DMARC requires alignment between the authentication domain from SPF or DKIM and the From header domain. SPF authenticates the MAIL FROM or EHLO domain, while DKIM authenticates the domain in the d= tag. Alignment can be strict, requiring an exact match, or relaxed, allowing subdomains. DMARC policies use tags to specify whether strict or relaxed alignment is needed for SPF (aspf) or DKIM (adkim) to pass DMARC verification. Alignment helps validate the authenticity of the From header by linking it to the authenticated domains.