This document discusses how platform ecosystems evolve over time through changes in platform design, governance strategies, and the exogenous evolution of users and developers. As platforms grow larger, they gain bargaining power over developers and provide less overall support. Late adopters on platforms have lower risk tolerance and preference for novelty compared to early adopters. The number of developers entering platforms increases over time, reducing the average potential market for each new app or game. This affects how developers create, capture, and distribute value on platforms over the lifecycle. Strategies are discussed for how developers and platform owners can address these evolutionary dynamics.