• Has become almost as important in plant import industry.
• Inevitable that many escape (some are turned loose).




     Biotic Rearrangement: The Sad
             Case of Florida
•   Dozens of species of exotic plants
    have become widespread, and
    most are expanding their ranges.
•   Spread of melaleuca tree has
    changed swamps to forests.
•   Aquatic weeds infest more than half
    a million acres of waters.
•   Many exotic animal species well
    established and some spreading
    rapidly.
•   Exotic fish more numerous and
    pose even more serious problems,
    competing with native species.
•   Greatest present and potential      Salt Cedars
    threat is so-called walking catfish
    from Southeast Asia.




                                                              27
Focus: Desert Adaptations of the
          Amazing Camel
• Dromedary (one-humped) camel has developed
  the most remarkable series of adjustments to
  desert environment.
• Anatomical adaptations include
• Light-colored and shiny summer coat reflects hot
  sunlight.
• Deeply cleft upper lip allows moisture loss from
  nose to be recycled back into mouth.
• Nostrils are horizontal slits that keep sand and
  dust out.




  Focus: Desert Adaptations of the
          Amazing Camel
• Eyes set beneath shaggy brows for shade, and double
  eyelids to protect eyes from sand.
• Broad and elastic feet for good traction and protection
  against the hot sand.
• Physiological adaptations include
• Highly fluctuating body temperatures that allow them to
  conserve moisture through minimal perspiration.
• Little production of urine and little moisture voided in
  their feces.
• Can stand long periods without water and can rapidly
  rehydrate when drinking.
• They cannot, however, store moisture in their hump.




      People and the Environment:
        Rainforest Loss in Brazil
• Brazil contains about one-third of the planet’s tropical rainforest.
• Settlement over the last 50 years in the forest has led to large tracts
  of it being cleared for settlement and logging.
• This clearing has been facilitated by the construction of the Cuiabá-
  Port Velho highway.
• By the late 1980s deforestation had increased substantially, and by
  2001 the amount removed was extraordinary.
• In 2004, more than 26,000 square kilometers of rainforest were lost.
• Much of the land is cleared for grazing land, which is then
  abandoned after a couple of years due to the poor fertility of tropical
  soils.
• This is leading to habitat and species loss in the region.




                                                                            28

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Ch 11

  • 1. Title Page Photo “In short, the animal and vegetable lines, diverging widely above, join below in a loop.” loop.” —Asa Gray (Brainyquote.com) Ecosystems and Biomes • Two organizing principles are ecosystem and biome. • Ecosystem: A Concept for All Scales – Ecosystem —the totality of interactions among organisms and the environment in the area of consideration. • Encompasses both the living and nonliving portion and how energy flows among them. • Weakness—there is an almost infinite variety in the magnitude of ecosystems that can be studied: – Range includes whole Earth itself to drop of water. Ecosystems and Biomes • Biome: A Scale for All Biogeographers – Biome—a large, recognizable assemblage of plants and animals in functional interaction with its environment. – Most appropriate scale for understanding world distribution patterns. 1
  • 2. Ecosystems and Biomes • Eleven major types – Often significant and even predictable relationships exist between the biota (particularly the flora) of a biome and the associated climate and soil types. • Ecotone—the transition zone between biotic communities in which the typical species of one community intermingle or interdigitate with those of Marsh/mangrove ecotone with fire, freeze, another. and hurricane damage. Taken September 20, 2001 Terrestrial Flora • Geographers interested in natural vegetation of landscape for three reasons: 1. Plants are likely to dominate a landscape (except where terrain is rugged, climate is harsh, or humans have intervened); 2. Vegetation is a sensitive indicator of other environmental attributes; 3. Vegetation is often instrumental to human settlement and activities. Characteristics of Plants • Most very hardy. • High survival potential dependent on: – Subsurface root system – Reproductive mechanism • Perennial—plant that can live more than a single year despite seasonal climatic variations. • Annual—plant that perishes during times of climatic stress but leaves behind a reservoir of seeds to germinate during the next favorable period. 2
  • 3. Characteristics of Plants • Common characteristics: – Roots (to gather nutrients and moisture and to anchor plant); – Stems and branches (to support and transport nutrients); – Leaves (to collect solar energy, exchange gases, and transpire water); – Reproductive organs. Floristic Terminology • Categorizing by reproduction: – Through spores Spores • Those that reproduce by spores are in two major groups: – Bryophytes—spore- bearing plants such as mosses and liverworts; never dominated in history, but can be very Bryophytes important in some localized situations. – Pteridophytes—spore- bearing plants such as ferns, horsetails, and clubmosses; used to dominate continental vegetation, but no more. Pteridophytes Floristic Terminology – Through seeds Seeds • Those that reproduce by seeds are in two major groups: Gymnosperms – Gymnosperms—seed- reproducing plants that carry their seeds in cones; also Angiosperms known as conifers. » Used to be more important, in geologic past. – Angiosperms—plants that have seeds encased in some sort of protective body, such as a fruit, a nut, or a seedpod. » Have dominated planet vegetation for last 50 million to 60 million years. 3
  • 4. – Other terms Trees Gymnosperms Softwood Coniferous Needleleaf Angiosperms Hardwood Deciduous Broadleaf – Fig. 11-3 Floristic Terminology • Categorizing by stem or trunk composition: – Woody plant—plant that has stem composed of hard fibrous material; refers mostly to trees and shrubs. – Herbaceous—refers to plants that have soft stems; mostly grasses, forbs, and lichens. • Categorizing by leaf retention: – Deciduous—refers to trees that experience an annual period in which all leaves die and usually fall from the tree, due either to a cold or dry season. – Evergreen—a tree or shrub that sheds its leaves on a sporadic or successive basis, but at any given time appears to be fully leaved. Floristic Terminology • Categorizing by leaf shape: – Broadleaf—tree that has flat and expansive leaves. • Majority are deciduous. • In rainy tropics, everything is evergreen. – Needleleaf—refers to trees adorned with thin slivers of tough, leathery, waxy needles rather than typical leaves. • Almost all are evergreen. 4
  • 5. Floristic Terminology • Categorizing by supposed structure—but this is unsatisfactory. – Hardwood—angiosperm tree that is usually broad- leaved and deciduous. Wood has a relatively complicated structure, but is not always hard. – Softwood —gymnosperm tree; nearly all such trees are needle-leaved evergreens with wood of simple cellular structure but not always soft. Environmental Adaptations • Two prominent adaptation strategies of plants to protect against environmental stress are – Xerophytic adaptations – Hygrophytic adaptations Environmental Adaptations • Xerophytic—refers to plants structurally adapting to withstand protracted dry conditions. – Roots, stems, leaves, reproductive cycle can all adapt in various ways. • Succulent—plant that has fleshy stem that stores water. 5
  • 6. Environmental Adaptations • Hygrophytic—refers to plants structurally adapting to withstand protracted wet conditions. • Hygrophytic Adapatation – Hygrophyte—plant that requires a saturated or semi-saturated environment (frequent soakings with water). Environmental Adaptations • Likely to have extensive root system for anchoring in soft ground. • Usually relies on buoyancy of water for support rather than stem. • Many have weak, pliable stems so can withstand currents. • Hydrophytes are often grouped in with this category. – Hydrophyte—a “water-loving” plant that is adapted to live in more or less permanently immersed in water. Environmental Adaptations • Hygrophytic Adapatation – Hygrophyte—plant that requires a saturated or semi-saturated environment (frequent soakings with water). – Likely to have extensive root system for anchoring in soft ground. – Usually relies on buoyancy of water for support rather than stem. – Many have weak, pliable stems so can withstand currents. • Hydrophytes are often grouped in with this category. – Hydrophyte—a “water-loving” plant that is adapted to live in more or less permanently immersed in water. 6
  • 7. The Critical Role of Competition • Competition is key in which plants grow where. – Even though all conditions (climatic, edaphic, etc.) are favorable, a plant Kudzu Vine may not take hold in one area because of competition. Spatial Groupings of Plants • Geographers usually more concerned with spatial groupings than individual plants. – Groups based on dominant members, dominant appearance, or both. Spatial Groupings of Plants • Floristic pattern of Earth is impermanent. • Change can be slow and orderly, as in lake infilling. • Change can be abrupt and chaotic, as in wildfire. – Climax vegetation—a stable plant association of relatively constant composition that develops at the end of a long succession of changes. 7
  • 8. Spatial Groupings of Plants • Is an association in equilibrium with prevailing environmental conditions. • Should persist until environmental disturbance/change occurs. • Seral association—various stages leading up to climax vegetation. Spatial Groupings of Plants • Geographers can face significant difficulties in recognizing spatial groupings. • As one tries to identify patterns and recognize relationships, must make generalizations. – When associations are portrayed on maps, boundaries usually represent approximations. – Human interference plays a major role. – Because of human impact, climax vegetation is now the exception rather than rule. • Maps often ignore human interference, so are actually maps of theoretical natural vegetation. Spatial Groupings of Plants • Many ways to classify plant associations. – Geographers usually place emphasis on structure and appearance of dominant plants. – Major associations include forests, woodlands, shrublands, grasslands, deserts, tundra, and wetlands. 8
  • 9. – Fig. 11-7 Spatial Groupings of Plants • Forest—an assemblage of trees growing closely together so that their individual leaf canopies generally overlap. – Likely to become climax association in any area where moisture is adequate and growing season isn’t very short. • Woodland—tree-dominated association in which the trees are spaced more widely apart than those of forests and do not have interlacing canopies. • Shrubland —plant association dominated by relatively short woody plants. – Wide latitudinal range but usually restricted to semiarid or arid areas. Spatial Groupings of Plants • Grassland—plant association dominated by grasses and forbs. – Prominent types are savanna, prairie, and steppe. – Associated with semiarid and subhumid climates. • Desert—actually a climate, not an association per se, but is typified by plants widely scattered on bare ground. • Tundra—a complex mix of very low-growing plants, including grasses, forbs, dwarf shrubs, mosses, and lichens, but no trees. – Only in the perennially cold climates of high latitudes or high altitudes. 9
  • 10. Spatial Groupings of Plants • Wetland—landscape characterized by shallow, standing water all or most of the year, with vegetation rising above the water level. – Have much more limited geographic Carson River, NV extent than any other above associations. Spatial Groupings of Plants • Various plant associations will exist in relatively narrow zones when mountain slopes have significant elevational changes in short horizontal distances. – Vertical zonation—the horizontal layering of different plant associations on a mountainside or hillside. • Elevation changes are counterpart of latitude changes. • Treeline elevation varies with latitude. • Southern and Northern hemispheres experience different elevation–latitude relationship, with Southern Hemisphere having lower treelines. • Reason for discrepancy is not understood yet. • Can have significant local variations caused by a variety of local environmental conditions. – Vertical Zonation • Most apparent in mountains due to changes in elevations over short distances – Fig. 11-10 10
  • 11. Spatial Groupings of Plants • Exposure to sunlight is often a critical determinant of vegetation composition. – Adret slope—Sun slope; slope where Sun’s rays arrive at a relatively direct angle. • Relatively hot and dry, and its vegetation is sparser and smaller than that on adjacent slopes with different exposures. • Likely to have species composition different from adjacent slopes. – Ubac slope—a slope where sunlight strikes at a low angle and hence is much less effective in heating and evaporating than on the adret slope, thus producing more luxuriant vegetation of a richer diversity. • Difference between adret and ubac decreases with increasing latitude. • Valley-bottom locations can have vegetation composition significantly different from slopes running to it. – Local Variations • Exposure to sunlight – Mountainous landscapes – Fig. 11-13 – Local Variation (continued) • Valley-bottom location – Fig. 11-14 11
  • 12. Spatial Groupings of Plants • Riparian vegetation— streamside growth, particularly prominent in relatively dry regions, where stream courses may be lined with trees, although no other trees are to be found in the landscape. Terrestrial Fauna • Animals occur in much greater variety than plants over Earth. • Animals, however, tend to be much less prominent than plants in the landscape. • They tend to be secretive and inconspicuous. • Also, environmental relationships are much less clearly evidenced by animals than plants. • Their inconspicuousness makes it more difficult to study them, and their mobility had lead to greater environmental adaptability among them. Characteristics of Animals • Variety of animal life is so great that it is difficult to find many unifying characteristics. – Two universal traits (though these aren’t always immediately recognizable): 1. Mobility The tree of life at Animal Kingdom 2. Need to eat plants and/or other animals 12
  • 13. • Characteristics of Animals – Great diversity – Two universal features • Motile • Heterotrophs – Consumers (incapable of manufacturing food from air, water and sunlight like plants do) – Fig. 11-15, 16, 19a, 21, & 27 Kinds of Animals • Size and habits are not valid indicators of animal’s significance to geographic study. • Minute and seemingly inconsequential organisms can play important roles. • Examples are carriers of disease, providers of scarce nutrients. • More than 90% of all animal species are invertebrates (without backbones). • Arthropods most prominent (insects, spiders, centipedes, millipedes, crustaceans). • Five groups of vertebrates, those with backbone: – Fish, amphibians, reptiles, birds, and mammals. • Kinds of Animals – Invertebrates – Vertebrates • Fishes • Amphibians • Reptiles • Birds • Mammals – Fig. 11-15, 16, 19a, 21, 27 & 11-17 13
  • 14. Kinds of Animals • Most mammals are placentals, having young grow and develop in mother’s body. – About 135 species are marsupials, in which mothers carry young, not fully developed at birth, in pouches. – Two species are monotremes—lay eggs. • Echidna and duckbill platypus. Environmental Adaptations • Three different kinds of evolutionary adaptation by animals: 1. Physiological 2. Behavioral 3. Reproductive Example of Animal Adaptations to Desert Life • Faunal diversity can be astounding in desert areas where water is permanent or prolonged. • Even in areas where open water is not available, there are pockets of localized favorable habitat that permit remnant populations to survive. • Most animals are completely nocturnal. • Animals are more conspicuous when cooler, such as at night and winter. • Some animals follow rains in nomadic fashion. • Most prominently displayed by birds. 14
  • 15. Example of Animal Adaptations to Desert Life • Some spend significant time underground. • Some bury selves to survive long dry spells, such as freshwater crayfish and crabs. • Text provides detailed discussion of anatomical and physiological adaptations. • A few species of rodents can exist from birth to death without ever taking a drink. • Get moisture from food. • Some species display ability to delay reproductive processes over long dry periods until more favorable conditions occur. • Australian desert kangaroos can delay implantation of fertilized blastocyst, so it remains in inactive state in uterus until better weather conditions occur. Competition Among Animals • Competition can be both direct and indirect. • Indirect—rivalry for space and resources. • Direct—antagonism of predation. • Many create social groups among own species. Competition Among Animals • Some across species, such as communal relationship among zebras, wildebeest and impalas in East African savannas. • Individual animals are concerned either largely or entirely with own survival. • Some animal species concerned with survival of mates. • Some concerned with survival of young (more common as maternal instinct, though some paternal too). • Still fewer concerned with survival for group. 15
  • 16. Cooperation among Animals • Symbiosis—association of two dissimilar organisms, in which they live together in some fashion. • Mutualism—symbiotic relationship in which the association is mutually beneficial to both organisms. • Commensalism—symbiotic relationship in which the association is neither Symbiosis beneficial nor injurious to either. • Parasitism—symbiotic relationship, in which the association benefits one, but harms the other; that is, one lives on or in the other, to Mutualism Commensalism Parasitism detriment of the host. Zoogeographic Regions • Animals’ distribution patterns more complex and irregular because of their mobility. • The broad distributions of animals nevertheless do reflect a general distribution of energy and food diversity. Zoogeographic Regions • Nine zoogeographic regions are generally recognized. • Represent average conditions and cannot portray some common pattern in which different groups of animals fit precisely. • Map on page 329. 16
  • 17.  Zoogeographic Regions • Reflective of the general distribution of energy and richness of food chemistry – Fig. 11-23 Zoogeographic Regions • Ethiopian Region – Has most diverse vertebrate fauna and greatest number of mammalian families. • Oriental Region – Similar to Ethiopian but with less diversity (save for birds and reptiles; large number of venomous snakes). • Palearctic Region – Poorer fauna than previous two; – Probably function of higher latitudes and more rigorous climate. • Palearctic Region – Separated from rest of Eurasia by mountains – Few endemic species, fewer species than in tropics – Fig. 11-23 17
  • 18. • Nearctic Region – Non-tropical portions of North America – Similar to Palearctic due to Bering land bridge – Fig. 11-23 Zoogeographic Regions • Nearctic Region – Faunal assemblage relatively poor (save for being well-represented with reptiles). – Largely a transitional zone between Palearctic and Neotropical groups. – Great similarity to Palearctic, so that some group together into superregion, Holoarctic. – Reflects how faunal dispersal occurred via Bering land bridge in geologic past. • Neotropical Region – Has rich and distinctive faunal assemblage: – Variety of habitats and isolation from other regions; – Has a larger number of endemic mammal families than any other region; – Bird fauna is exceedingly diverse and conspicuous. Zoogeographic Regions • Madagascar Region – Dominated by relic assemblage of unusual forms. – Primitive primates (lemurs). • New Zealand region – Fauna dominated by birds (mostly flightless). – Almost no terrestrial vertebrates. – No mammals and only a few reptiles and amphibians. 18
  • 19. Zoogeographic Regions • Pacific Islands Region – Limited faunal assemblage. • Australian Region – Has most distinctive fauna of any region. – Lack of variety is made up by animals’ uniqueness. – The Unique Biota of Australia – More than 90% of the native tree species in Australia are of the single genus, Eucalyptus. • Above should either be “a single genus, Euc” or “the single genus Euc” (no comma) – These trees, which there are greater than 400 species of, are native only to Australia. – The shrubs and bushes of Australia are dominated by a single genus, Acacia. – The dominance of this genus is attributed to isolation. – Australia Region • Australia and adjacent islands • Most distinctive fauna of any region due to the region’s lengthy isolation – Few placental mammals • Its unique biota are also primarily a result of isolation – Fig. 11-17: Kangaroo. Fig. 11-18: Monotremes (egg-laying mammals) Echidna and duckbill platypus. Zoogeographic Regions • Australian fauna is also unique because of isolation. – Australian fauna is dominated by a single primitive mammalian order, marsupials. – The continent is also the only home to the primitive monotremes. – Marsupials and monotremes were able to flourish in relative isolation from competitive and predatory pressures that influenced animal evolution in other parts of the world. – As such, indigenous placental mammals are lacking on the continent. 19
  • 20. The Major Biomes • Named for dominant vegetation, but encompasses fauna as well as interrelationships with soil, climate, and topography.  Major Biomes • Summary of each biome follows… – Distribution (map) – Climate types – Main vegetation types – Limiting factors to flora and fauna – Fig. 11-25 (left panel, p. 330) – Fig. 11-25 (right panel, p. 331) 20
  • 21. The Major Biomes • Tropical Rainforest – Selva — tropical rainforest; a distinctive assemblage of tropical vegetation that is dominated by a great variety of tall, high-crowned trees. • Probably most complex of all terrestrial ecosystems. – Distribution closely related to climate. – Consistent rainfall and relatively high temperatures. – Layered structure, with second layer being a branch canopy formed by the high trees that crest above the canopy. – Undergrowth relatively sparse because of lack of light. – Interior is region of heavy shade, high humidity, windless air, continuous warmth, aroma of mold and decomposition. – Fauna is largely arboreal—tree dwelling. – Canopy serves as principal food source. The Major Biomes • Tropical Deciduous Forest – Not closely correlated with specific climatic types; distribution more irregular and fragmented. – Compared to rainforest, canopy is less dense, trees are shorter, and there is less diversity of tree species (but greater variety of shrubs and other lesser plants). – Response to either less total precipitation or less periodic precipitation. – Many trees shed leaves at same time, so more sunlight can penetrate. – Produces classic jungle conditions. The Major Biomes • Tropical Scrub – Widespread in drier portions of A climatic realm (covers extensive areas in tropics and subtropics). – Dominated by low-growing scraggly trees and tall bushes, usually with extensive understory of grasses. – Plant species diversity less than that in tropical rainforest and tropical deciduous. – Faunal diversity very different from tropical rainforest and tropical deciduous. – Moderately rich assemblage of ground-dwelling mammals and reptiles, and of birds and insects. 21
  • 22. The Major Biomes • Tropical Savanna – Distribution of biome doesn’t exactly correlate with distribution of tropical savanna climate. – Incomplete correlation most noticeable where seasonal rainfall contrasts are greatest (which is associated with annual shifting of the intertropical convergence zone [ITCZ]). – Dominated by tall grasses. – Some regions actually former tropical deciduous forest and even tropical rainforest, but humans converted it through fires and by grazing domestic animals. – Has a very pronounced seasonal rhythm: wet season, dry season, and wildfire season. – Savanna fauna varies according to continent. – Africa has most remarkable, diverse large wildlife. – Latin America has only sparse population of large wildlife. The Major Biomes • Desert – Occurs extensively in midlatitude locations in Asia, North America, and South America with a fairly close correlation to Bwh and Bwk climates. – Vegetation surprisingly variable. – Shrubs are typical, with succulents common in drier parts. – Trees can be found, particularly in Australia. – Most deserts have moderately diverse faunal assemblage. – Variety of large mammals is limited. The Major Biomes • Mediterranean Woodland and Shrub – Six widely scattered and relatively small areas in midlatitudes. – Have pronounced dry-summer wet-winter precipitation. – Dominant vegetation associations are physically similar, but taxonomically quite varied. – Dominated by dense growth of woody shrubs, but also have open grassy woodlands. – Plant species vary from region to region, but in all, the trees and shrubs are primarily broadleaf evergreens. – Many plants are adapted to rapid recovery after wildfire. – Fauna not particularly distinctive. – Seed-eating, burrowing rodents common. – General overlap of animals between this biome and adjacent ones. 22
  • 23. The Major Biomes • Midlatitude Grassland – Locational coincidence between this biome and steppe climatic type is very pronounced in Northern Hemisphere. – Less distinct climatic correlations in Southern Hemisphere. – Occurs widely in midlatitudes of North America and Eurasia. – Low precipitation and/or frequency of fire prevent growth of tree or shrub seedlings. – Characteristics of grasses depends on moisture: taller in wetter area (prairie), shorter in dryer (steppe), and sometimes not continuous, but grow in discrete tufts. – Before human encroachment, fauna comprised of large numbers of relatively few species, with migratory larger herbivores. The Major Biomes • Midlatitude Deciduous Forest – Used to be far more extensive in all Northern Hemisphere continents and to some extent in tracts in Southern Hemisphere. – Humans have cleared away large portions for agriculture. – Fairly dense growth of tall broadleaf trees with complete canopy in summer. – Winter very different, with seasonal fall of leaves. – Tree species vary greatly from region to region. – Generally has the richest assemblage of fauna in midlatitudes. – Seasonal variation to fauna (hibernation and migration). The Major Biomes • Boreal Forest – An extensive needleleaf forest in subarctic regions of North America and Eurasia; also called taiga. – One of most extensive biomes, occupying vast expanse of northern North America and Eurasia. – Close correlation with subarctic climatic type. – Has perhaps simplest assemblage of plants. – Most trees are conifers, though in some places deciduous trees interrupt the coniferous cover. – Trees become spindlier, short, and openly spaced in north. – Bogs and swamps numerous because of permanently frozen subsoil and derangement of normal surface drainage from past glaciers. – Faunal diversity limited because of limited food supply. – Populations of some species can fluctuate enormously in space of year or so. – Insects absent in winter but superabundant in brief summer. 23
  • 24. The Major Biomes • Tundra – Distribution along northern edge of Northern Hemisphere continents. – Essentially a cold desert or grassland. – No trees, but considerable mixture of species (grasses, mosses, lichens, flowering herbs, and a scattering of low shrubs). – Dominant animal life consists of bird and insects during summer. – Few species of mammals and freshwater fishes and almost no reptiles or amphibians. The Major Biomes • Alpine Tundra – Found in many high-elevation regions. – Above timberline there is sparse vegetation cover, consisting mostly of herbaceous plants, grasses, and low shrubs. Human Modification of Natural Distribution Patterns • Human activities severely alter natural distribution patterns of biota. • Humans directly influence biotic distributions in three ways: 1. Physical removal of organisms 2. Habitat modification 3. Artificial translocation of organisms 24
  • 25. Physical Removal of Organisms • Humans severely modify landscape, affecting both plants and animal inhabitants. • Cut down, plow up, pave over, burn out, – Fig. 11-41. Central America – one of highest poison, shoot, trap, rates of deforestation otherwise eradicate. (due mainly to expansion of cattle ranching) Habitat Modification • Humans affect native plants and animals by changing their habitat. • Humans change soil environment through farming, grazing, engineering, and construction. • Humans degrade atmospheric environment through pollution. • Humans impound, divert, • Removal for agriculture often and pollute waters. • results in soil erosion and • low crop yields as well as • wildlife habitat destruction. Tropical Rainforest Removal • One of Earth’s most serious environmental problems, as of last decade or so. • Rate of deforestation = 51 acres (21 hectares) per minute. • More than half of original African rainforest is now gone, about 45% of Asia’s and close to 40% of Latin America’s. • Current situation varies in five major rainforest regions. • In mid-1980s, extinction rates = about 1 species per day. • In mid 1990s, extinction rates = about 2 species per hour. 25
  • 26. Tropical Rainforest Removal • Loss of forest also contributes to: – Accelerated soil erosion – Drought – Flooding – Water-quality degradation – Declining agricultural productivity – Greater poverty for rural inhabitants – Increasing atmospheric carbon dioxide (greenhouse effect). – Anticipated economic benefits are usually illusory. Tropical Rainforest Removal • Continuous heavy (and expensive) fertilization necessary for sustainable agriculture. • Losing potential valuable resources—pharmaceutical products, new food crops, natural insecticides, industrial material, and crop hybridizations (for resisting disease, insects, parasites, and other environmental stresses). • Development of agroforestry (the planting of crops with trees) to counteract some of the destruction. • UNESCO project to set aside reserves to protect biodiversity. • At present about 300 preserves have been established in more than 75 countries, encompassing 12 million hectares. • Artificial Translocation of Organisms Tropical Rainforest Removal • Humans have introduced many wild plants and animals into “new” habitats. • Exotic species—organism that is introduced into “new” habitats in which it did not naturally occur. • Sometimes deliberate, sometimes accidental. • Exotics have had great impacts (cats on flightless bird populations in New Zealand, European flea in all parts of Walking Catfish, Florida world). 26
  • 27. Biotic Rearrangement: The Sad Case of Florida • Human actions in introducing exotic (nonnative) species result in usually one of two extremes: • Either exotic dies out in a short time; or • It flourishes extraordinarily. • When flourishes, can occasionally have a salutary effect, but many cases either unsatisfactory or absolutely disastrous. Biotic Rearrangement: The Sad Case of Florida • Florida presents perhaps the most frightening case study. • In last decade or so, state has experienced one of highest in-migration of people anywhere. • Has created massive disruptions in ecosystems, and exotics are most likely to prosper when ecosystem is unstable. • Artificial drainage provides routes for easy dispersal of aquatic organisms. • Has become the major world center for animal-import industry. • Has become almost as important in plant import industry. • Inevitable that many escape (some are turned loose). Biotic Rearrangement: The Sad Case of Florida • Dozens of species of exotic plants have become widespread, and most are expanding their ranges. • Spread of melaleuca tree has changed swamps to forests. • Aquatic weeds infest more than half a million acres of waters. • Many exotic animal species well established and some spreading rapidly. • Exotic fish more numerous and pose even more serious problems, competing with native species. • Greatest present and potential Salt Cedars threat is so-called walking catfish from Southeast Asia. 27
  • 28. Focus: Desert Adaptations of the Amazing Camel • Dromedary (one-humped) camel has developed the most remarkable series of adjustments to desert environment. • Anatomical adaptations include • Light-colored and shiny summer coat reflects hot sunlight. • Deeply cleft upper lip allows moisture loss from nose to be recycled back into mouth. • Nostrils are horizontal slits that keep sand and dust out. Focus: Desert Adaptations of the Amazing Camel • Eyes set beneath shaggy brows for shade, and double eyelids to protect eyes from sand. • Broad and elastic feet for good traction and protection against the hot sand. • Physiological adaptations include • Highly fluctuating body temperatures that allow them to conserve moisture through minimal perspiration. • Little production of urine and little moisture voided in their feces. • Can stand long periods without water and can rapidly rehydrate when drinking. • They cannot, however, store moisture in their hump. People and the Environment: Rainforest Loss in Brazil • Brazil contains about one-third of the planet’s tropical rainforest. • Settlement over the last 50 years in the forest has led to large tracts of it being cleared for settlement and logging. • This clearing has been facilitated by the construction of the Cuiabá- Port Velho highway. • By the late 1980s deforestation had increased substantially, and by 2001 the amount removed was extraordinary. • In 2004, more than 26,000 square kilometers of rainforest were lost. • Much of the land is cleared for grazing land, which is then abandoned after a couple of years due to the poor fertility of tropical soils. • This is leading to habitat and species loss in the region. 28