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Introduction 
Data simulations can reveal larger pictures about our 
world. My program simulates the relationship between 
Bruchid Beetles (predator) and Panama palm tree endocarps 
(prey). By running simulations with parameters that match 
the generation time and offspring count of my studied 
populations, I can determine the effect of Beetle predation 
on clumping patterns. 
I chose to focus my simulations on Panama palm trees 
because:
• The seeds retain scars from predation events, such as claw 
marks and emergence holes.
• Data from endocarps is more reliable because it’s easier to 
count and track all of the endocarps over multiple 
generations.
• Significant study of palms in Panama has given enough 
characterization about their predators to create complex 
simulations.
I chose to use the Bruchid Beetles as predators because:
• They leave large emergence holes.
• Their generation time is the same as one season for 
seedlings.
Simulating the Effect of Predator‐Prey Ratios on Clumping in 
Seedling Distributions of Panama Palm Trees
Methods
My simulation features:
• Random generation of tree locations
• Offspring distributed with an inverse logarithmic 
probability density curve
• Beetles taking Random Walks
• Multiple generations of Beetles and trees
• A function to recognize the strength of clumping 
within a population
Results
I’ve developed the following skillsets:
• Programming in R statistical software
• Using Git Version control software
Hypothesis:
• I hypothesize that small Beetle‐seedling 
ratios will not affect clumping, but large 
ones will significantly reduce clumping.
Figure 2 – Simulations of Seed Dispersal
Figure 3 – Predation of 
Endocarps by Bruchid Beetles
Figure 1‐ Versatile User Interface
Justin Tirrell
Utah State University
Biology
tirrelljustin95@gmail.com
Left: Coordinate plane showing the locations of trees, Beetles, Seeds, and Dead seeds.
Right: Data table representing the factors to be included in future models. 
Program created using R Statistical Software
Abstract
Palm trees provide a unique opportunity to study what conditions optimize 
the probability that a seed will grow successfully. The seeds of palm trees, 
endocarps, are large and easy to locate. When they don't grow, predators 
leave marks on them that tell the story of their fate. The focus of my 
experiment is to determine how the current distribution pattern of parent 
palm trees in Panama palm trees affects the future distribution of seedlings. I 
have programmed a versatile model that takes the assumption that Bruchid
Beetles are the sole predators acting on the seeds, and that these fall from 
the trees in an inverse logarithmic density pattern. The Beetles are 
programmed to move to a random seed within an arbitrary distance of their 
start point. If no seeds are near enough to them, then they starve. I 
hypothesize that the Beetles will decrease clumping within five generations.1
Justin Tirrell, Eric Sodja, Cole Carlson, Noelle Beckman
Department of Biology, Utah State University.
Objectives
Simulations in R Statistical Software will determine: 
• How the ratio of Bruchid Beetles to palm trees may be 
expected to influence the distribution pattern of 
mature palm trees in succeeding generations.
• How the initial distribution of palm trees may be 
expected to influence the distribution pattern of 
mature palm trees in succeeding generations.
Figure 3 Image Attributions:
“Attalea sp. , fruits and seeds” by Roger Culos
“Male Callosobruchus maculatus” by limbatus
“Attalea brasiliensis” by  João Medeiros
Palm trees in Panama drop 
large seeds called endocarps.
Bruchid Beetles lay their 
eggs on the endocarps. 
Beetle larvae prey on 
the endocarps, leaving 
large emergence holes 
when mature.
User Interface:  This simulation can take input from the user that describes the dynamics of the existing population. 
Citations:
1. Wright, S. J. (1983). The Dispersion of Eggs by a Bruchid Beetle among Scheelea Palm Seeds and the Effect of Distance to the Parent Palm.
Ecology, 64(5), 1016–1021. https://doi.org/10.2307/1937808

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  • 1. Introduction  Data simulations can reveal larger pictures about our  world. My program simulates the relationship between  Bruchid Beetles (predator) and Panama palm tree endocarps  (prey). By running simulations with parameters that match  the generation time and offspring count of my studied  populations, I can determine the effect of Beetle predation  on clumping patterns.  I chose to focus my simulations on Panama palm trees  because: • The seeds retain scars from predation events, such as claw  marks and emergence holes. • Data from endocarps is more reliable because it’s easier to  count and track all of the endocarps over multiple  generations. • Significant study of palms in Panama has given enough  characterization about their predators to create complex  simulations. I chose to use the Bruchid Beetles as predators because: • They leave large emergence holes. • Their generation time is the same as one season for  seedlings. Simulating the Effect of Predator‐Prey Ratios on Clumping in  Seedling Distributions of Panama Palm Trees Methods My simulation features: • Random generation of tree locations • Offspring distributed with an inverse logarithmic  probability density curve • Beetles taking Random Walks • Multiple generations of Beetles and trees • A function to recognize the strength of clumping  within a population Results I’ve developed the following skillsets: • Programming in R statistical software • Using Git Version control software Hypothesis: • I hypothesize that small Beetle‐seedling  ratios will not affect clumping, but large  ones will significantly reduce clumping. Figure 2 – Simulations of Seed Dispersal Figure 3 – Predation of  Endocarps by Bruchid Beetles Figure 1‐ Versatile User Interface Justin Tirrell Utah State University Biology tirrelljustin95@gmail.com Left: Coordinate plane showing the locations of trees, Beetles, Seeds, and Dead seeds. Right: Data table representing the factors to be included in future models.  Program created using R Statistical Software Abstract Palm trees provide a unique opportunity to study what conditions optimize  the probability that a seed will grow successfully. The seeds of palm trees,  endocarps, are large and easy to locate. When they don't grow, predators  leave marks on them that tell the story of their fate. The focus of my  experiment is to determine how the current distribution pattern of parent  palm trees in Panama palm trees affects the future distribution of seedlings. I  have programmed a versatile model that takes the assumption that Bruchid Beetles are the sole predators acting on the seeds, and that these fall from  the trees in an inverse logarithmic density pattern. The Beetles are  programmed to move to a random seed within an arbitrary distance of their  start point. If no seeds are near enough to them, then they starve. I  hypothesize that the Beetles will decrease clumping within five generations.1 Justin Tirrell, Eric Sodja, Cole Carlson, Noelle Beckman Department of Biology, Utah State University. Objectives Simulations in R Statistical Software will determine:  • How the ratio of Bruchid Beetles to palm trees may be  expected to influence the distribution pattern of  mature palm trees in succeeding generations. • How the initial distribution of palm trees may be  expected to influence the distribution pattern of  mature palm trees in succeeding generations. Figure 3 Image Attributions: “Attalea sp. , fruits and seeds” by Roger Culos “Male Callosobruchus maculatus” by limbatus “Attalea brasiliensis” by  João Medeiros Palm trees in Panama drop  large seeds called endocarps. Bruchid Beetles lay their  eggs on the endocarps.  Beetle larvae prey on  the endocarps, leaving  large emergence holes  when mature. User Interface:  This simulation can take input from the user that describes the dynamics of the existing population.  Citations: 1. Wright, S. J. (1983). The Dispersion of Eggs by a Bruchid Beetle among Scheelea Palm Seeds and the Effect of Distance to the Parent Palm. Ecology, 64(5), 1016–1021. https://doi.org/10.2307/1937808