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Evolution of Man
 
• The rise of man: - Fossils of Man’s ancestors are difficult 
to find, because only a few fossil exist. But we can trace 
Man’s evolution to some extent.
• Early Apes:-
• The primates first evolved in the Palaeocene epoch, about 
60 million years ago. Apes, such as Aegyptopithecus and 
Propliopithecus,  appeared  during  the  Oligocene  epoch, 
about 35 million years ago. 
• Towards the end of the Oligocene epoch, scientists believe 
that the ancestors of man and those of the true apes split 
into two separate groups. 
• Fossil  pieces  of  jaws  of  an  early  man  –  like  ape  called 
Ramapithecus  dating  from  about  10  million  years  ago, 
show several man like characteristics. 
• Australopithecines:-
• After Ramapithecus, Man’s evolution becomes rather 
indistinct, we know that the Australopithecines evolved 
during the Pliocene epoch and finally became extinct 
during the Pleistocene epoch. 
• Australopithecines included Australopithecus robustus, a 
large heavy boned type and Australopithecus afrianas also 
called Australopithecus grucilis, which is a lighter form. 
Some people believe that Man evolved from A. africanus. 
• But recent finds in East Africa suggest that early Man have 
lived  alongside  the  Australopithecines,  even  in  the  early 
Pliocene period (chart). 
• True Hominids:-
• By the beginning of the Pleistocene epoch, true hominids 
(man- like creatures) definitely existed. Homo habilis 
(“handy man”) is the name given to several fossils that 
date from this time. But some people believe that some of 
these finds belong to the Australopithecine group and that 
the rest are early forms of Homo erectus (upright ape 
man). 
• Even if Homo habelis did exist, he may have became 
extinct. A homo scull, known by its find number, 1470, 
found at Olduvai, Tanzania, may have been a more 
advanced form of man than Homo habilis. 
• Whatever their origins, the Homo erectus group was well 
established in the middle Pleistocene epoch. At least two 
lines of Homo sapiens evolved from it. 
• Homo sapiens:-
• Neandertal Man, or Homo sapiens neanderthalensis, who 
appeared around 100,000 years ago, became extinct at the 
end of the Pleistocene epoch. Modern Man, Homo sapiens
sapiens, appeared right at the end of the epoch. 
• But it is uncertain which of the earlier fossils of Homo
sapiens are on the direct line of evolution from Homo
erectus to modern man. Steinhein Man and Swanscombe 
man could be ancestors of both modern man and the 
Neanderthals. 
• Some  fossils,  such  as  those  from  Broken  hill  (now 
Kabwe), Zambia and from the Tabun cave at mt. Carmel 
Israel, show Neanderthal like features so they are probably 
not the ancestors of modern man.
• A hominid is any member of the biological family 
Hominidae. (the "great apes"), including the extinct and 
extant humans, chimpanzees, gorillas , and Orangutans
• . This classification has been revised
• several times in the last few decades. These various 
revisions have led to a varied use of the word "hominid":
• The original meaning of Hominidae referred only to the 
modern meaning of Hominina, including only humans and 
their closest relatives.
• The  meaning  of  the  taxon  changed  gradually,  leading  to 
the modern meaning of "hominid" in which it includes all 
great apes. The Primatological term is easily confused with 
a number of very similar words:
• A hominoid is a member of the superfamily Hominoidea: extant 
members are the lesser apes (gibbons) and great apes. 
• A hominid is a member of the family Hominidae: all of the 
great apes. 
• A hominine is a member of the subfamily Homininae: gorillas, 
chimpanzees, humans (excludes Orangutans). 
• A hominin is a member of the tribe Hominini: chimpanzees and 
humans. 
• A  Hominan  is  a  member  of  the  sub-tribe  Hominina:  modern 
humans and their extinct relatives. 
• A  human  is  a  member  of  Homo sapiens sapiens or  Homo
sapiens idaltu and  might  also  be  used  to  refer  to  any  extinct 
member  of  the  genus  Homo or  members  from  other  hominan 
genera. In more hypothetical and fictional contexts it might also 
refer to a being out of an other evolutionary lineage who looks 
very similar to a member of Homo sapiens sapiens. 
• A Humanoid is a vaguely human-shaped entity. 
• Certain morphologicalcharacteristics are still used 
conventionally to support the idea that hominid should only 
denote humans and human ancestors, namely bipedalism and 
large brains. 
• These points of departure between human beings and the other 
great apes are important, but according to geneticically based 
taxonomic classification, is not enough to divide us into separate 
families. 
• Genetics, rather than morphology, as the standard is now 
generally accepted as the critical test of relatedness and in this 
respect humans and the other great apes ought to be of the same 
family. 
• The terms hominid and "great ape" are now effectively 
coterminous. Anthropologists use the term to mean humans and 
their direct and near-direct bipedal ancestors. 
• Whether  the  critical  standard  should  be  strictly  genetic  or 
morphologic,  or  a  combination  of  the  two  is  yet  to  be 
determined.
• Hominini  is  the  tribe  of  Homininae  that  only  includes 
humans  (Homo),  chimpanzees  (Pan),  and  their  extinct 
ancestors. Members of the tribe are called  hominins (cf. 
Hominidae, "hominids").
• Through DNA comparison, scientists believe the 
Pan/Homo divergence was completed between 5.4 to 6.3 
million years ago, 
• After an unusual process of speciation that ranged over 
four million years. It is interesting to note that no fossil 
species on the Pan side of the split have been determined; 
all of the extinct genera listed to the right are ancestral to 
Homo, or are offshoots of such. 
• However, both Orrorin and Sahelanthropus existed around 
the  time  of  the  split,  and  so  may  be  ancestral  to  both 
humans and chimpanzees.
• In the proposal of Mann and Weiss (1996), the tribe Hominini 
includes Pan as well as Homo as separate subtribes. Homo (and, 
by inference, all bipedal apes) is by itself only in the subtribe 
Hominina, while Pan is in the Panina subtribe. 
• Scientific classification 
• Kingdom:  Animalia
• Phylum:  Chordata
• Class:  Mammalia
• Order:  Primates
• Superfamily:  Hominoidea
• Family:  Hominidae
• Subfamily:  Homininae
• Tribe:  Hominini
 
• WHY STUDY BONES
• Archaeologists study people in the past, so study of the
physical remains of actual people is fundamental.
• The study of human remains adds another dimension to the
past which study of the material culture cannot usually
provide, namely the health and physique of a population.
• The basic requirements of the archaeologist dictate the
major aspects of work carried out by the archaeological
human bone specialist: age and sex structure of a
population; physical size and appearance; general stresses
and strains of daily life; and diseases in the past.
• The techniques of analysing bones
• Introduction
• The following is an introduction to the work of the
archaeological human bone specialist, osteoarchaeologist,
or physical/biological anthropologist. The general
techniques are used widely by both archaeological and
forensic specialists in human remains. Further and more
detailed information is available in the books referenced at
the end of this page.
• Age
• The most basic distinction is between child and adult.
Although clearly children are generally smaller than
adults, this is not a reliable method of distinction beyond
the age of c.12 years.
The ends of the long bones and
parts of some other bones are
detached in children's skeletons.
This allows the bones to grow.
These detached parts are known as
epiphyses, and they all fuse at
different ages, from c.15 years
onwards.
• The stage of fusion of the bones can therefore be used to age an
adolescent. Before this, the most reliable method is to study the
developmental stage of the teeth.
• As a rough guide, the first permanent molar erupts at the age of six
years, the second at twelve years, and the third at c.18-21 years - you
need to know the difference between milk molars and permanent
molars first though! If the teeth are not present then the long bone
lengths are also a good indicator of age in younger children.
• Ageing skeletons becomes more difficult as the individual gets older.
Once all the bones have fused there is really no reliable simple method
to use.
• Tooth wear gives some indication, but only if members of the group
being studied were eating a fairly coarse diet. Other methods of
ageing, such as changes to the face of the pubic symphysis (the joint at
the front of the pelvis) or cranial suture closure, have largely been
discredited in recent years.
• The best we can do is to place skeletons into simple age categories -
young, middle-aged or old.
• Sex
• It is not generally possible to sex the skeleton of a child, unless it has passed
puberty and is starting to show some of the characteristics appropriate to its
sex.
• Adults are more easy to deal with, as long as you remember that humans are
continuously variable and there is not always an easy distinction to be made
between a small man and a large woman.
• The best indicators of sex in the skeleton are to be found in the pelvis. This
is because one of the major biological differences between men and women,
that of having babies, largely determines the shape of that part of the body.
• A woman's pelvis is wide and bowl-shaped whilst a man's is tall and narrow.
As the pelvis is made up of three bones (two innominates and the sacrum),
this can be seen in the width of the sacrum, the sciatic notch and the sub-
pubic angle, and sex can usually be determined even if part of the pelvis is
destroyed.
The next best indicator of sex is the skull. The face shows subtle
differences in the shape of the eye sockets (orbits), the angle of
the forehead is more vertical in women, the brow ridges and
mastoid processes are smaller, and the muscle markings are less
robust.
In fact this is the main criterion for determining sex if only the
long bones and a few fragments of skull remain - as a general
rule men tend to be more physically robust than women.
• Now compare these measurements with the chart
that follows. (Remember to take your
measurements in millimeters.)
Measureme
nts
Female
Probably
Female
Indefinite
sex
Probably
male
Male
Diameter of
head of
femur
<41.5 41.5 to 43.5 43.5 to 44.5 44.5 to 45.5 >45.5
Trochanteric
oblique
length
<390 390 to 405 405 to 430 430 to 450 >450
 –Evolution of Man
 –Evolution of Man
 –Evolution of Man
 –Evolution of Man
 –Evolution of Man
 –Evolution of Man

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–Evolution of Man

  • 2. • The rise of man: - Fossils of Man’s ancestors are difficult  to find, because only a few fossil exist. But we can trace  Man’s evolution to some extent. • Early Apes:- • The primates first evolved in the Palaeocene epoch, about  60 million years ago. Apes, such as Aegyptopithecus and  Propliopithecus,  appeared  during  the  Oligocene  epoch,  about 35 million years ago.  • Towards the end of the Oligocene epoch, scientists believe  that the ancestors of man and those of the true apes split  into two separate groups.  • Fossil  pieces  of  jaws  of  an  early  man  –  like  ape  called  Ramapithecus  dating  from  about  10  million  years  ago,  show several man like characteristics. 
  • 3.
  • 4. • Australopithecines:- • After Ramapithecus, Man’s evolution becomes rather  indistinct, we know that the Australopithecines evolved  during the Pliocene epoch and finally became extinct  during the Pleistocene epoch.  • Australopithecines included Australopithecus robustus, a  large heavy boned type and Australopithecus afrianas also  called Australopithecus grucilis, which is a lighter form.  Some people believe that Man evolved from A. africanus.  • But recent finds in East Africa suggest that early Man have  lived  alongside  the  Australopithecines,  even  in  the  early  Pliocene period (chart). 
  • 5. • True Hominids:- • By the beginning of the Pleistocene epoch, true hominids  (man- like creatures) definitely existed. Homo habilis  (“handy man”) is the name given to several fossils that  date from this time. But some people believe that some of  these finds belong to the Australopithecine group and that  the rest are early forms of Homo erectus (upright ape  man).  • Even if Homo habelis did exist, he may have became  extinct. A homo scull, known by its find number, 1470,  found at Olduvai, Tanzania, may have been a more  advanced form of man than Homo habilis.  • Whatever their origins, the Homo erectus group was well  established in the middle Pleistocene epoch. At least two  lines of Homo sapiens evolved from it. 
  • 6. • Homo sapiens:- • Neandertal Man, or Homo sapiens neanderthalensis, who  appeared around 100,000 years ago, became extinct at the  end of the Pleistocene epoch. Modern Man, Homo sapiens sapiens, appeared right at the end of the epoch.  • But it is uncertain which of the earlier fossils of Homo sapiens are on the direct line of evolution from Homo erectus to modern man. Steinhein Man and Swanscombe  man could be ancestors of both modern man and the  Neanderthals.  • Some  fossils,  such  as  those  from  Broken  hill  (now  Kabwe), Zambia and from the Tabun cave at mt. Carmel  Israel, show Neanderthal like features so they are probably  not the ancestors of modern man.
  • 7. • A hominid is any member of the biological family  Hominidae. (the "great apes"), including the extinct and  extant humans, chimpanzees, gorillas , and Orangutans • . This classification has been revised • several times in the last few decades. These various  revisions have led to a varied use of the word "hominid": • The original meaning of Hominidae referred only to the  modern meaning of Hominina, including only humans and  their closest relatives. • The  meaning  of  the  taxon  changed  gradually,  leading  to  the modern meaning of "hominid" in which it includes all  great apes. The Primatological term is easily confused with  a number of very similar words:
  • 8. • A hominoid is a member of the superfamily Hominoidea: extant  members are the lesser apes (gibbons) and great apes.  • A hominid is a member of the family Hominidae: all of the  great apes.  • A hominine is a member of the subfamily Homininae: gorillas,  chimpanzees, humans (excludes Orangutans).  • A hominin is a member of the tribe Hominini: chimpanzees and  humans.  • A  Hominan  is  a  member  of  the  sub-tribe  Hominina:  modern  humans and their extinct relatives.  • A  human  is  a  member  of  Homo sapiens sapiens or  Homo sapiens idaltu and  might  also  be  used  to  refer  to  any  extinct  member  of  the  genus  Homo or  members  from  other  hominan  genera. In more hypothetical and fictional contexts it might also  refer to a being out of an other evolutionary lineage who looks  very similar to a member of Homo sapiens sapiens.  • A Humanoid is a vaguely human-shaped entity. 
  • 9. • Certain morphologicalcharacteristics are still used  conventionally to support the idea that hominid should only  denote humans and human ancestors, namely bipedalism and  large brains.  • These points of departure between human beings and the other  great apes are important, but according to geneticically based  taxonomic classification, is not enough to divide us into separate  families.  • Genetics, rather than morphology, as the standard is now  generally accepted as the critical test of relatedness and in this  respect humans and the other great apes ought to be of the same  family.  • The terms hominid and "great ape" are now effectively  coterminous. Anthropologists use the term to mean humans and  their direct and near-direct bipedal ancestors.  • Whether  the  critical  standard  should  be  strictly  genetic  or  morphologic,  or  a  combination  of  the  two  is  yet  to  be  determined.
  • 10. • Hominini  is  the  tribe  of  Homininae  that  only  includes  humans  (Homo),  chimpanzees  (Pan),  and  their  extinct  ancestors. Members of the tribe are called  hominins (cf.  Hominidae, "hominids"). • Through DNA comparison, scientists believe the  Pan/Homo divergence was completed between 5.4 to 6.3  million years ago,  • After an unusual process of speciation that ranged over  four million years. It is interesting to note that no fossil  species on the Pan side of the split have been determined;  all of the extinct genera listed to the right are ancestral to  Homo, or are offshoots of such.  • However, both Orrorin and Sahelanthropus existed around  the  time  of  the  split,  and  so  may  be  ancestral  to  both  humans and chimpanzees.
  • 11. • In the proposal of Mann and Weiss (1996), the tribe Hominini  includes Pan as well as Homo as separate subtribes. Homo (and,  by inference, all bipedal apes) is by itself only in the subtribe  Hominina, while Pan is in the Panina subtribe.  • Scientific classification  • Kingdom:  Animalia • Phylum:  Chordata • Class:  Mammalia • Order:  Primates • Superfamily:  Hominoidea • Family:  Hominidae • Subfamily:  Homininae • Tribe:  Hominini  
  • 12.
  • 13.
  • 14.
  • 15.
  • 16.
  • 17.
  • 18.
  • 19.
  • 20.
  • 21. • WHY STUDY BONES • Archaeologists study people in the past, so study of the physical remains of actual people is fundamental. • The study of human remains adds another dimension to the past which study of the material culture cannot usually provide, namely the health and physique of a population. • The basic requirements of the archaeologist dictate the major aspects of work carried out by the archaeological human bone specialist: age and sex structure of a population; physical size and appearance; general stresses and strains of daily life; and diseases in the past.
  • 22. • The techniques of analysing bones • Introduction • The following is an introduction to the work of the archaeological human bone specialist, osteoarchaeologist, or physical/biological anthropologist. The general techniques are used widely by both archaeological and forensic specialists in human remains. Further and more detailed information is available in the books referenced at the end of this page. • Age • The most basic distinction is between child and adult. Although clearly children are generally smaller than adults, this is not a reliable method of distinction beyond the age of c.12 years.
  • 23. The ends of the long bones and parts of some other bones are detached in children's skeletons. This allows the bones to grow. These detached parts are known as epiphyses, and they all fuse at different ages, from c.15 years onwards.
  • 24. • The stage of fusion of the bones can therefore be used to age an adolescent. Before this, the most reliable method is to study the developmental stage of the teeth. • As a rough guide, the first permanent molar erupts at the age of six years, the second at twelve years, and the third at c.18-21 years - you need to know the difference between milk molars and permanent molars first though! If the teeth are not present then the long bone lengths are also a good indicator of age in younger children. • Ageing skeletons becomes more difficult as the individual gets older. Once all the bones have fused there is really no reliable simple method to use. • Tooth wear gives some indication, but only if members of the group being studied were eating a fairly coarse diet. Other methods of ageing, such as changes to the face of the pubic symphysis (the joint at the front of the pelvis) or cranial suture closure, have largely been discredited in recent years. • The best we can do is to place skeletons into simple age categories - young, middle-aged or old.
  • 25. • Sex • It is not generally possible to sex the skeleton of a child, unless it has passed puberty and is starting to show some of the characteristics appropriate to its sex. • Adults are more easy to deal with, as long as you remember that humans are continuously variable and there is not always an easy distinction to be made between a small man and a large woman. • The best indicators of sex in the skeleton are to be found in the pelvis. This is because one of the major biological differences between men and women, that of having babies, largely determines the shape of that part of the body. • A woman's pelvis is wide and bowl-shaped whilst a man's is tall and narrow. As the pelvis is made up of three bones (two innominates and the sacrum), this can be seen in the width of the sacrum, the sciatic notch and the sub- pubic angle, and sex can usually be determined even if part of the pelvis is destroyed.
  • 26. The next best indicator of sex is the skull. The face shows subtle differences in the shape of the eye sockets (orbits), the angle of the forehead is more vertical in women, the brow ridges and mastoid processes are smaller, and the muscle markings are less robust. In fact this is the main criterion for determining sex if only the long bones and a few fragments of skull remain - as a general rule men tend to be more physically robust than women.
  • 27.
  • 28.
  • 29.
  • 30.
  • 31.
  • 32.
  • 33.
  • 34.
  • 35.
  • 36. • Now compare these measurements with the chart that follows. (Remember to take your measurements in millimeters.) Measureme nts Female Probably Female Indefinite sex Probably male Male Diameter of head of femur <41.5 41.5 to 43.5 43.5 to 44.5 44.5 to 45.5 >45.5 Trochanteric oblique length <390 390 to 405 405 to 430 430 to 450 >450