3. The plant body has a hierarchy of
organs, tissues, and cells
Like multicellular animals, plants have organs
composed of different tissues, which in turn are
composed of cells.
Three Basic Plant Organs (Roots, Stems and
Leaves)
4. ORGANS
distinct functional structure consisting of multiple
types of tissues (Dermal, Vascular and Ground
tissues)
TISSUES
collection of one or more cell types that performs
a specific function within an organ
5. Each of these three
categories forms a system
(Roots system and
Shoots System)
Roots rely on sugar
produced by
photosynthesis in the
shoot system.
Shoots rely on water and
minerals absorbed by the
root system.
7. Structures found in plant cells but
not in animals cells.
LARGE CENTRAL VACUOLE –
surrounded by its own membrane
and contains water and dissolved
substances. Its primary role is to
maintain pressure against the
inside of the cell wall, giving the
cell shape and helping to support
the plant.
8. Cell wall – located outside
the cell membrane. It
consists mainly of cellulose
and lignin, which makes it
more rigid. The cell wall
shapes, supports, and
protects the cell. It prevents
the cell from absorbing too
much water and bursting. It
also keeps large, damaging
molecules out of the cell.
9. Platids are membrane-
bound organelles with
their own DNA.
Chloroplasts contain
the green pigment
chlorophyll and carry out
photosynthesis.
Chromoplasts make
and store other
pigments. They give
flower petals their bright
colors.
11. Parenchyma cells in a privet
(Ligustrum) leaf (LM)
25 ÎĽm
Parenchyma Cells
• have thin primary (1o) cell walls without a
secondary (2o) cell wall
• the least differentiated plant cell type
• the most
metabolically active
plant cell type
• are capable of
undergoing cell
division and further
differentiation
12. Collenchyma Cells
• flexible support in newly formed shoot
provide structures without restraining
growth
• flexible 1o cell
walls with
irregular
2o wall
thickening
Collenchyma cells
(in Helianthus stem) (LM) 5 ÎĽm
13. Sclerenchyma Cells
• provide rigid support due to thick 2o cell walls
containing lignin that are dead at maturity
5 ÎĽm
Sclereid cells in pear (LM)
Cell wall
Fiber cells (cross section from ash tree) (LM)
25 ÎĽm
• 2 types of
sclerenchyma
cells:
• sclereid cells
with very thick
2o cell walls
• long and
slender fiber
cells arranged
in threads
14. Water-Conducting Xylem Cells
2 types of xylem
cells, both of
which are dead at
maturity:
TRACHEIDS
100 ÎĽm
Tracheids and vessels
(colorized SEM)
Vessel Tracheids
Perforation
plate
Vessel
element
Vessel elements, with
perforated end walls
Tracheids
Pits
• found in all xylem
vessels
• long, thin with
tapered ends
VESSEL ELEMENTS
• wider, less tapered
• perforated ends
15. Sugar-Conducting Phloem Cells
2 types of phloem cells, both of which are
alive at maturity:
SIEVE CELLS
• found in seedless
vascular plants &
gymnosperms
SIEVE-TUBE ELEMENTS
• cells that form sieve
tubes in angiosperms
• have sieve plates
between elements &
supporting
companion cells
19. An overview of primary and secondary growth
Shoot tip (shoot
apical meristem
and young leaves)
Lateral meristems:
Axillary bud
meristem
Vascular cambium
Cork cambium
Root apical
meristems
Primary growth in stems
Epidermis
Cortex
Primary phloem
Primary xylem
Pith
Secondary growth in stems
Periderm
Cork
cambium
Cortex
Primary
phloem
Secondary
phloem
Pith
Primary
xylem
Secondary
xylem
Vascular cambium
20. Intercalary meristems - occur
only in monocots, at the
bases of leaf blades and at
nodes (the areas where
leaves attach to a stem). This
tissue enables the monocot
leaf blade to increase in
length from the leaf base; for
example, it allows lawn grass
leaves to elongate even after
repeated mowing.
21. •Meristems produce cells that quickly differentiate, or
specialize, and become permanent tissue. Such cells
take on specific roles and lose their ability to divide
further.
•They differentiate into three main types: dermal,
vascular, and ground tissue.
•Dermal tissue covers and protects the plant.
•Vascular tissue transports water, minerals, and
sugars to different parts of the plant.
•Ground tissue serves as a site for photosynthesis,
provides a supporting matrix for the vascular tissue,
and helps to store water and sugars.
22.
23. •Secondary tissues are either simple (composed of
similar cell types) or complex (composed of different cell
types).
•Dermal tissue, for example, is a simple tissue that covers
covers the outer surface of the plant and controls gas
exchange. Vascular tissue is an example of a complex
tissue, and is made of two specialized conducting tissues:
xylem and phloem.
•Xylem tissue transports water and nutrients from the
roots to different parts of the plant, and includes three
different cell types: vessel elements and tracheids (both
(both of which conduct water), and xylem parenchyma.
•Phloem tissue transports organic compounds from the
site of photosynthesis to other parts of the plant,
consists of four different cell types: sieve cells (which
conduct photosynthates), companion cells, phloem
parenchyma, and phloem fibers.
•Unlike xylem conducting cells, phloem conducting cells
are alive at maturity. The xylem and phloem always lie
adjacent to each other.
•In stems, the xylem and the phloem form a structure
called a vascular bundle; in roots, this is termed the
vascular stele or vascular cylinder.
24. •Meristems produce cells that quickly
differentiate, or specialize, and become
permanent tissue. Such cells take on
specific roles and lose their ability to divide
further.
•They differentiate into three main types:
dermal, vascular, and ground tissue.
25. 3 Basic Plant Tissue Types
Dermal tissue
• outer, protective covering
of the plant
Vascular tissue
• transports water,
minerals, and sugars to
different parts of the plant
Ground tissue
• serves as a site for
photosynthesis, provides a
supporting matrix for the
vascular tissue, and helps to
store water and sugars.
Dermal
tissue
Ground
tissue Vascular
tissue
each of these tissues forms
a continuous tissue system
throughout the plant
26. More on Dermal Tissue…
In nonwoody plants and structures (e.g., leaves) the
dermal tissue is epidermis.
• epidermis is frequently covered with a waxy cuticle to
minimize water loss
• some plants also have trichomes in epidermal tissue
which provide protection from water loss, intense light
and insects
In woody plants the epidermis develops into a
protective laver called periderm (part of the bark).
Trichomes
300
ÎĽm
27. ď‚· The epidermis of a leaf
also contains openings
known as stomata,
through which the
exchange of gases takes
place.
ď‚· Two cells, known as guard
cells, surround each leaf
stoma, controlling its
opening and closing and
thus regulating the uptake
of carbon dioxide and the
release of oxygen and
water vapor.
28. More on Vascular Tissue…
Plant vascular tissue consists of phloem & xylem.
Xylem
• transports water & minerals upward from the root
system to the organs and tissues of the shoot system
Phloem
• transports photosynthetic
products (e.g., sugars)
downward to the roots and
other parts of the plant
Phloem & xylem are organized
into vascular bundles or
cylinders called steles.
29. VASCULAR TISSUE
ď‚· When the stem is viewed in
cross section, the vascular
bundles of dicot stems are
arranged in a ring. In plants
with stems that live for more
than one year, the individual
bundles grow together and
produce the characteristic
growth rings.
ď‚· In monocot stems, the
vascular bundles are
randomly scattered
throughout the ground
tissue.
30. Water-Conducting Cells of the Xylem
ď‚· Xylem tissue has three types of cells:
xylem parenchyma, tracheids, and
vessel elements.
ď‚· Tracheids are xylem cells with thick
secondary cell walls that are lignified.
ď‚· Water moves from one tracheid to
another through regions on the side
walls known as pits, where secondary
walls are absent.
ď‚·Vessel elements are xylem cells with
thinner walls; they are shorter than
tracheids. Each vessel element is
connected to the next by means of a
perforation plate at the end walls of
the element. Water moves through
the perforation plates to travel up the
plant.
31. Differentiated Plant Cells in the Xylem - Dead at Maturity
Perforation
plate
Vessel
element
Vessel elements, with
perforated end walls
Tracheids
Pits
Tracheids and vessels
(colorized SEM)
Vessel Tracheids 100 µm
32. Sugar-Conducting Cells of the Phloem
ď‚· Phloem tissue is composed of
sieve-tube cells, companion
phloem parenchyma, and
fibers.
ď‚· Sieve-tube elements are alive at
functional maturity, though they
lack organelles.
ď‚· Sieve plates are the porous end
walls that allow fluid to flow
between cells along the sieve
tube.
ď‚· Each sieve-tube element has a
companion cell whose nucleus
and ribosomes serve both cells.
33. Differentiated
Plant Cells
Sieve-tube element (left)
and companion cell:
cross section (TEM)
3 µm
Sieve-tube elements:
longitudinal view (LM)
Sieve plate
Companion
cells
Sieve-tube
elements
Plasmodesma
Sieve
plate
Nucleus of
companion
cells
Sieve-tube elements:
longitudinal view Sieve plate with pores (SEM)
10 µm
30 µm
36. More on Ground Tissue…
Tissues that are not dermal or vascular are ground
tissue which come in 2 general types.
Pith
• ground tissue found
internal to the
vascular tissue
Cortex
• ground tissue found
between the dermal
and vascular tissue
Ground tissues include cells involved in storage,
transport, structural support and photosynthesis.
39. PLANTS ORGANS
SHOOT
SYSTEM
ď‚·The shoot system consists of
two portions: the vegetative
(non-reproductive) parts of the
plant, such as the leaves and the
stems, and the reproductive
parts of the plant, which include
flowers and fruits.
ROOT
SYSTEM
• The root system, which
supports the plants and
absorbs water and minerals, is
usually underground.
40. Three Basic Plant Organs
Plant organs evolved to obtain nutrients, water
and energy on land – below & above ground
ROOTS
• absorb water, minerals and other nutrients from
the soil
• anchor & support plant in the ground
STEMS
• structural support of plant above ground
• transport of water & nutrients throughout the
plant
LEAVES
• harvesting light & CO2 for photosynthesis
41. Stem Structure and Function
Stem structures include:
points of leaf attachment
called nodes
internodes – the stems
between each node
apical buds at the shoot tips
where growth occurs
axillary buds which give rise to
lateral branches, thorns or flowers
The petiole is the stalk
connecting the leaf to the stem
42. STEM ANATOMY
The stem and other plant organs arise
from the ground tissue, and are primarily
made up of simple tissues formed from
three types of parenchyma, collenchyma,
and sclerenchyma cells
43. •
PARENCHYMA CELLS are
the most common plant cells.
•They are found in the stem,
the root, the inside of the leaf,
and the pulp of the fruit.
•Parenchyma cells are
responsible for metabolic
functions, such as
photosynthesis, and they help
repair and heal wounds.
•Some parenchyma cells also
store starch.
44. ď‚· COLLENCHYMA CELLS
are elongated cells with
unevenly thickened
walls.
ď‚· They provide structural
support, mainly to the
stem and leaves. These
cells are alive at maturity
and are usually found
below the epidermis.
 The “strings” of a celery
stalk are an example of
collenchyma cells.
45. ď‚· SCLERENCHYMA CELLS also
provide support to the plant, but
unlike collenchyma cells, many
of them are dead at maturity.
ď‚· There are two types of
sclerenchyma cells: fibers and
sclereids. Both types have
secondary cell walls that are
thickened with deposits of lignin,
an organic compound that is a
key component of wood.
ď‚· Fibers are long, slender cells;
sclereids are smaller-sized.
Sclereids give pears their gritty
texture.
ď‚· Humans use sclerenchyma
fibers to make linen and rope.
46. Evolutionary Adaptations of Stems
Stems can be modified to serve a variety of
functions:
• rhizomes which grow just beneath
the soil surface and give rise to
vertical shoots from axillary buds
• stolons that function as “runners”
along the soil surface giving rise to
new plantlets
• tubers that serve
as storage “sinks”
for carbohydrates
Root
Rhizome
Rhizomes
Stolons
Stolo
Tubers
47.
48. • Some aerial modifications of stems are tendrils and
thorns.
• Tendrils seek support by climbing on other surfaces.
• Thorns are modified branches appearing as sharp
outgrowths that protect the plant
64. CO2
O2
H2O
Minerals
ROOTS
ď‚· The roots have three
major functions:
1.anchoring the plant to
the soil,
2.absorbing water and
minerals and
transporting them
upwards,
3.storing the products of
photosynthesis.
*over-watering can suffocate a plant!
65. ď‚· Some roots are modified to absorb
moisture and exchange gases.
ď‚· Most roots are underground.
ď‚· Some plants, however, also have
adventitious roots, which emerge above
the ground from the shoot.
66. TYPES OF ROOTS SYSTEM
DICOTS
A tap root system has a main
root that grows down vertically,
and from which many smaller
lateral roots arise.
A tap root system penetrates
deep into the soil.
MONOCOTS
A fibrous root system is
located closer to the soil
surface, and forms a dense
network of roots that also
helps prevent soil erosion.
67. ď‚· Some plants have a combination of tap
roots and fibrous roots.
ď‚· Plants that grow in dry areas often have
deep root systems, whereas plants
growing in areas with abundant water
are likely to have shallower root
systems.
68. ROOT GROWTH AND ANATOMY
ď‚· Root growth begins with seed
germination.
ď‚· When the plant embryo emerges
from the seed, the radicle of the
embryo forms the root system.
ď‚· The tip of the root is protected by the
root cap, a structure exclusive to
roots and unlike any other plant
structure.
ď‚· The root cap is continuously replaced
because it gets damaged easily as the
root pushes through soil.
69. ď‚· The root tip can be divided into three
zones: a zone of cell division, a zone
of elongation, and a zone of
maturation and differentiation.
ď‚· The zone of cell division is closest to
the root tip; it is made up of the
actively dividing cells of the root
meristem.
ď‚· The zone of elongation is where the
newly formed cells increase in length,
thereby lengthening the root.
ď‚· Beginning at the first root hair is the
zone of cell maturation where the
cells begin to differentiate into special
cell types.
70. ď‚· Inside the root, the ground tissue forms two regions:
the cortex and the pith.
ď‚· Both regions include cells that store photosynthetic
products.
ď‚· The cortex is between the epidermis and the vascular
tissue, whereas the pith lies between the vascular
tissue and the center of the root.
71. ď‚· The vascular tissue in the root is arranged in the inner
portion of the root, which is called the stele.
ď‚· A layer of cells known as the endodermis separates the
stele from the ground tissue in the outer portion of
the root.
ď‚· The endodermis is exclusive to roots, and serves as a
checkpoint for materials entering the root’s vascular
system.
72. ď‚· A waxy substance called suberin is present on the
walls of the endodermal cells.
ď‚· This waxy region, known as the Casparian strip,
forces water and solutes to cross the plasma
membranes of endodermal cells instead of slipping
between the cells.
ď‚· This ensures that only materials required by the root
pass through the endodermis, while toxic substances
and pathogens are generally excluded.
73.  The outermost cell layer of the root’s vascular tissue is
the pericycle, an area that can give rise to lateral
roots.
ď‚· In dicot roots, the xylem and phloem of the stele are
arranged alternately in an X shape, whereas in
monocot roots, the vascular tissue is arranged in a
ring around the pith.
74. ROOT MODIFICATIONS
ď‚· Root structures may be modified for specific
purposes. For example, some roots are bulbous and
store starch.
ď‚· Aerial roots and prop roots are two forms of
aboveground roots that provide additional support to
anchor the plant.
ď‚· Tap roots, such as carrots, turnips, and beets, are
examples of roots that are modified for food storage
79. ď‚· Epiphytic roots enable a plant to
grow on another plant.
ď‚· For example, the epiphytic roots
of orchids develop a spongy
tissue to absorb moisture.
81. Meristem Tissue
Unlike animals, plants are capable of indeterminate
growth – growth throughout the life of the plant.
This unlimited growth potential is due to meristem
tissue – a special, undifferentiated tissue with
unlimited replicative potential.
• in contrast, animals and some plant structures (e.g.,
flowers, thorns) exhibit determinate growth in which
they stop growing when they reach a certain size
There are 2 types of meristems:
• APICAL MERISTEM • LATERAL MERISTEM
82. Root apical
meristems
Axillary bud
meristem
Shoot tip
(shoot apical
meristem and
young leaves)
Apical meristem is located at
the tips of roots and shoots
and is responsible for growth
in length – what is called
primary growth.
• in non-woody (herbaceous)
plants, most if not all growth is
due to apical meristem
• in woody plants (e.g., trees),
there is also growth in width,
what is referred to as
secondary growth…
84. Primary Growth of Roots
100 ÎĽm
Mitotic
cells
Zone of cell
division
(including
apical
meristem)
Root cap
Zone of
elongation
Zone of
differentiation
Dermal
Ground
Vascular
Vascular cylinder
Cortex
Epidermis
Root hair
Root tips have a protective, non-dividing root cap.
Just underneath the
root cap is the
Zone of Cell Division
which contains the
apical meristem cells.
Beyond the Zone of
Cell Division are 2
zones in successive
developmental stages:
Zone of Elongation
• pushes root into soil
Zone of Differentiation
• cells adopt specific fates
85. 100 ÎĽm
(a) Root with xylem and phloem in
the center (typical of eudicots)
Xylem
Phloem
Dermal
Ground
Vascular
Pericycle
Core of
parenchyma
cells
Vascular cylinder
Endodermis
Cortex
Epidermis
Endodermis
Pericycle
Xylem
Phloem
70 ÎĽm
In most eudicot
roots, there is a
central vascular
cylinder (stele)
with a “X-shaped”
arrangement of
xylem as seen in
cross section with
phloem filling in
between the
“arms” of the X.
Eudicot
Roots
87. Lateral Root Growth
100 ÎĽm Epidermis
Lateral root
Emerging
lateral
root
Cortex
Vascular
cylinder
1
Pericycle
2 3
Lateral root growth occurs from the meristematic
pericycle, the outermost layer of cells in the
vascular cylinder just inside the endodermis, the
innermost layer of cortex.
88. Primary Growth of Shoots
Leaf primordia
Young leaf
Shoot apical
meristem
Developing
vascular
strand
Axillary bud
meristems
0.25 mm
Primary growth of shoot structures occurs from:
• apical meristem
which lengthens
the stem and
gives rise to leaf
primordia
• axial meristem
which gives rise
to new branches
from the main
stem
89. Organization of Eudicot Stems
1 mm
Dermal
Ground
Vascular
(a) Cross section of stem with
vascular bundles forming a
ring (typical of eudicots) (LM)
Cortex
Pith
Vascular
bundle
Epidermis
Xylem
Phloem
Sclerenchyma
(fiber cells) Ground tissue
connecting
pith to cortex
In most eudicot
stems, the vascular
tissue consists of
bundles of phloem
and xylem arranged
in a ring around the
central pith tissue.
• the xylem is always
located inside the
phloem adjacent to
the pith
90. 1 mm
Dermal
Ground
Vascular
(b) Cross section of stem with
scattered vascular bundles
(typical of monocots) (LM)
Epidermis
Vascular
bundles
Ground
tissue
Organization of Monocot Stems
In most monocot
stems, the vascular
tissue consists of
bundles of phloem
and xylem scattered
throughout the
ground tissue.
92. Epidermis
• outer cell layer on both sides of leaf
• secrete waxy cuticle to waterproof the leaf
Mesophyll (ground tissue of leaf)
• loosely packed photosynthetic parenchyma cells
• palisade or spongy arrangement
Vascular Bundles
• phloem & xylem
• surrounded by bundle sheath cells
Stomata (singular = “stoma”)
• openings for gas exchange, transpiration
• regulated by guard cells
93. All gymnosperms and most eudicots undergo
growth in diameter or width – 2o growth.
• most monocots undergo primary growth only
VASCULAR CAMBIUM
• a single-celled ring of meristem between primary xylem
and phloem
• produces new (secondary) xylem toward the inside and
new (secondary) phloem toward the outside
CORK CAMBIUM
• produces cork cells periderm in place of the original
epidermis to produce a protective outer layer