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Role of commitments and
emotions for long-term relations
a modelling perspective
Tom Lenaerts
MLG AI lab
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
coordination
cooperation
Human society
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Simple interactions
Complex patterns
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Global
local
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Catastrophes
Errors
Attacks
Mistakes
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Why don’t things fail more often?
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Mechanisms that induce
Trust
Translation to ICT solutions
Inherent to every human
being
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Good Agreements Make Good Friends
The Anh Han1,2
, Luı´s Moniz Pereira3
, Francisco C. Santos4,5
& Tom Lenaerts1,2
1
AI lab, Computer Science Department, Vrije Universiteit Brussel, Pleinlaan 2, 1050 Brussels, Belgium, 2
MLG, De´partement
d’Informatique, Universite´ Libre de Bruxelles, Boulevard du Triomphe CP212, 1050 Brussels, Belgium, 3
Centro de Inteligeˆncia
Artificial (CENTRIA), Departamento de Informa´tica, Faculdade de Cieˆncias e Tecnologia, Universidade Nova de Lisboa, 2829-516
Caparica, Portugal, 4
INESC-ID and Instituto Superior Te´nico, Universidade de Lisboa, IST-Taguspark, 2744-016 Porto Salvo,
Portugal, 5
ATP-group, CMAF, Instituto para a Investigaça˜o Interdisciplinar, P-1649-003 Lisboa Codex, Portugal.
When starting a new collaborative endeavor, it pays to establish upfront how strongly your partner commits
to the common goal and what compensation can be expected in case the collaboration is violated. Diverse
examples in biological and social contexts have demonstrated the pervasiveness of making prior agreements
on posterior compensations, suggesting that this behavior could have been shaped by natural selection.
Here, we analyze the evolutionary relevance of such a commitment strategy and relate it to the costly
punishment strategy, where no prior agreements are made. We show that when the cost of arranging a
commitment deal lies within certain limits, substantial levels of cooperation can be achieved. Moreover,
these levels are higher than that achieved by simple costly punishment, especially when one insists on
sharing the arrangement cost. Not only do we show that good agreements make good friends, agreements
based on shared costs result in even better outcomes.
C
onventional wisdom suggests that cooperative interactions have a bigger chance of surviving when all
participants are aware of the expectations and the possible consequences of their actions. All parties then
clearly know to what they commit and can refuse such a commitment whenever the offer is made. A
classical example of such an agreement is marriage1,2
. In that case mutual commitment ensures some stability in
the relationship, reducing the fear of exploitation and providing security against potential cataclysms. Clearly
such agreements can be beneficial in many situations, which are not limited to the type of formal and explicit
contracts as is the case for marriage. Commitments may even be arranged in a much more implicit manner as is
the case for members of the same religion3,4
, or by some elaborate signaling mechanism as is the case in primates’
use of signaling to synchronize expectations and the consequences of defaulting on commitment in their different
ventures5
.
Here we investigate analytically and numerically whether costly commitment strategies, in which players
propose, initiate and honor a deal, are viable strategies for the evolution of cooperative behavior, using the
symmetric, pairwise, and non-repeated Prisoner’s Dilemma (PD) game to model a social dilemma. Next to
the traditional cooperate (C) and defect (D) options, a player can propose its co-player to commit to cooperation
before playing the PD game, willing to pay a personal cost ð Þ to make it credible. If the co-player accepts the
arrangement and also plays C, they both receive their rewards for mutual cooperation. Yet if the co-player plays D,
then he or she will have to provide the proposer with a compensation at a personal cost (d). Finally, when the co-
player does not accept the deal, the game is not played and hence both obtain no payoff.
Although there is a kind of punishment associated with the agreement, the notion of compensating a partner
when not honoring a negotiated deal is not entirely equivalent to the general notion of punishment as has been
studied in Evolutionary Game Theory6–13
so far. In the current work, both parties are aware of the stakes before
they start the interaction: the person who accepts to commit knows upfront what to expect from the person that
proposes the commitment and what will happen if he or she does not act appropriately. Even more, the co-player
has the possibility not to accept such an agreement and continue interacting without any prior commitment with
the other players, and with no posterior repercussions from commitment proposers. In the current literature,
punishment (with or without cost) is imposed as a result of ‘‘bad’’ behavior, which can only be escaped by not
participating in the game at all9,12,14
. As such, the present work differs from peer and pool punishment9,12
in that
the latter imposes the commitment to the other players, i.e. defectors will always be punished even when they did
not want to play with punishers. In addition, there is no notion of compensation incorporated in the model that
remunerates the proposer when her accepted deal is violated, in contradistinction to our own model. Moreover,
because the creation of the agreement occurs explicitly in the current work, players can behave conditionally
(even without considering previous interactions, whether direct or indirect), and that is not considered in the
OPEN
SUBJECT AREAS:
BIOLOGICAL PHYSICS
BEHAVIOURAL METHODS
EVOLUTIONARY THEORY
SOCIAL EVOLUTION
Received
2 July 2013
Accepted
30 August 2013
Published
18 September 2013
Correspondence and
requests for materials
should be addressed to
T.A.H. (h.anh@ai.vub.
ac.be) or T.L. (Tom.
Lenaerts@ulb.ac.be)
SCIENTIFIC REPORTS | 3 : 2695 | DOI: 10.1038/srep02695 1
Our (cognitive) capacity for
commitment may have evolved
by natural selection
Emotions manage mutually
beneficial relationships
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Commitment ?
“A commitment is an act or signal that gives up options in
order to influence someone’s behaviour by changing
incentives and expectations”
“They can be enforced by external incentives,
but also by some combination of reputation
and emotion”
“Commitments can be promises to help, or
threats to harm”
“Our (cognitive) capacity for commitment
may have evolved by natural selection”
prisoners’ dilemma
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
R = reward
S = suckers payoff
T = temptation to defect
P = punishment
fear = P >S
greed = T > R
R
D
C
C
D
S
PT
T > R > P > S
C.H. Coombs (1973) A reparameterization of the prisoner’s dilemma
game. Behavioral Science 18:424-428
prisoners’ dilemma
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
R
D
C
C
D
S
PT
T > R > P > S
C.H. Coombs (1973) A reparameterization of the prisoner’s dilemma
game. Behavioral Science 18:424-428
7
3
3 7
0
1
1
0
C
D
C D
7
1
1
7
Nash equilibrium
Q1
Q2
Propose
(at a cost ε) NOT propose
accept
NOT
accept
IN OUT
OUT
Adding commitment
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Propose = promise to play C or
threat to not play at all if not playing C
Accept = agree to also play C
(following promise or threat)
COMMIT
C
D
C D
FREE
FREE
IN OUT
propose accept
NOT propose
Q3
C D C D
Q3 COMMIT
NOT
play
propose NOT accept
COMMIT
propose/
accept
C
C
accept
D
FREE
FREE
FAKE
pay compensation
(δ)
accept
FAKE
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
R-ε/2 R-ε 0 S+δ-ε R-ε
R R S S S
0 T P P P
T-δ T P P P
R T P P P
COMMIT
FAKE
FREE
COMMIT FAKE FREE
C
D
C D
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
T=b;
R=b-c;
P=0;
S=-c
Translation to Donation
game
b=benefit of cooperation
c=cost of cooperation
b-c
-ε/2
b-c-ε 0 -c+δ-ε b-c-ε
b-c b-c -c -c -c
0 b 0 0 0
b-δ b 0 0 0
b-c b 0 0 0
COMMIT
FAKE
FREE
COMMIT FAKE FREE
C
D
C D
δ > c+3ε/4
ε < 2(b-c)/3
Cooperation will evolve
when …
T=b;
R=b-c;
P=0;
S=-c
Translation to Donation
game
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
b=benefit of cooperation
c=cost of cooperation
Frequencyofstrategy
ε
δ=4
COMMIT
FREE
D
δ=4
ε=0.25
FAKE
Same conclusions hold for 2 or more players
β=0.1
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
F
P
F
P
A
DC
P
D
F
P
Commitments are often
long-term…
Iterated prisoners’ dilemma
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
With a probability ω the interaction continues
time
C C C C
C C C
P1
P2 C C C C
C C C …
…
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
How to deal with mistakes ? (intentional or not)
Should we collect the compensation or continue
the agreement?
Should one take revenge or apologise and
forgive?
With a probability ω the interaction continues
time
C D C C
C C C
P1
P2 D D C C
C C D …
…
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Individuals prefer to
take revenge
time
C C C D
C C D
P1
P2 C C C C
D D D …
…
0.4
−0.2
0
0.2
0.4 a
t
)−Fr(NC,−,AllD)
NN
AllD
PN
AllD
TFT
abundancerelative
tothedefectors
0.0
0.2
0.4
-0.2
log(noise)
-3 -2 -1
D
P-C-D
P-C-TFT
pay compen-
sation δ
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
When apologising is
not possible
time
C C C C
C C D
P1
P2 C C C C
C C C …
…
Oeps!
sorry
really sorry
no
problem
apology
cost (γ)
0 2 4 6 8
−0.1
0
0.1
0.2
0.3
0.4
γ
Fr(S)−Fr(NC,−,AllD,−)
α=0.01
0 2 4 6 8
−0.1
0
0.1
0.2
0.3
0.4
γ
Fr(S)−Fr(NC,−,AllD,−)
α=0.1
(P,C,AllD,q=1) (P,C,AllD,q=0) (P,D,AllD,q=1) (A,D,AllD,q=1)
abundancerelative
tothedefectors
0.0
0.1
0.3
0.2
0.4
-0.1
0 2 4 6 8
D
P-C-D+A
P-C-D + NA
apology cost
P-D-D+A
A-D-D+A
These are like
the fake players
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
When apologising is
possible
Apology needs to
be sincere
To err is humane, to forgive …
Alexander Pope, An essay on Criticism (1711)
evolutionary viable when
the apology is sincere
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Science=collaboration
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
The Anh Han (UK) Luis Moniz Pereira (P) Luis Martinez (I)
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Produces equivalent results as in the mathematical
model
0
0.5
1
1.5
2
2.5
3
3.5
4
4.5
5
x 10
−3b/c=1.5, =0.01
0 2 4 6 8
0
2
4
6
8
b/c=2, =0.01
0 2 4 6 8
0
2
4
6
8
b/c=4, =0.01
0 2 4 6 8
0
2
4
6
8
b/c=1.5, =0.1
0 2 4 6 8
0
2
4
6
8
b/c=2, =0.1
0 2 4 6 8
0
2
4
6
8
b/c=4, =0.1
0 2 4 6 8
0
2
4
6
8
Agent-based modelling
An agent’s
genotype is
defined by:
apology cost (γ)
forgiveness
threshold (τγ)
© Tom Lenaerts, 2017 AISB in Bath
S
T
R+1RR-1
P-1
P+1
P
7
3
3 7
0
1
1
0
C
D
C D T > R > P > S
7
3
3 7
1
0
0
1
C
D
C D T > R > S > P
3
7
7 3
0
1
1
0
C
D
C D
R > T > P > S
3
7
7 3
1
0
0
1
C
D
C D
R > T > S > P
Space of social dilemmas
© Tom Lenaerts, 2017 AISB in Bath
S
T
R+1RR-1
P-1
P+1
P
C D
C
D
S-P
R-S-T+P
C
D
An evolutionary game theory
perspective ….
REPLICATOR DYNAMICS
C D
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Finite population dynamics
D
D
C
D
CD
DC
DD
t µ
CD
O
1-µ
C
C
p=[1+eβ(f(D)-f(C))]-1
D
C
D
CD
DC
DD
t+1
D
A moran process (birth-death process)
CC
selection strength
© Tom Lenaerts, 2016
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Assuming that mutations are rare, we have either …
n(C)=Z
C
C
C
C
C
CC
CC
C
C
C
n(C)=0
D
D
D
D
DD
DD
D
D
D
D
n(C)=1
C
D
D
D
D
DD
DD
D
D
D
...
fixation probability
ρ of C in D
population
n(C)=Z-1
C
C
C
C
C
CD
CC
C
C
C
fixation probability
ρ of D in C
population
Finite population dynamics
© Tom Lenaerts, 2016
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Which produces a reduced Markov chain
For which the stationary distributions can
be calculated = how likely it is to end up in either
monomorphic state
DC
Fudenberg, D., & Imhof, L.A. (2006). Imitation processes with small mutations. J. Econ.Theo, 131,251–262.
Imhof, L.A., Fudenberg, D., & Nowak, M.A. (2005). Evolutionary cycles of cooperation and defection. Proc
Nat Acad Sci USA, 102(31), 10797–10800.
ρCD
ρDC
Dynamics in finite populations
x% y%=1-x
© Tom Lenaerts, 2016
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Dynamics in finite populations
4
3
3 4
0
1
1
0
C
DC
D
DC
1.6ρN
0.6ρN
26.5%
73.5%
with β=0.01
10
-5
10
-4
10
-3
10
-2
10
-1
10
0
10
1
Selection strength (β)
0
0.1
0.2
0.3
0.4
0.5
0.6
0.7
0.8
0.9
1
Stationarydistribution
C
D
for varying β
ρN=drift=1/N
How can we
achieve
cooperation?
© Tom Lenaerts, 2016
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn
Rules for the evolution of
cooperation
Nowak, M.A. Five Rules for the Evolution of Cooperation. Science 314, 1560–1563 (2006).
provided that
b/c > 1 + (n/m) (5)
Evolutionary Success
Before proceeding to a comparative analysis of
the five mechanisms, let me introduce some
The entries denote the payoff for the row
player. Without any mechanism for the evolution
of cooperation, defectors dominate cooperators,
which means a < g and b < d. A mechanism for
the evolution of cooperation can change these
inequalities.
1) If a > g, then cooperation is an evo-
lutionarily stable strategy (ESS). An infinitely
large population of cooperators cannot be in-
vaded by defectors under deterministic selec-
tion dynamics (32).
2) If a + b > g + d, then cooperators are
risk-dominant (RD). If both strategies are
ESS, then the risk-dominant strategy has the
bigger basin of attraction.
3) If a + 2b > g + 2d, then cooperators are
advantageous (AD). This concept is important
for stochastic game dynamics in finite pop-
ulations. Here, the crucial quantity is the fix-
ation probability of a strategy, defined as the
probability that the lineage arising from a
single mutant of that strategy will take over
the entire population consisting of the other
strategy. An AD strategy has a fixation proba-
bility greater than the inverse of the popu-
lation size, 1/N. The condition can also be
expressed as a 1/3 rule: If the fitness of the in-
vading strategy at a frequency of 1/3 is greater
than the fitness of the resident, then the fix-
ation probability of the invader is greater than
1/N. This condition holds in the limit of weak
selection (52).
We have encountered
evolution of cooperati
mathematical formali
mechanisms are very
each theory is a simp
coherent mathematica
the derivation of all fi
is that each mechanis
game between two st
payoff matrix (Table
can derive the relevan
of cooperation.
For kin selection
inclusive fitness prop
(31). The relatedness
Therefore, your payof
to mine. A second me
to a different matrix
direct reciprocity, the
while the defectors
expected number of r
tit-for-tat players coop
tat versus always-defe
first move and then
iprocity, the probabili
reputation is given b
unless the reputation
dicates a defector. A
network reciprocity, i
expected frequency o
by a standard replicat
formed payoff matrix
the payoff matrices o
Kin selection
Network reciprocity
Direct reciprocity
Indirect reciprocity
1 r
the five mechanisms, let me introduce some inequalities.
1) If a > g, then coope
lutionarily stable strategy (E
large population of cooperat
vaded by defectors under de
tion dynamics (32).
2) If a + b > g + d, the
risk-dominant (RD). If bo
ESS, then the risk-dominant
bigger basin of attraction.
3) If a + 2b > g + 2d, the
advantageous (AD). This con
for stochastic game dynami
ulations. Here, the crucial q
ation probability of a strateg
probability that the lineage
single mutant of that strateg
the entire population consis
strategy. An AD strategy has
bility greater than the inve
lation size, 1/N. The condi
expressed as a 1/3 rule: If the
vading strategy at a frequenc
than the fitness of the resid
ation probability of the invad
1/N. This condition holds in
selection (52).
Kin selection
Network reciprocity
Direct reciprocity
Indirect reciprocity
1 r
Table 1. Each mechanism ca
interaction between cooperato
essary conditions for evolution
Before proceeding to a comparative analysis of
the five mechanisms, let me introduce some
the evolution of cooperation ca
inequalities.
1) If a > g, then cooperat
lutionarily stable strategy (ESS
large population of cooperators
vaded by defectors under deter
tion dynamics (32).
2) If a + b > g + d, then c
risk-dominant (RD). If both
ESS, then the risk-dominant s
bigger basin of attraction.
3) If a + 2b > g + 2d, then
advantageous (AD). This conce
for stochastic game dynamics
ulations. Here, the crucial quan
ation probability of a strategy,
probability that the lineage a
single mutant of that strategy
the entire population consistin
strategy. An AD strategy has a
bility greater than the inverse
lation size, 1/N. The conditio
expressed as a 1/3 rule: If the fi
vading strategy at a frequency o
than the fitness of the residen
ation probability of the invader
1/N. This condition holds in th
selection (52).
Kin selection
Network reciprocity
Direct reciprocity
Indirect reciprocity
1 r
Table 1. Each mechanism can
interaction between cooperators
Network reciprocity
Indirect reciprocity
Group selection
Cooperators Defectors
Fig. 3. Five mechanisms for the evolution of
cooperation. Kin selection operates when the
probability that the lineage arising from a
single mutant of that strategy will take over
the entire population consisting of the other
strategy. An AD strategy has a fixation proba-
bility greater than the inverse of the popu-
lation size, 1/N. The condition can also be
expressed as a 1/3 rule: If the fitness of the in-
vading strategy at a frequency of 1/3 is greater
than the fitness of the resident, then the fix-
ation probability of the invader is greater than
1/N. This condition holds in the limit of weak
selection (52).
tit-for-tat players cooperate
tat versus always-defect co
first move and then defec
iprocity, the probability of
reputation is given by q.
unless the reputation of
dicates a defector. A defe
network reciprocity, it can
expected frequency of coo
by a standard replicator e
formed payoff matrix (54)
the payoff matrices of the
Network reciprocity
Group selection
Cooperators Defectors
Fig. 3. Five mechanisms for the evolution of
cooperation. Kin selection operates when the
donor and the recipient of an altruistic act are
genetic relatives. Direct reciprocity requires re-
peated encounters between the same two individ-
uals. Indirect reciprocity is based on reputation; a
helpful individual is more likely to receive help.
Network reciprocity means that clusters of coop-
erators outcompete defectors. Group selection is
Table 1. Each mechanism can be described by a simple 2 × 2 payoff matri
interaction between cooperators and defectors. From these matrices we can di
essary conditions for evolution of cooperation. The parameters c and b denote,
for the donor and the benefit for the recipient. For network reciprocity, we us
[(b − c)k − 2c]/[(k + 1)(k − 2)]. All conditions can be expressed as the
exceeding a critical value. See (53) for further explanations of the underlyin
Cooperation is…
0
)1)((
rcbD
cbrrcbC
0
)1/()(
bD
cwcbC
0)1(
)1(
qbD
qccbC
0HbD
cHcbC
)())(( cnmcbnmcbC
DC
Kin
selection
Direct
reciprocity
Indirect
reciprocity
Network
reciprocity
Group
rc
b 1
rc
b 1
rc
b 1
r…
w…
q…
k…
n…
wc
b 1
w
w
c
b 2
w
w
c
b 23
qc
b 1
q
q
c
b 2
q
q
c
b 23
k
c
b
k
c
b
k
c
b
nb
1
nb
1
ESS RD AD
nb
1
Payoff matrix
Assortment
© Tom Lenaerts, 2018 FutureICT2.0 Tallinn

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Role of commitment and emotions for long-term relations

  • 1. Role of commitments and emotions for long-term relations a modelling perspective Tom Lenaerts MLG AI lab
  • 2. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn coordination cooperation Human society
  • 3. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Simple interactions Complex patterns
  • 4. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Global local
  • 5. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Catastrophes Errors Attacks Mistakes
  • 6. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Why don’t things fail more often?
  • 7. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Mechanisms that induce Trust Translation to ICT solutions Inherent to every human being
  • 8. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Good Agreements Make Good Friends The Anh Han1,2 , Luı´s Moniz Pereira3 , Francisco C. Santos4,5 & Tom Lenaerts1,2 1 AI lab, Computer Science Department, Vrije Universiteit Brussel, Pleinlaan 2, 1050 Brussels, Belgium, 2 MLG, De´partement d’Informatique, Universite´ Libre de Bruxelles, Boulevard du Triomphe CP212, 1050 Brussels, Belgium, 3 Centro de Inteligeˆncia Artificial (CENTRIA), Departamento de Informa´tica, Faculdade de Cieˆncias e Tecnologia, Universidade Nova de Lisboa, 2829-516 Caparica, Portugal, 4 INESC-ID and Instituto Superior Te´nico, Universidade de Lisboa, IST-Taguspark, 2744-016 Porto Salvo, Portugal, 5 ATP-group, CMAF, Instituto para a Investigaça˜o Interdisciplinar, P-1649-003 Lisboa Codex, Portugal. When starting a new collaborative endeavor, it pays to establish upfront how strongly your partner commits to the common goal and what compensation can be expected in case the collaboration is violated. Diverse examples in biological and social contexts have demonstrated the pervasiveness of making prior agreements on posterior compensations, suggesting that this behavior could have been shaped by natural selection. Here, we analyze the evolutionary relevance of such a commitment strategy and relate it to the costly punishment strategy, where no prior agreements are made. We show that when the cost of arranging a commitment deal lies within certain limits, substantial levels of cooperation can be achieved. Moreover, these levels are higher than that achieved by simple costly punishment, especially when one insists on sharing the arrangement cost. Not only do we show that good agreements make good friends, agreements based on shared costs result in even better outcomes. C onventional wisdom suggests that cooperative interactions have a bigger chance of surviving when all participants are aware of the expectations and the possible consequences of their actions. All parties then clearly know to what they commit and can refuse such a commitment whenever the offer is made. A classical example of such an agreement is marriage1,2 . In that case mutual commitment ensures some stability in the relationship, reducing the fear of exploitation and providing security against potential cataclysms. Clearly such agreements can be beneficial in many situations, which are not limited to the type of formal and explicit contracts as is the case for marriage. Commitments may even be arranged in a much more implicit manner as is the case for members of the same religion3,4 , or by some elaborate signaling mechanism as is the case in primates’ use of signaling to synchronize expectations and the consequences of defaulting on commitment in their different ventures5 . Here we investigate analytically and numerically whether costly commitment strategies, in which players propose, initiate and honor a deal, are viable strategies for the evolution of cooperative behavior, using the symmetric, pairwise, and non-repeated Prisoner’s Dilemma (PD) game to model a social dilemma. Next to the traditional cooperate (C) and defect (D) options, a player can propose its co-player to commit to cooperation before playing the PD game, willing to pay a personal cost ð Þ to make it credible. If the co-player accepts the arrangement and also plays C, they both receive their rewards for mutual cooperation. Yet if the co-player plays D, then he or she will have to provide the proposer with a compensation at a personal cost (d). Finally, when the co- player does not accept the deal, the game is not played and hence both obtain no payoff. Although there is a kind of punishment associated with the agreement, the notion of compensating a partner when not honoring a negotiated deal is not entirely equivalent to the general notion of punishment as has been studied in Evolutionary Game Theory6–13 so far. In the current work, both parties are aware of the stakes before they start the interaction: the person who accepts to commit knows upfront what to expect from the person that proposes the commitment and what will happen if he or she does not act appropriately. Even more, the co-player has the possibility not to accept such an agreement and continue interacting without any prior commitment with the other players, and with no posterior repercussions from commitment proposers. In the current literature, punishment (with or without cost) is imposed as a result of ‘‘bad’’ behavior, which can only be escaped by not participating in the game at all9,12,14 . As such, the present work differs from peer and pool punishment9,12 in that the latter imposes the commitment to the other players, i.e. defectors will always be punished even when they did not want to play with punishers. In addition, there is no notion of compensation incorporated in the model that remunerates the proposer when her accepted deal is violated, in contradistinction to our own model. Moreover, because the creation of the agreement occurs explicitly in the current work, players can behave conditionally (even without considering previous interactions, whether direct or indirect), and that is not considered in the OPEN SUBJECT AREAS: BIOLOGICAL PHYSICS BEHAVIOURAL METHODS EVOLUTIONARY THEORY SOCIAL EVOLUTION Received 2 July 2013 Accepted 30 August 2013 Published 18 September 2013 Correspondence and requests for materials should be addressed to T.A.H. (h.anh@ai.vub. ac.be) or T.L. (Tom. Lenaerts@ulb.ac.be) SCIENTIFIC REPORTS | 3 : 2695 | DOI: 10.1038/srep02695 1 Our (cognitive) capacity for commitment may have evolved by natural selection Emotions manage mutually beneficial relationships
  • 9. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Commitment ? “A commitment is an act or signal that gives up options in order to influence someone’s behaviour by changing incentives and expectations” “They can be enforced by external incentives, but also by some combination of reputation and emotion” “Commitments can be promises to help, or threats to harm” “Our (cognitive) capacity for commitment may have evolved by natural selection”
  • 10. prisoners’ dilemma © Tom Lenaerts, 2018 FutureICT2.0 Tallinn R = reward S = suckers payoff T = temptation to defect P = punishment fear = P >S greed = T > R R D C C D S PT T > R > P > S C.H. Coombs (1973) A reparameterization of the prisoner’s dilemma game. Behavioral Science 18:424-428
  • 11. prisoners’ dilemma © Tom Lenaerts, 2018 FutureICT2.0 Tallinn R D C C D S PT T > R > P > S C.H. Coombs (1973) A reparameterization of the prisoner’s dilemma game. Behavioral Science 18:424-428 7 3 3 7 0 1 1 0 C D C D 7 1 1 7 Nash equilibrium
  • 12. Q1 Q2 Propose (at a cost ε) NOT propose accept NOT accept IN OUT OUT Adding commitment © Tom Lenaerts, 2018 FutureICT2.0 Tallinn Propose = promise to play C or threat to not play at all if not playing C Accept = agree to also play C (following promise or threat) COMMIT C D C D FREE FREE
  • 13. IN OUT propose accept NOT propose Q3 C D C D Q3 COMMIT NOT play propose NOT accept COMMIT propose/ accept C C accept D FREE FREE FAKE pay compensation (δ) accept FAKE © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 14. R-ε/2 R-ε 0 S+δ-ε R-ε R R S S S 0 T P P P T-δ T P P P R T P P P COMMIT FAKE FREE COMMIT FAKE FREE C D C D © Tom Lenaerts, 2018 FutureICT2.0 Tallinn T=b; R=b-c; P=0; S=-c Translation to Donation game b=benefit of cooperation c=cost of cooperation
  • 15. b-c -ε/2 b-c-ε 0 -c+δ-ε b-c-ε b-c b-c -c -c -c 0 b 0 0 0 b-δ b 0 0 0 b-c b 0 0 0 COMMIT FAKE FREE COMMIT FAKE FREE C D C D δ > c+3ε/4 ε < 2(b-c)/3 Cooperation will evolve when … T=b; R=b-c; P=0; S=-c Translation to Donation game © Tom Lenaerts, 2018 FutureICT2.0 Tallinn b=benefit of cooperation c=cost of cooperation
  • 16. Frequencyofstrategy ε δ=4 COMMIT FREE D δ=4 ε=0.25 FAKE Same conclusions hold for 2 or more players β=0.1 © Tom Lenaerts, 2018 FutureICT2.0 Tallinn F P F P A DC P D F P
  • 17. Commitments are often long-term… Iterated prisoners’ dilemma © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 18. With a probability ω the interaction continues time C C C C C C C P1 P2 C C C C C C C … … © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 19. How to deal with mistakes ? (intentional or not) Should we collect the compensation or continue the agreement? Should one take revenge or apologise and forgive? With a probability ω the interaction continues time C D C C C C C P1 P2 D D C C C C D … … © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 20. Individuals prefer to take revenge time C C C D C C D P1 P2 C C C C D D D … … 0.4 −0.2 0 0.2 0.4 a t )−Fr(NC,−,AllD) NN AllD PN AllD TFT abundancerelative tothedefectors 0.0 0.2 0.4 -0.2 log(noise) -3 -2 -1 D P-C-D P-C-TFT pay compen- sation δ © Tom Lenaerts, 2018 FutureICT2.0 Tallinn When apologising is not possible
  • 21. time C C C C C C D P1 P2 C C C C C C C … … Oeps! sorry really sorry no problem apology cost (γ) 0 2 4 6 8 −0.1 0 0.1 0.2 0.3 0.4 γ Fr(S)−Fr(NC,−,AllD,−) α=0.01 0 2 4 6 8 −0.1 0 0.1 0.2 0.3 0.4 γ Fr(S)−Fr(NC,−,AllD,−) α=0.1 (P,C,AllD,q=1) (P,C,AllD,q=0) (P,D,AllD,q=1) (A,D,AllD,q=1) abundancerelative tothedefectors 0.0 0.1 0.3 0.2 0.4 -0.1 0 2 4 6 8 D P-C-D+A P-C-D + NA apology cost P-D-D+A A-D-D+A These are like the fake players © Tom Lenaerts, 2018 FutureICT2.0 Tallinn When apologising is possible Apology needs to be sincere
  • 22. To err is humane, to forgive … Alexander Pope, An essay on Criticism (1711) evolutionary viable when the apology is sincere © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 23. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 24. Science=collaboration © Tom Lenaerts, 2018 FutureICT2.0 Tallinn The Anh Han (UK) Luis Moniz Pereira (P) Luis Martinez (I)
  • 25. © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 26.
  • 27. Produces equivalent results as in the mathematical model 0 0.5 1 1.5 2 2.5 3 3.5 4 4.5 5 x 10 −3b/c=1.5, =0.01 0 2 4 6 8 0 2 4 6 8 b/c=2, =0.01 0 2 4 6 8 0 2 4 6 8 b/c=4, =0.01 0 2 4 6 8 0 2 4 6 8 b/c=1.5, =0.1 0 2 4 6 8 0 2 4 6 8 b/c=2, =0.1 0 2 4 6 8 0 2 4 6 8 b/c=4, =0.1 0 2 4 6 8 0 2 4 6 8 Agent-based modelling An agent’s genotype is defined by: apology cost (γ) forgiveness threshold (τγ) © Tom Lenaerts, 2017 AISB in Bath
  • 28. S T R+1RR-1 P-1 P+1 P 7 3 3 7 0 1 1 0 C D C D T > R > P > S 7 3 3 7 1 0 0 1 C D C D T > R > S > P 3 7 7 3 0 1 1 0 C D C D R > T > P > S 3 7 7 3 1 0 0 1 C D C D R > T > S > P Space of social dilemmas © Tom Lenaerts, 2017 AISB in Bath
  • 29. S T R+1RR-1 P-1 P+1 P C D C D S-P R-S-T+P C D An evolutionary game theory perspective …. REPLICATOR DYNAMICS C D © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 30. Finite population dynamics D D C D CD DC DD t µ CD O 1-µ C C p=[1+eβ(f(D)-f(C))]-1 D C D CD DC DD t+1 D A moran process (birth-death process) CC selection strength © Tom Lenaerts, 2016 © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 31. Assuming that mutations are rare, we have either … n(C)=Z C C C C C CC CC C C C n(C)=0 D D D D DD DD D D D D n(C)=1 C D D D D DD DD D D D ... fixation probability ρ of C in D population n(C)=Z-1 C C C C C CD CC C C C fixation probability ρ of D in C population Finite population dynamics © Tom Lenaerts, 2016 © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 32. Which produces a reduced Markov chain For which the stationary distributions can be calculated = how likely it is to end up in either monomorphic state DC Fudenberg, D., & Imhof, L.A. (2006). Imitation processes with small mutations. J. Econ.Theo, 131,251–262. Imhof, L.A., Fudenberg, D., & Nowak, M.A. (2005). Evolutionary cycles of cooperation and defection. Proc Nat Acad Sci USA, 102(31), 10797–10800. ρCD ρDC Dynamics in finite populations x% y%=1-x © Tom Lenaerts, 2016 © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 33. Dynamics in finite populations 4 3 3 4 0 1 1 0 C DC D DC 1.6ρN 0.6ρN 26.5% 73.5% with β=0.01 10 -5 10 -4 10 -3 10 -2 10 -1 10 0 10 1 Selection strength (β) 0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1 Stationarydistribution C D for varying β ρN=drift=1/N How can we achieve cooperation? © Tom Lenaerts, 2016 © Tom Lenaerts, 2018 FutureICT2.0 Tallinn
  • 34. Rules for the evolution of cooperation Nowak, M.A. Five Rules for the Evolution of Cooperation. Science 314, 1560–1563 (2006). provided that b/c > 1 + (n/m) (5) Evolutionary Success Before proceeding to a comparative analysis of the five mechanisms, let me introduce some The entries denote the payoff for the row player. Without any mechanism for the evolution of cooperation, defectors dominate cooperators, which means a < g and b < d. A mechanism for the evolution of cooperation can change these inequalities. 1) If a > g, then cooperation is an evo- lutionarily stable strategy (ESS). An infinitely large population of cooperators cannot be in- vaded by defectors under deterministic selec- tion dynamics (32). 2) If a + b > g + d, then cooperators are risk-dominant (RD). If both strategies are ESS, then the risk-dominant strategy has the bigger basin of attraction. 3) If a + 2b > g + 2d, then cooperators are advantageous (AD). This concept is important for stochastic game dynamics in finite pop- ulations. Here, the crucial quantity is the fix- ation probability of a strategy, defined as the probability that the lineage arising from a single mutant of that strategy will take over the entire population consisting of the other strategy. An AD strategy has a fixation proba- bility greater than the inverse of the popu- lation size, 1/N. The condition can also be expressed as a 1/3 rule: If the fitness of the in- vading strategy at a frequency of 1/3 is greater than the fitness of the resident, then the fix- ation probability of the invader is greater than 1/N. This condition holds in the limit of weak selection (52). We have encountered evolution of cooperati mathematical formali mechanisms are very each theory is a simp coherent mathematica the derivation of all fi is that each mechanis game between two st payoff matrix (Table can derive the relevan of cooperation. For kin selection inclusive fitness prop (31). The relatedness Therefore, your payof to mine. A second me to a different matrix direct reciprocity, the while the defectors expected number of r tit-for-tat players coop tat versus always-defe first move and then iprocity, the probabili reputation is given b unless the reputation dicates a defector. A network reciprocity, i expected frequency o by a standard replicat formed payoff matrix the payoff matrices o Kin selection Network reciprocity Direct reciprocity Indirect reciprocity 1 r the five mechanisms, let me introduce some inequalities. 1) If a > g, then coope lutionarily stable strategy (E large population of cooperat vaded by defectors under de tion dynamics (32). 2) If a + b > g + d, the risk-dominant (RD). If bo ESS, then the risk-dominant bigger basin of attraction. 3) If a + 2b > g + 2d, the advantageous (AD). This con for stochastic game dynami ulations. Here, the crucial q ation probability of a strateg probability that the lineage single mutant of that strateg the entire population consis strategy. An AD strategy has bility greater than the inve lation size, 1/N. The condi expressed as a 1/3 rule: If the vading strategy at a frequenc than the fitness of the resid ation probability of the invad 1/N. This condition holds in selection (52). Kin selection Network reciprocity Direct reciprocity Indirect reciprocity 1 r Table 1. Each mechanism ca interaction between cooperato essary conditions for evolution Before proceeding to a comparative analysis of the five mechanisms, let me introduce some the evolution of cooperation ca inequalities. 1) If a > g, then cooperat lutionarily stable strategy (ESS large population of cooperators vaded by defectors under deter tion dynamics (32). 2) If a + b > g + d, then c risk-dominant (RD). If both ESS, then the risk-dominant s bigger basin of attraction. 3) If a + 2b > g + 2d, then advantageous (AD). This conce for stochastic game dynamics ulations. Here, the crucial quan ation probability of a strategy, probability that the lineage a single mutant of that strategy the entire population consistin strategy. An AD strategy has a bility greater than the inverse lation size, 1/N. The conditio expressed as a 1/3 rule: If the fi vading strategy at a frequency o than the fitness of the residen ation probability of the invader 1/N. This condition holds in th selection (52). Kin selection Network reciprocity Direct reciprocity Indirect reciprocity 1 r Table 1. Each mechanism can interaction between cooperators Network reciprocity Indirect reciprocity Group selection Cooperators Defectors Fig. 3. Five mechanisms for the evolution of cooperation. Kin selection operates when the probability that the lineage arising from a single mutant of that strategy will take over the entire population consisting of the other strategy. An AD strategy has a fixation proba- bility greater than the inverse of the popu- lation size, 1/N. The condition can also be expressed as a 1/3 rule: If the fitness of the in- vading strategy at a frequency of 1/3 is greater than the fitness of the resident, then the fix- ation probability of the invader is greater than 1/N. This condition holds in the limit of weak selection (52). tit-for-tat players cooperate tat versus always-defect co first move and then defec iprocity, the probability of reputation is given by q. unless the reputation of dicates a defector. A defe network reciprocity, it can expected frequency of coo by a standard replicator e formed payoff matrix (54) the payoff matrices of the Network reciprocity Group selection Cooperators Defectors Fig. 3. Five mechanisms for the evolution of cooperation. Kin selection operates when the donor and the recipient of an altruistic act are genetic relatives. Direct reciprocity requires re- peated encounters between the same two individ- uals. Indirect reciprocity is based on reputation; a helpful individual is more likely to receive help. Network reciprocity means that clusters of coop- erators outcompete defectors. Group selection is Table 1. Each mechanism can be described by a simple 2 × 2 payoff matri interaction between cooperators and defectors. From these matrices we can di essary conditions for evolution of cooperation. The parameters c and b denote, for the donor and the benefit for the recipient. For network reciprocity, we us [(b − c)k − 2c]/[(k + 1)(k − 2)]. All conditions can be expressed as the exceeding a critical value. See (53) for further explanations of the underlyin Cooperation is… 0 )1)(( rcbD cbrrcbC 0 )1/()( bD cwcbC 0)1( )1( qbD qccbC 0HbD cHcbC )())(( cnmcbnmcbC DC Kin selection Direct reciprocity Indirect reciprocity Network reciprocity Group rc b 1 rc b 1 rc b 1 r… w… q… k… n… wc b 1 w w c b 2 w w c b 23 qc b 1 q q c b 2 q q c b 23 k c b k c b k c b nb 1 nb 1 ESS RD AD nb 1 Payoff matrix Assortment © Tom Lenaerts, 2018 FutureICT2.0 Tallinn