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PLANT STRUCTURE
AND GROWTH
1.Vascular Plant Structure
2.Vascular Plant Growth
VASCULAR PLANT
STRUCTURE
The plant body has a hierarchy of
organs, tissues, and cells
Like multicellular animals, plants have organs
composed of different tissues, which in turn are
composed of cells.
Three Basic Plant Organs (Roots, Stems and
Leaves)
ORGANS
distinct functional structure consisting of multiple
types of tissues (Dermal, Vascular and Ground
tissues)
TISSUES
collection of one or more cell types that performs
a specific function within an organ
Each of these three
categories forms a system
(Roots system and
Shoots System)
Roots rely on sugar
produced by
photosynthesis in the
shoot system.
Shoots rely on water and
minerals absorbed by the
root system.
PLANT CELL STRUCTURES
Structures found in plant cells but
not in animals cells.
LARGE CENTRAL VACUOLE –
surrounded by its own membrane
and contains water and dissolved
substances. Its primary role is to
maintain pressure against the
inside of the cell wall, giving the
cell shape and helping to support
the plant.
Cell wall – located outside
the cell membrane. It
consists mainly of cellulose
and lignin, which makes it
more rigid. The cell wall
shapes, supports, and
protects the cell. It prevents
the cell from absorbing too
much water and bursting. It
also keeps large, damaging
molecules out of the cell.
Platids are membrane-
bound organelles with
their own DNA.
Chloroplasts contain
the green pigment
chlorophyll and carry out
photosynthesis.
Chromoplasts make
and store other
pigments. They give
flower petals their bright
colors.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
Common Types of Plant Cells - are specialized of
cells in structure and function.
• Some major types of plant cells:
– Parenchyma - ground: thin flexible cell walls:
photosynthesis, storage.
– Collenchyma - ground: thicker cell walls for
flexible support.
– Sclerenchyma - ground: thick secondary cell
walls reinforced with lignin for rigid, sturdy
support.
– Xylem - vascular: water-conducting cells.
– Phloem - vascular: sugar-conducting cells.
Parenchyma cells in a privet
(Ligustrum) leaf (LM)
25 ÎĽm
Parenchyma Cells
• have thin primary (1o) cell walls without a
secondary (2o) cell wall
• the least differentiated plant cell type
• the most
metabolically active
plant cell type
• are capable of
undergoing cell
division and further
differentiation
Collenchyma Cells
• flexible support in newly formed shoot
provide structures without restraining
growth
• flexible 1o cell
walls with
irregular
2o wall
thickening
Collenchyma cells
(in Helianthus stem) (LM) 5 ÎĽm
Sclerenchyma Cells
• provide rigid support due to thick 2o cell walls
containing lignin that are dead at maturity
5 ÎĽm
Sclereid cells in pear (LM)
Cell wall
Fiber cells (cross section from ash tree) (LM)
25 ÎĽm
• 2 types of
sclerenchyma
cells:
• sclereid cells
with very thick
2o cell walls
• long and
slender fiber
cells arranged
in threads
Water-Conducting Xylem Cells
2 types of xylem
cells, both of
which are dead at
maturity:
TRACHEIDS
100 ÎĽm
Tracheids and vessels
(colorized SEM)
Vessel Tracheids
Perforation
plate
Vessel
element
Vessel elements, with
perforated end walls
Tracheids
Pits
• found in all xylem
vessels
• long, thin with
tapered ends
VESSEL ELEMENTS
• wider, less tapered
• perforated ends
Sugar-Conducting Phloem Cells
2 types of phloem cells, both of which are
alive at maturity:
SIEVE CELLS
• found in seedless
vascular plants &
gymnosperms
SIEVE-TUBE ELEMENTS
• cells that form sieve
tubes in angiosperms
• have sieve plates
between elements &
supporting
companion cells
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• Meristems are growth regions - have perpetual
embryonic tissue that allows for indeterminate
growth.
• Apical meristems are located at the tips of
roots and shoots and at the axillary buds of
shoots.
• Apical meristems elongate shoots and roots, a
process called primary growth.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• Lateral meristems add thickness to woody
plants, a process called secondary growth.
• There are two lateral meristems: the vascular
cambium and the cork cambium.
• The vascular cambium adds layers of
vascular tissue called secondary xylem = wood
and secondary phloem.
• The cork cambium replaces the epidermis
with periderm, which is thicker and tougher.
An overview of primary and secondary growth
Shoot tip (shoot
apical meristem
and young leaves)
Lateral meristems:
Axillary bud
meristem
Vascular cambium
Cork cambium
Root apical
meristems
Primary growth in stems
Epidermis
Cortex
Primary phloem
Primary xylem
Pith
Secondary growth in stems
Periderm
Cork
cambium
Cortex
Primary
phloem
Secondary
phloem
Pith
Primary
xylem
Secondary
xylem
Vascular cambium
Intercalary meristems - occur
only in monocots, at the
bases of leaf blades and at
nodes (the areas where
leaves attach to a stem). This
tissue enables the monocot
leaf blade to increase in
length from the leaf base; for
example, it allows lawn grass
leaves to elongate even after
repeated mowing.
•Meristems produce cells that quickly differentiate, or
specialize, and become permanent tissue. Such cells
take on specific roles and lose their ability to divide
further.
•They differentiate into three main types: dermal,
vascular, and ground tissue.
•Dermal tissue covers and protects the plant.
•Vascular tissue transports water, minerals, and
sugars to different parts of the plant.
•Ground tissue serves as a site for photosynthesis,
provides a supporting matrix for the vascular tissue,
and helps to store water and sugars.
•Secondary tissues are either simple (composed of
similar cell types) or complex (composed of different cell
types).
•Dermal tissue, for example, is a simple tissue that covers
covers the outer surface of the plant and controls gas
exchange. Vascular tissue is an example of a complex
tissue, and is made of two specialized conducting tissues:
xylem and phloem.
•Xylem tissue transports water and nutrients from the
roots to different parts of the plant, and includes three
different cell types: vessel elements and tracheids (both
(both of which conduct water), and xylem parenchyma.
•Phloem tissue transports organic compounds from the
site of photosynthesis to other parts of the plant,
consists of four different cell types: sieve cells (which
conduct photosynthates), companion cells, phloem
parenchyma, and phloem fibers.
•Unlike xylem conducting cells, phloem conducting cells
are alive at maturity. The xylem and phloem always lie
adjacent to each other.
•In stems, the xylem and the phloem form a structure
called a vascular bundle; in roots, this is termed the
vascular stele or vascular cylinder.
•Meristems produce cells that quickly
differentiate, or specialize, and become
permanent tissue. Such cells take on
specific roles and lose their ability to divide
further.
•They differentiate into three main types:
dermal, vascular, and ground tissue.
3 Basic Plant Tissue Types
Dermal tissue
• outer, protective covering
of the plant
Vascular tissue
• transports water,
minerals, and sugars to
different parts of the plant
Ground tissue
• serves as a site for
photosynthesis, provides a
supporting matrix for the
vascular tissue, and helps to
store water and sugars.
Dermal
tissue
Ground
tissue Vascular
tissue
each of these tissues forms
a continuous tissue system
throughout the plant
More on Dermal Tissue…
In nonwoody plants and structures (e.g., leaves) the
dermal tissue is epidermis.
• epidermis is frequently covered with a waxy cuticle to
minimize water loss
• some plants also have trichomes in epidermal tissue
which provide protection from water loss, intense light
and insects
In woody plants the epidermis develops into a
protective laver called periderm (part of the bark).
Trichomes
300
ÎĽm
ď‚· The epidermis of a leaf
also contains openings
known as stomata,
through which the
exchange of gases takes
place.
ď‚· Two cells, known as guard
cells, surround each leaf
stoma, controlling its
opening and closing and
thus regulating the uptake
of carbon dioxide and the
release of oxygen and
water vapor.
More on Vascular Tissue…
Plant vascular tissue consists of phloem & xylem.
Xylem
• transports water & minerals upward from the root
system to the organs and tissues of the shoot system
Phloem
• transports photosynthetic
products (e.g., sugars)
downward to the roots and
other parts of the plant
Phloem & xylem are organized
into vascular bundles or
cylinders called steles.
VASCULAR TISSUE
ď‚· When the stem is viewed in
cross section, the vascular
bundles of dicot stems are
arranged in a ring. In plants
with stems that live for more
than one year, the individual
bundles grow together and
produce the characteristic
growth rings.
ď‚· In monocot stems, the
vascular bundles are
randomly scattered
throughout the ground
tissue.
Water-Conducting Cells of the Xylem
ď‚· Xylem tissue has three types of cells:
xylem parenchyma, tracheids, and
vessel elements.
ď‚· Tracheids are xylem cells with thick
secondary cell walls that are lignified.
ď‚· Water moves from one tracheid to
another through regions on the side
walls known as pits, where secondary
walls are absent.
ď‚·Vessel elements are xylem cells with
thinner walls; they are shorter than
tracheids. Each vessel element is
connected to the next by means of a
perforation plate at the end walls of
the element. Water moves through
the perforation plates to travel up the
plant.
Differentiated Plant Cells in the Xylem - Dead at Maturity
Perforation
plate
Vessel
element
Vessel elements, with
perforated end walls
Tracheids
Pits
Tracheids and vessels
(colorized SEM)
Vessel Tracheids 100 µm
Sugar-Conducting Cells of the Phloem
ď‚· Phloem tissue is composed of
sieve-tube cells, companion
phloem parenchyma, and
fibers.
ď‚· Sieve-tube elements are alive at
functional maturity, though they
lack organelles.
ď‚· Sieve plates are the porous end
walls that allow fluid to flow
between cells along the sieve
tube.
ď‚· Each sieve-tube element has a
companion cell whose nucleus
and ribosomes serve both cells.
Differentiated
Plant Cells
Sieve-tube element (left)
and companion cell:
cross section (TEM)
3 µm
Sieve-tube elements:
longitudinal view (LM)
Sieve plate
Companion
cells
Sieve-tube
elements
Plasmodesma
Sieve
plate
Nucleus of
companion
cells
Sieve-tube elements:
longitudinal view Sieve plate with pores (SEM)
10 µm
30 µm
Sieve-tube elements:
longitudinal view (LM)
Sieve plate
Companion
cells
Sieve-tube
elements
30 µm
Sieve-tube
element
Plasmodesma
Sieve
plate
Nucleus of
companion
cells
Sieve-tube elements:
longitudinal view Sieve plate with pores (SEM)
10 µm
More on Ground Tissue…
Tissues that are not dermal or vascular are ground
tissue which come in 2 general types.
Pith
• ground tissue found
internal to the
vascular tissue
Cortex
• ground tissue found
between the dermal
and vascular tissue
Ground tissues include cells involved in storage,
transport, structural support and photosynthesis.
SUMMARY
PLANTS ORGANS
SHOOT
SYSTEM
ď‚·The shoot system consists of
two portions: the vegetative
(non-reproductive) parts of the
plant, such as the leaves and the
stems, and the reproductive
parts of the plant, which include
flowers and fruits.
ROOT
SYSTEM
• The root system, which
supports the plants and
absorbs water and minerals, is
usually underground.
Three Basic Plant Organs
Plant organs evolved to obtain nutrients, water
and energy on land – below & above ground
ROOTS
• absorb water, minerals and other nutrients from
the soil
• anchor & support plant in the ground
STEMS
• structural support of plant above ground
• transport of water & nutrients throughout the
plant
LEAVES
• harvesting light & CO2 for photosynthesis
Stem Structure and Function
Stem structures include:
points of leaf attachment
called nodes
internodes – the stems
between each node
apical buds at the shoot tips
where growth occurs
axillary buds which give rise to
lateral branches, thorns or flowers
The petiole is the stalk
connecting the leaf to the stem
STEM ANATOMY
The stem and other plant organs arise
from the ground tissue, and are primarily
made up of simple tissues formed from
three types of parenchyma, collenchyma,
and sclerenchyma cells
•
PARENCHYMA CELLS are
the most common plant cells.
•They are found in the stem,
the root, the inside of the leaf,
and the pulp of the fruit.
•Parenchyma cells are
responsible for metabolic
functions, such as
photosynthesis, and they help
repair and heal wounds.
•Some parenchyma cells also
store starch.
ď‚· COLLENCHYMA CELLS
are elongated cells with
unevenly thickened
walls.
ď‚· They provide structural
support, mainly to the
stem and leaves. These
cells are alive at maturity
and are usually found
below the epidermis.
 The “strings” of a celery
stalk are an example of
collenchyma cells.
ď‚· SCLERENCHYMA CELLS also
provide support to the plant, but
unlike collenchyma cells, many
of them are dead at maturity.
ď‚· There are two types of
sclerenchyma cells: fibers and
sclereids. Both types have
secondary cell walls that are
thickened with deposits of lignin,
an organic compound that is a
key component of wood.
ď‚· Fibers are long, slender cells;
sclereids are smaller-sized.
Sclereids give pears their gritty
texture.
ď‚· Humans use sclerenchyma
fibers to make linen and rope.
Evolutionary Adaptations of Stems
Stems can be modified to serve a variety of
functions:
• rhizomes which grow just beneath
the soil surface and give rise to
vertical shoots from axillary buds
• stolons that function as “runners”
along the soil surface giving rise to
new plantlets
• tubers that serve
as storage “sinks”
for carbohydrates
Root
Rhizome
Rhizomes
Stolons
Stolo
Tubers
• Some aerial modifications of stems are tendrils and
thorns.
• Tendrils seek support by climbing on other surfaces.
• Thorns are modified branches appearing as sharp
outgrowths that protect the plant
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
Leaves = the main photosynthetic organs
• Leaves are usually green due to the presence of
chlorophyll in the leaf cells.
• Each variation helps a plant species maximize its
chances of survival in a particular habitat.
• Usually, the leaves of plants growing in tropical
rainforests have larger surface areas than those of
plants growing in deserts or very cold conditions,
which are likely to have a smaller surface area to
minimize water loss.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
STRUCTURE OF A LEAF
• Each leaf typically has a leaf blade called
the lamina.
• Some leaves are attached to the plant stem
by a petiole.
• Leaves that do not have a petiole and are
directly attached to the plant stem are called
sessile leaves.
• Small green appendages usually found at
the base of the petiole are known as
stipules.
• Most leaves have a midrib, which travels
the length of the leaf and branches to each
side to produce veins of vascular tissue.
• The edge of the leaf is called the margin.
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• Within each leaf, the vascular tissue forms veins.
• The arrangement of veins in a leaf is called the
venation pattern.
• Monocots and dicots differ in their patterns of
venation.
• Monocots have parallel venation; the veins run in
straight lines
• In dicots, however, the veins of the leaf have a net-
like appearance, forming a pattern known as
reticulate venation.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• One extant plant, the Ginkgo biloba, has
dichotomous venation where the veins fork.
a) Tulip, a monocot, has leaves with parallel venation.
The netlike venation in this (b) linden leaf distinguishes it
as a dicot. The (c) Ginkgo biloba tree has dichotomous
venation.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
LEAF ARRANGEMENT
• The arrangement of leaves on a stem is known as
PHYLLOTAXY.
• Leaves are classified as either ALTERNATE, SPIRAL,
OR OPPOSITE.
• Plants that have only one leaf per
node have leaves that are said to
be either alternate—meaning
the leaves alternate on each side
of the stem in a flat plane—or
spiral, meaning the leaves are
arrayed in a spiral along the stem.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• In an opposite leaf arrangement, two leaves
arise at the same point, with the leaves connecting
opposite each other along the branch.
• If there are three or more leaves connected at a
node, the leaf arrangement is classified as
WHORLED
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
LEAF FORM
• Leaves may be simple or
compound.
• In a simple leaf, the blade is
either completely undivided—
as in the banana leaf—or it
has lobes, but the separation
does not reach the midrib, as
in the maple leaf.
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• In a compound leaf, the leaf blade is completely
divided, forming leaflets, as in the locust tree. Each
leaflet may have its own stalk, but is attached to the
rachis.
• A palmately compound leaf resembles the palm of a
hand, with leaflets radiating outwards from one point
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
ď‚· Pinnately compound leaves take their name from
their feather-like appearance; the leaflets are
arranged along the midrib.
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• The honey locust has double compound
leaves, in which leaflets branch from the
veins.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
LEAF STRUCTURE AND FUNCTIONS
• The outermost layer of the leaf is the epidermis; it is
present on both sides of the leaf and is called the upper
and lower epidermis, respectively.
• The upper side the adaxial surface (or adaxis) and the
lower side the abaxial surface (or abaxis).
• The epidermis helps in the regulation of gas exchange.
• It contains stomata: openings through which the exchange
of gases takes place.
• Two guard cells surround each stoma, regulating its
opening and closing
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
(a) The surface of this sumac leaf. At 5,000x
magnification, the guard cells of (b) a single stoma from
lyre-leaved sand cress have the appearance of lips that
surround the opening. In this (c) light micrograph cross-
section of an A. lyrata leaf, the guard cell pair is visible
along with the large, sub-stomatal air space in the leaf.
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• The epidermis is usually one cell layer thick;
however, in plants that grow in very hot or very
cold conditions, the epidermis may be several
layers thick to protect against excessive water loss
from transpiration.
• A waxy layer known as the cuticle covers the
leaves of all plant species which reduces the rate
of water loss from the leaf surface.
• Other leaves may have small hairs (trichomes) on
the leaf surface, which help to deter herbivore by
restricting insect movements, or by storing toxic or
bad-tasting compounds; they can also reduce the
rate of transpiration by blocking air flow across the
leaf surface
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• Below the epidermis of dicot leaves are layers of
cells known as the mesophyll, or “middle leaf.”
ď‚· The mesophyll of most leaves typically contains two arrangements of
parenchyma cells: the palisade parenchyma and spongy parenchyma.
ď‚· The palisade parenchyma has column-shaped, tightly packed cells, and
may be present in one, two, or three layers.
ď‚· Below the palisade parenchyma are loosely arranged cells of an irregular
shape. These are the cells of the spongy parenchyma
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
• In aquatic plants, the intercellular spaces in
the spongy parenchyma help the leaf float.
Both layers of the mesophyll contain many
chloroplasts.
• Guard cells are the only epidermal cells to
contain chloroplasts.
CO2
O2
H2O
Minerals
ROOTS
ď‚· The roots have three
major functions:
1.anchoring the plant to
the soil,
2.absorbing water and
minerals and
transporting them
upwards,
3.storing the products of
photosynthesis.
*over-watering can suffocate a plant!
ď‚· Some roots are modified to absorb
moisture and exchange gases.
ď‚· Most roots are underground.
ď‚· Some plants, however, also have
adventitious roots, which emerge above
the ground from the shoot.
TYPES OF ROOTS SYSTEM
DICOTS
A tap root system has a main
root that grows down vertically,
and from which many smaller
lateral roots arise.
A tap root system penetrates
deep into the soil.
MONOCOTS
A fibrous root system is
located closer to the soil
surface, and forms a dense
network of roots that also
helps prevent soil erosion.
ď‚· Some plants have a combination of tap
roots and fibrous roots.
ď‚· Plants that grow in dry areas often have
deep root systems, whereas plants
growing in areas with abundant water
are likely to have shallower root
systems.
ROOT GROWTH AND ANATOMY
ď‚· Root growth begins with seed
germination.
ď‚· When the plant embryo emerges
from the seed, the radicle of the
embryo forms the root system.
ď‚· The tip of the root is protected by the
root cap, a structure exclusive to
roots and unlike any other plant
structure.
ď‚· The root cap is continuously replaced
because it gets damaged easily as the
root pushes through soil.
ď‚· The root tip can be divided into three
zones: a zone of cell division, a zone
of elongation, and a zone of
maturation and differentiation.
ď‚· The zone of cell division is closest to
the root tip; it is made up of the
actively dividing cells of the root
meristem.
ď‚· The zone of elongation is where the
newly formed cells increase in length,
thereby lengthening the root.
ď‚· Beginning at the first root hair is the
zone of cell maturation where the
cells begin to differentiate into special
cell types.
ď‚· Inside the root, the ground tissue forms two regions:
the cortex and the pith.
ď‚· Both regions include cells that store photosynthetic
products.
ď‚· The cortex is between the epidermis and the vascular
tissue, whereas the pith lies between the vascular
tissue and the center of the root.
ď‚· The vascular tissue in the root is arranged in the inner
portion of the root, which is called the stele.
ď‚· A layer of cells known as the endodermis separates the
stele from the ground tissue in the outer portion of
the root.
ď‚· The endodermis is exclusive to roots, and serves as a
checkpoint for materials entering the root’s vascular
system.
ď‚· A waxy substance called suberin is present on the
walls of the endodermal cells.
ď‚· This waxy region, known as the Casparian strip,
forces water and solutes to cross the plasma
membranes of endodermal cells instead of slipping
between the cells.
ď‚· This ensures that only materials required by the root
pass through the endodermis, while toxic substances
and pathogens are generally excluded.
 The outermost cell layer of the root’s vascular tissue is
the pericycle, an area that can give rise to lateral
roots.
ď‚· In dicot roots, the xylem and phloem of the stele are
arranged alternately in an X shape, whereas in
monocot roots, the vascular tissue is arranged in a
ring around the pith.
ROOT MODIFICATIONS
ď‚· Root structures may be modified for specific
purposes. For example, some roots are bulbous and
store starch.
ď‚· Aerial roots and prop roots are two forms of
aboveground roots that provide additional support to
anchor the plant.
ď‚· Tap roots, such as carrots, turnips, and beets, are
examples of roots that are modified for food storage
Many
plants
have
modified
roots
Prop roots
“Strangling”
aerial roots
Storage roots
Buttress roots
Pneumatophores
Modified roots
Prop roots - support tall top heavy plants
Modified
Roots
Pneumatophores - “air roots” enable root systems to capture oxygen
Modified Roots
Buttress roots - support tall trunks of some tropical trees “like butresses.”
ď‚· Epiphytic roots enable a plant to
grow on another plant.
ď‚· For example, the epiphytic roots
of orchids develop a spongy
tissue to absorb moisture.
Vascular Plant
Growth
Meristem Tissue
Unlike animals, plants are capable of indeterminate
growth – growth throughout the life of the plant.
This unlimited growth potential is due to meristem
tissue – a special, undifferentiated tissue with
unlimited replicative potential.
• in contrast, animals and some plant structures (e.g.,
flowers, thorns) exhibit determinate growth in which
they stop growing when they reach a certain size
There are 2 types of meristems:
• APICAL MERISTEM • LATERAL MERISTEM
Root apical
meristems
Axillary bud
meristem
Shoot tip
(shoot apical
meristem and
young leaves)
Apical meristem is located at
the tips of roots and shoots
and is responsible for growth
in length – what is called
primary growth.
• in non-woody (herbaceous)
plants, most if not all growth is
due to apical meristem
• in woody plants (e.g., trees),
there is also growth in width,
what is referred to as
secondary growth…
Vascular
cambium
Cork cambium
Lateral
meristems
Primary
xylem
Secondary
xylem
Pith Cortex
Primary
phloem
Secondary
phloem
Vascular
cambium
Secondary growth in stems
Cork cambium
Periderm
Primary growth in stems
Epidermis
Cortex
Primary phloem
Primary xylem
Pith
Lateral Meristem
Secondary growth in width is due to
2 types of lateral meristem:
• vascular cambium which adds new
layers of phloem & xylem
• cork cambium which replaces the
epidermis with protective periderm
Primary Growth of Roots
100 ÎĽm
Mitotic
cells
Zone of cell
division
(including
apical
meristem)
Root cap
Zone of
elongation
Zone of
differentiation
Dermal
Ground
Vascular
Vascular cylinder
Cortex
Epidermis
Root hair
Root tips have a protective, non-dividing root cap.
Just underneath the
root cap is the
Zone of Cell Division
which contains the
apical meristem cells.
Beyond the Zone of
Cell Division are 2
zones in successive
developmental stages:
Zone of Elongation
• pushes root into soil
Zone of Differentiation
• cells adopt specific fates
100 ÎĽm
(a) Root with xylem and phloem in
the center (typical of eudicots)
Xylem
Phloem
Dermal
Ground
Vascular
Pericycle
Core of
parenchyma
cells
Vascular cylinder
Endodermis
Cortex
Epidermis
Endodermis
Pericycle
Xylem
Phloem
70 ÎĽm
In most eudicot
roots, there is a
central vascular
cylinder (stele)
with a “X-shaped”
arrangement of
xylem as seen in
cross section with
phloem filling in
between the
“arms” of the X.
Eudicot
Roots
Xylem
Phloem
Dermal
Ground
Vascular
Pericycle
Core of
parenchyma
cells
Vascular cylinder
Endodermis
Cortex
Epidermis
100 ÎĽm
(b) Root with parenchyma in the
center (typical of monocots)
In most monocot
roots, there is a
core of parenchyma
cells surrounded by
a ring of alternating
phloem and xylem
vessels.
Monocot
Roots
Lateral Root Growth
100 ÎĽm Epidermis
Lateral root
Emerging
lateral
root
Cortex
Vascular
cylinder
1
Pericycle
2 3
Lateral root growth occurs from the meristematic
pericycle, the outermost layer of cells in the
vascular cylinder just inside the endodermis, the
innermost layer of cortex.
Primary Growth of Shoots
Leaf primordia
Young leaf
Shoot apical
meristem
Developing
vascular
strand
Axillary bud
meristems
0.25 mm
Primary growth of shoot structures occurs from:
• apical meristem
which lengthens
the stem and
gives rise to leaf
primordia
• axial meristem
which gives rise
to new branches
from the main
stem
Organization of Eudicot Stems
1 mm
Dermal
Ground
Vascular
(a) Cross section of stem with
vascular bundles forming a
ring (typical of eudicots) (LM)
Cortex
Pith
Vascular
bundle
Epidermis
Xylem
Phloem
Sclerenchyma
(fiber cells) Ground tissue
connecting
pith to cortex
In most eudicot
stems, the vascular
tissue consists of
bundles of phloem
and xylem arranged
in a ring around the
central pith tissue.
• the xylem is always
located inside the
phloem adjacent to
the pith
1 mm
Dermal
Ground
Vascular
(b) Cross section of stem with
scattered vascular bundles
(typical of monocots) (LM)
Epidermis
Vascular
bundles
Ground
tissue
Organization of Monocot Stems
In most monocot
stems, the vascular
tissue consists of
bundles of phloem
and xylem scattered
throughout the
ground tissue.
Leaf Structure
50
ÎĽm
100
ÎĽm
Guard
cells
Vein
Cuticle
Dermal
Ground
Vascular
Lower
epidermis
Spongy
mesophyll
Palisade
mesophyll
Upper
epidermis
Phloem
Xylem
Bundle-
sheath
cell
(a) Cutaway drawing of leaf tissues
Sclerenchyma
fibers
Stoma
Cuticle
Epidermal
cell
Guard
cells
Stomatal
pore
(b) Surface view of a spiderwort
(Tradescantia) leaf (LM)
Vein Air spaces Guard cells
(c) Cross section of a lilac
(Syringa) leaf (LM)
Epidermis
• outer cell layer on both sides of leaf
• secrete waxy cuticle to waterproof the leaf
Mesophyll (ground tissue of leaf)
• loosely packed photosynthetic parenchyma cells
• palisade or spongy arrangement
Vascular Bundles
• phloem & xylem
• surrounded by bundle sheath cells
Stomata (singular = “stoma”)
• openings for gas exchange, transpiration
• regulated by guard cells
All gymnosperms and most eudicots undergo
growth in diameter or width – 2o growth.
• most monocots undergo primary growth only
VASCULAR CAMBIUM
• a single-celled ring of meristem between primary xylem
and phloem
• produces new (secondary) xylem toward the inside and
new (secondary) phloem toward the outside
CORK CAMBIUM
• produces cork cells periderm in place of the original
epidermis to produce a protective outer layer
Thank
You!

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  • 1. PLANT STRUCTURE AND GROWTH 1.Vascular Plant Structure 2.Vascular Plant Growth
  • 3. The plant body has a hierarchy of organs, tissues, and cells Like multicellular animals, plants have organs composed of different tissues, which in turn are composed of cells. Three Basic Plant Organs (Roots, Stems and Leaves)
  • 4. ORGANS distinct functional structure consisting of multiple types of tissues (Dermal, Vascular and Ground tissues) TISSUES collection of one or more cell types that performs a specific function within an organ
  • 5. Each of these three categories forms a system (Roots system and Shoots System) Roots rely on sugar produced by photosynthesis in the shoot system. Shoots rely on water and minerals absorbed by the root system.
  • 7. Structures found in plant cells but not in animals cells. LARGE CENTRAL VACUOLE – surrounded by its own membrane and contains water and dissolved substances. Its primary role is to maintain pressure against the inside of the cell wall, giving the cell shape and helping to support the plant.
  • 8. Cell wall – located outside the cell membrane. It consists mainly of cellulose and lignin, which makes it more rigid. The cell wall shapes, supports, and protects the cell. It prevents the cell from absorbing too much water and bursting. It also keeps large, damaging molecules out of the cell.
  • 9. Platids are membrane- bound organelles with their own DNA. Chloroplasts contain the green pigment chlorophyll and carry out photosynthesis. Chromoplasts make and store other pigments. They give flower petals their bright colors.
  • 10. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Common Types of Plant Cells - are specialized of cells in structure and function. • Some major types of plant cells: – Parenchyma - ground: thin flexible cell walls: photosynthesis, storage. – Collenchyma - ground: thicker cell walls for flexible support. – Sclerenchyma - ground: thick secondary cell walls reinforced with lignin for rigid, sturdy support. – Xylem - vascular: water-conducting cells. – Phloem - vascular: sugar-conducting cells.
  • 11. Parenchyma cells in a privet (Ligustrum) leaf (LM) 25 ÎĽm Parenchyma Cells • have thin primary (1o) cell walls without a secondary (2o) cell wall • the least differentiated plant cell type • the most metabolically active plant cell type • are capable of undergoing cell division and further differentiation
  • 12. Collenchyma Cells • flexible support in newly formed shoot provide structures without restraining growth • flexible 1o cell walls with irregular 2o wall thickening Collenchyma cells (in Helianthus stem) (LM) 5 ÎĽm
  • 13. Sclerenchyma Cells • provide rigid support due to thick 2o cell walls containing lignin that are dead at maturity 5 ÎĽm Sclereid cells in pear (LM) Cell wall Fiber cells (cross section from ash tree) (LM) 25 ÎĽm • 2 types of sclerenchyma cells: • sclereid cells with very thick 2o cell walls • long and slender fiber cells arranged in threads
  • 14. Water-Conducting Xylem Cells 2 types of xylem cells, both of which are dead at maturity: TRACHEIDS 100 ÎĽm Tracheids and vessels (colorized SEM) Vessel Tracheids Perforation plate Vessel element Vessel elements, with perforated end walls Tracheids Pits • found in all xylem vessels • long, thin with tapered ends VESSEL ELEMENTS • wider, less tapered • perforated ends
  • 15. Sugar-Conducting Phloem Cells 2 types of phloem cells, both of which are alive at maturity: SIEVE CELLS • found in seedless vascular plants & gymnosperms SIEVE-TUBE ELEMENTS • cells that form sieve tubes in angiosperms • have sieve plates between elements & supporting companion cells
  • 16.
  • 17. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • Meristems are growth regions - have perpetual embryonic tissue that allows for indeterminate growth. • Apical meristems are located at the tips of roots and shoots and at the axillary buds of shoots. • Apical meristems elongate shoots and roots, a process called primary growth.
  • 18. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • Lateral meristems add thickness to woody plants, a process called secondary growth. • There are two lateral meristems: the vascular cambium and the cork cambium. • The vascular cambium adds layers of vascular tissue called secondary xylem = wood and secondary phloem. • The cork cambium replaces the epidermis with periderm, which is thicker and tougher.
  • 19. An overview of primary and secondary growth Shoot tip (shoot apical meristem and young leaves) Lateral meristems: Axillary bud meristem Vascular cambium Cork cambium Root apical meristems Primary growth in stems Epidermis Cortex Primary phloem Primary xylem Pith Secondary growth in stems Periderm Cork cambium Cortex Primary phloem Secondary phloem Pith Primary xylem Secondary xylem Vascular cambium
  • 20. Intercalary meristems - occur only in monocots, at the bases of leaf blades and at nodes (the areas where leaves attach to a stem). This tissue enables the monocot leaf blade to increase in length from the leaf base; for example, it allows lawn grass leaves to elongate even after repeated mowing.
  • 21. •Meristems produce cells that quickly differentiate, or specialize, and become permanent tissue. Such cells take on specific roles and lose their ability to divide further. •They differentiate into three main types: dermal, vascular, and ground tissue. •Dermal tissue covers and protects the plant. •Vascular tissue transports water, minerals, and sugars to different parts of the plant. •Ground tissue serves as a site for photosynthesis, provides a supporting matrix for the vascular tissue, and helps to store water and sugars.
  • 22.
  • 23. •Secondary tissues are either simple (composed of similar cell types) or complex (composed of different cell types). •Dermal tissue, for example, is a simple tissue that covers covers the outer surface of the plant and controls gas exchange. Vascular tissue is an example of a complex tissue, and is made of two specialized conducting tissues: xylem and phloem. •Xylem tissue transports water and nutrients from the roots to different parts of the plant, and includes three different cell types: vessel elements and tracheids (both (both of which conduct water), and xylem parenchyma. •Phloem tissue transports organic compounds from the site of photosynthesis to other parts of the plant, consists of four different cell types: sieve cells (which conduct photosynthates), companion cells, phloem parenchyma, and phloem fibers. •Unlike xylem conducting cells, phloem conducting cells are alive at maturity. The xylem and phloem always lie adjacent to each other. •In stems, the xylem and the phloem form a structure called a vascular bundle; in roots, this is termed the vascular stele or vascular cylinder.
  • 24. •Meristems produce cells that quickly differentiate, or specialize, and become permanent tissue. Such cells take on specific roles and lose their ability to divide further. •They differentiate into three main types: dermal, vascular, and ground tissue.
  • 25. 3 Basic Plant Tissue Types Dermal tissue • outer, protective covering of the plant Vascular tissue • transports water, minerals, and sugars to different parts of the plant Ground tissue • serves as a site for photosynthesis, provides a supporting matrix for the vascular tissue, and helps to store water and sugars. Dermal tissue Ground tissue Vascular tissue each of these tissues forms a continuous tissue system throughout the plant
  • 26. More on Dermal Tissue… In nonwoody plants and structures (e.g., leaves) the dermal tissue is epidermis. • epidermis is frequently covered with a waxy cuticle to minimize water loss • some plants also have trichomes in epidermal tissue which provide protection from water loss, intense light and insects In woody plants the epidermis develops into a protective laver called periderm (part of the bark). Trichomes 300 ÎĽm
  • 27. ď‚· The epidermis of a leaf also contains openings known as stomata, through which the exchange of gases takes place. ď‚· Two cells, known as guard cells, surround each leaf stoma, controlling its opening and closing and thus regulating the uptake of carbon dioxide and the release of oxygen and water vapor.
  • 28. More on Vascular Tissue… Plant vascular tissue consists of phloem & xylem. Xylem • transports water & minerals upward from the root system to the organs and tissues of the shoot system Phloem • transports photosynthetic products (e.g., sugars) downward to the roots and other parts of the plant Phloem & xylem are organized into vascular bundles or cylinders called steles.
  • 29. VASCULAR TISSUE ď‚· When the stem is viewed in cross section, the vascular bundles of dicot stems are arranged in a ring. In plants with stems that live for more than one year, the individual bundles grow together and produce the characteristic growth rings. ď‚· In monocot stems, the vascular bundles are randomly scattered throughout the ground tissue.
  • 30. Water-Conducting Cells of the Xylem ď‚· Xylem tissue has three types of cells: xylem parenchyma, tracheids, and vessel elements. ď‚· Tracheids are xylem cells with thick secondary cell walls that are lignified. ď‚· Water moves from one tracheid to another through regions on the side walls known as pits, where secondary walls are absent. ď‚·Vessel elements are xylem cells with thinner walls; they are shorter than tracheids. Each vessel element is connected to the next by means of a perforation plate at the end walls of the element. Water moves through the perforation plates to travel up the plant.
  • 31. Differentiated Plant Cells in the Xylem - Dead at Maturity Perforation plate Vessel element Vessel elements, with perforated end walls Tracheids Pits Tracheids and vessels (colorized SEM) Vessel Tracheids 100 µm
  • 32. Sugar-Conducting Cells of the Phloem ď‚· Phloem tissue is composed of sieve-tube cells, companion phloem parenchyma, and fibers. ď‚· Sieve-tube elements are alive at functional maturity, though they lack organelles. ď‚· Sieve plates are the porous end walls that allow fluid to flow between cells along the sieve tube. ď‚· Each sieve-tube element has a companion cell whose nucleus and ribosomes serve both cells.
  • 33. Differentiated Plant Cells Sieve-tube element (left) and companion cell: cross section (TEM) 3 µm Sieve-tube elements: longitudinal view (LM) Sieve plate Companion cells Sieve-tube elements Plasmodesma Sieve plate Nucleus of companion cells Sieve-tube elements: longitudinal view Sieve plate with pores (SEM) 10 µm 30 µm
  • 34. Sieve-tube elements: longitudinal view (LM) Sieve plate Companion cells Sieve-tube elements 30 µm
  • 36. More on Ground Tissue… Tissues that are not dermal or vascular are ground tissue which come in 2 general types. Pith • ground tissue found internal to the vascular tissue Cortex • ground tissue found between the dermal and vascular tissue Ground tissues include cells involved in storage, transport, structural support and photosynthesis.
  • 38.
  • 39. PLANTS ORGANS SHOOT SYSTEM ď‚·The shoot system consists of two portions: the vegetative (non-reproductive) parts of the plant, such as the leaves and the stems, and the reproductive parts of the plant, which include flowers and fruits. ROOT SYSTEM • The root system, which supports the plants and absorbs water and minerals, is usually underground.
  • 40. Three Basic Plant Organs Plant organs evolved to obtain nutrients, water and energy on land – below & above ground ROOTS • absorb water, minerals and other nutrients from the soil • anchor & support plant in the ground STEMS • structural support of plant above ground • transport of water & nutrients throughout the plant LEAVES • harvesting light & CO2 for photosynthesis
  • 41. Stem Structure and Function Stem structures include: points of leaf attachment called nodes internodes – the stems between each node apical buds at the shoot tips where growth occurs axillary buds which give rise to lateral branches, thorns or flowers The petiole is the stalk connecting the leaf to the stem
  • 42. STEM ANATOMY The stem and other plant organs arise from the ground tissue, and are primarily made up of simple tissues formed from three types of parenchyma, collenchyma, and sclerenchyma cells
  • 43. • PARENCHYMA CELLS are the most common plant cells. •They are found in the stem, the root, the inside of the leaf, and the pulp of the fruit. •Parenchyma cells are responsible for metabolic functions, such as photosynthesis, and they help repair and heal wounds. •Some parenchyma cells also store starch.
  • 44. ď‚· COLLENCHYMA CELLS are elongated cells with unevenly thickened walls. ď‚· They provide structural support, mainly to the stem and leaves. These cells are alive at maturity and are usually found below the epidermis. ď‚· The “strings” of a celery stalk are an example of collenchyma cells.
  • 45. ď‚· SCLERENCHYMA CELLS also provide support to the plant, but unlike collenchyma cells, many of them are dead at maturity. ď‚· There are two types of sclerenchyma cells: fibers and sclereids. Both types have secondary cell walls that are thickened with deposits of lignin, an organic compound that is a key component of wood. ď‚· Fibers are long, slender cells; sclereids are smaller-sized. Sclereids give pears their gritty texture. ď‚· Humans use sclerenchyma fibers to make linen and rope.
  • 46. Evolutionary Adaptations of Stems Stems can be modified to serve a variety of functions: • rhizomes which grow just beneath the soil surface and give rise to vertical shoots from axillary buds • stolons that function as “runners” along the soil surface giving rise to new plantlets • tubers that serve as storage “sinks” for carbohydrates Root Rhizome Rhizomes Stolons Stolo Tubers
  • 47.
  • 48. • Some aerial modifications of stems are tendrils and thorns. • Tendrils seek support by climbing on other surfaces. • Thorns are modified branches appearing as sharp outgrowths that protect the plant
  • 49. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Leaves = the main photosynthetic organs • Leaves are usually green due to the presence of chlorophyll in the leaf cells. • Each variation helps a plant species maximize its chances of survival in a particular habitat. • Usually, the leaves of plants growing in tropical rainforests have larger surface areas than those of plants growing in deserts or very cold conditions, which are likely to have a smaller surface area to minimize water loss.
  • 50. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings STRUCTURE OF A LEAF • Each leaf typically has a leaf blade called the lamina. • Some leaves are attached to the plant stem by a petiole. • Leaves that do not have a petiole and are directly attached to the plant stem are called sessile leaves. • Small green appendages usually found at the base of the petiole are known as stipules. • Most leaves have a midrib, which travels the length of the leaf and branches to each side to produce veins of vascular tissue. • The edge of the leaf is called the margin.
  • 51. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • Within each leaf, the vascular tissue forms veins. • The arrangement of veins in a leaf is called the venation pattern. • Monocots and dicots differ in their patterns of venation. • Monocots have parallel venation; the veins run in straight lines • In dicots, however, the veins of the leaf have a net- like appearance, forming a pattern known as reticulate venation.
  • 52. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • One extant plant, the Ginkgo biloba, has dichotomous venation where the veins fork. a) Tulip, a monocot, has leaves with parallel venation. The netlike venation in this (b) linden leaf distinguishes it as a dicot. The (c) Ginkgo biloba tree has dichotomous venation.
  • 53. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings LEAF ARRANGEMENT • The arrangement of leaves on a stem is known as PHYLLOTAXY. • Leaves are classified as either ALTERNATE, SPIRAL, OR OPPOSITE. • Plants that have only one leaf per node have leaves that are said to be either alternate—meaning the leaves alternate on each side of the stem in a flat plane—or spiral, meaning the leaves are arrayed in a spiral along the stem.
  • 54. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • In an opposite leaf arrangement, two leaves arise at the same point, with the leaves connecting opposite each other along the branch. • If there are three or more leaves connected at a node, the leaf arrangement is classified as WHORLED
  • 55. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings LEAF FORM • Leaves may be simple or compound. • In a simple leaf, the blade is either completely undivided— as in the banana leaf—or it has lobes, but the separation does not reach the midrib, as in the maple leaf.
  • 56. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • In a compound leaf, the leaf blade is completely divided, forming leaflets, as in the locust tree. Each leaflet may have its own stalk, but is attached to the rachis. • A palmately compound leaf resembles the palm of a hand, with leaflets radiating outwards from one point
  • 57. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings ď‚· Pinnately compound leaves take their name from their feather-like appearance; the leaflets are arranged along the midrib.
  • 58. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • The honey locust has double compound leaves, in which leaflets branch from the veins.
  • 59. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings LEAF STRUCTURE AND FUNCTIONS • The outermost layer of the leaf is the epidermis; it is present on both sides of the leaf and is called the upper and lower epidermis, respectively. • The upper side the adaxial surface (or adaxis) and the lower side the abaxial surface (or abaxis). • The epidermis helps in the regulation of gas exchange. • It contains stomata: openings through which the exchange of gases takes place. • Two guard cells surround each stoma, regulating its opening and closing
  • 60. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings (a) The surface of this sumac leaf. At 5,000x magnification, the guard cells of (b) a single stoma from lyre-leaved sand cress have the appearance of lips that surround the opening. In this (c) light micrograph cross- section of an A. lyrata leaf, the guard cell pair is visible along with the large, sub-stomatal air space in the leaf.
  • 61. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • The epidermis is usually one cell layer thick; however, in plants that grow in very hot or very cold conditions, the epidermis may be several layers thick to protect against excessive water loss from transpiration. • A waxy layer known as the cuticle covers the leaves of all plant species which reduces the rate of water loss from the leaf surface. • Other leaves may have small hairs (trichomes) on the leaf surface, which help to deter herbivore by restricting insect movements, or by storing toxic or bad-tasting compounds; they can also reduce the rate of transpiration by blocking air flow across the leaf surface
  • 62. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • Below the epidermis of dicot leaves are layers of cells known as the mesophyll, or “middle leaf.” ď‚· The mesophyll of most leaves typically contains two arrangements of parenchyma cells: the palisade parenchyma and spongy parenchyma. ď‚· The palisade parenchyma has column-shaped, tightly packed cells, and may be present in one, two, or three layers. ď‚· Below the palisade parenchyma are loosely arranged cells of an irregular shape. These are the cells of the spongy parenchyma
  • 63. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • In aquatic plants, the intercellular spaces in the spongy parenchyma help the leaf float. Both layers of the mesophyll contain many chloroplasts. • Guard cells are the only epidermal cells to contain chloroplasts.
  • 64. CO2 O2 H2O Minerals ROOTS ď‚· The roots have three major functions: 1.anchoring the plant to the soil, 2.absorbing water and minerals and transporting them upwards, 3.storing the products of photosynthesis. *over-watering can suffocate a plant!
  • 65. ď‚· Some roots are modified to absorb moisture and exchange gases. ď‚· Most roots are underground. ď‚· Some plants, however, also have adventitious roots, which emerge above the ground from the shoot.
  • 66. TYPES OF ROOTS SYSTEM DICOTS A tap root system has a main root that grows down vertically, and from which many smaller lateral roots arise. A tap root system penetrates deep into the soil. MONOCOTS A fibrous root system is located closer to the soil surface, and forms a dense network of roots that also helps prevent soil erosion.
  • 67. ď‚· Some plants have a combination of tap roots and fibrous roots. ď‚· Plants that grow in dry areas often have deep root systems, whereas plants growing in areas with abundant water are likely to have shallower root systems.
  • 68. ROOT GROWTH AND ANATOMY ď‚· Root growth begins with seed germination. ď‚· When the plant embryo emerges from the seed, the radicle of the embryo forms the root system. ď‚· The tip of the root is protected by the root cap, a structure exclusive to roots and unlike any other plant structure. ď‚· The root cap is continuously replaced because it gets damaged easily as the root pushes through soil.
  • 69. ď‚· The root tip can be divided into three zones: a zone of cell division, a zone of elongation, and a zone of maturation and differentiation. ď‚· The zone of cell division is closest to the root tip; it is made up of the actively dividing cells of the root meristem. ď‚· The zone of elongation is where the newly formed cells increase in length, thereby lengthening the root. ď‚· Beginning at the first root hair is the zone of cell maturation where the cells begin to differentiate into special cell types.
  • 70. ď‚· Inside the root, the ground tissue forms two regions: the cortex and the pith. ď‚· Both regions include cells that store photosynthetic products. ď‚· The cortex is between the epidermis and the vascular tissue, whereas the pith lies between the vascular tissue and the center of the root.
  • 71. ď‚· The vascular tissue in the root is arranged in the inner portion of the root, which is called the stele. ď‚· A layer of cells known as the endodermis separates the stele from the ground tissue in the outer portion of the root. ď‚· The endodermis is exclusive to roots, and serves as a checkpoint for materials entering the root’s vascular system.
  • 72. ď‚· A waxy substance called suberin is present on the walls of the endodermal cells. ď‚· This waxy region, known as the Casparian strip, forces water and solutes to cross the plasma membranes of endodermal cells instead of slipping between the cells. ď‚· This ensures that only materials required by the root pass through the endodermis, while toxic substances and pathogens are generally excluded.
  • 73. ď‚· The outermost cell layer of the root’s vascular tissue is the pericycle, an area that can give rise to lateral roots. ď‚· In dicot roots, the xylem and phloem of the stele are arranged alternately in an X shape, whereas in monocot roots, the vascular tissue is arranged in a ring around the pith.
  • 74. ROOT MODIFICATIONS ď‚· Root structures may be modified for specific purposes. For example, some roots are bulbous and store starch. ď‚· Aerial roots and prop roots are two forms of aboveground roots that provide additional support to anchor the plant. ď‚· Tap roots, such as carrots, turnips, and beets, are examples of roots that are modified for food storage
  • 76. Modified roots Prop roots - support tall top heavy plants
  • 77. Modified Roots Pneumatophores - “air roots” enable root systems to capture oxygen
  • 78. Modified Roots Buttress roots - support tall trunks of some tropical trees “like butresses.”
  • 79. ď‚· Epiphytic roots enable a plant to grow on another plant. ď‚· For example, the epiphytic roots of orchids develop a spongy tissue to absorb moisture.
  • 81. Meristem Tissue Unlike animals, plants are capable of indeterminate growth – growth throughout the life of the plant. This unlimited growth potential is due to meristem tissue – a special, undifferentiated tissue with unlimited replicative potential. • in contrast, animals and some plant structures (e.g., flowers, thorns) exhibit determinate growth in which they stop growing when they reach a certain size There are 2 types of meristems: • APICAL MERISTEM • LATERAL MERISTEM
  • 82. Root apical meristems Axillary bud meristem Shoot tip (shoot apical meristem and young leaves) Apical meristem is located at the tips of roots and shoots and is responsible for growth in length – what is called primary growth. • in non-woody (herbaceous) plants, most if not all growth is due to apical meristem • in woody plants (e.g., trees), there is also growth in width, what is referred to as secondary growth…
  • 83. Vascular cambium Cork cambium Lateral meristems Primary xylem Secondary xylem Pith Cortex Primary phloem Secondary phloem Vascular cambium Secondary growth in stems Cork cambium Periderm Primary growth in stems Epidermis Cortex Primary phloem Primary xylem Pith Lateral Meristem Secondary growth in width is due to 2 types of lateral meristem: • vascular cambium which adds new layers of phloem & xylem • cork cambium which replaces the epidermis with protective periderm
  • 84. Primary Growth of Roots 100 ÎĽm Mitotic cells Zone of cell division (including apical meristem) Root cap Zone of elongation Zone of differentiation Dermal Ground Vascular Vascular cylinder Cortex Epidermis Root hair Root tips have a protective, non-dividing root cap. Just underneath the root cap is the Zone of Cell Division which contains the apical meristem cells. Beyond the Zone of Cell Division are 2 zones in successive developmental stages: Zone of Elongation • pushes root into soil Zone of Differentiation • cells adopt specific fates
  • 85. 100 ÎĽm (a) Root with xylem and phloem in the center (typical of eudicots) Xylem Phloem Dermal Ground Vascular Pericycle Core of parenchyma cells Vascular cylinder Endodermis Cortex Epidermis Endodermis Pericycle Xylem Phloem 70 ÎĽm In most eudicot roots, there is a central vascular cylinder (stele) with a “X-shaped” arrangement of xylem as seen in cross section with phloem filling in between the “arms” of the X. Eudicot Roots
  • 86. Xylem Phloem Dermal Ground Vascular Pericycle Core of parenchyma cells Vascular cylinder Endodermis Cortex Epidermis 100 ÎĽm (b) Root with parenchyma in the center (typical of monocots) In most monocot roots, there is a core of parenchyma cells surrounded by a ring of alternating phloem and xylem vessels. Monocot Roots
  • 87. Lateral Root Growth 100 ÎĽm Epidermis Lateral root Emerging lateral root Cortex Vascular cylinder 1 Pericycle 2 3 Lateral root growth occurs from the meristematic pericycle, the outermost layer of cells in the vascular cylinder just inside the endodermis, the innermost layer of cortex.
  • 88. Primary Growth of Shoots Leaf primordia Young leaf Shoot apical meristem Developing vascular strand Axillary bud meristems 0.25 mm Primary growth of shoot structures occurs from: • apical meristem which lengthens the stem and gives rise to leaf primordia • axial meristem which gives rise to new branches from the main stem
  • 89. Organization of Eudicot Stems 1 mm Dermal Ground Vascular (a) Cross section of stem with vascular bundles forming a ring (typical of eudicots) (LM) Cortex Pith Vascular bundle Epidermis Xylem Phloem Sclerenchyma (fiber cells) Ground tissue connecting pith to cortex In most eudicot stems, the vascular tissue consists of bundles of phloem and xylem arranged in a ring around the central pith tissue. • the xylem is always located inside the phloem adjacent to the pith
  • 90. 1 mm Dermal Ground Vascular (b) Cross section of stem with scattered vascular bundles (typical of monocots) (LM) Epidermis Vascular bundles Ground tissue Organization of Monocot Stems In most monocot stems, the vascular tissue consists of bundles of phloem and xylem scattered throughout the ground tissue.
  • 91. Leaf Structure 50 ÎĽm 100 ÎĽm Guard cells Vein Cuticle Dermal Ground Vascular Lower epidermis Spongy mesophyll Palisade mesophyll Upper epidermis Phloem Xylem Bundle- sheath cell (a) Cutaway drawing of leaf tissues Sclerenchyma fibers Stoma Cuticle Epidermal cell Guard cells Stomatal pore (b) Surface view of a spiderwort (Tradescantia) leaf (LM) Vein Air spaces Guard cells (c) Cross section of a lilac (Syringa) leaf (LM)
  • 92. Epidermis • outer cell layer on both sides of leaf • secrete waxy cuticle to waterproof the leaf Mesophyll (ground tissue of leaf) • loosely packed photosynthetic parenchyma cells • palisade or spongy arrangement Vascular Bundles • phloem & xylem • surrounded by bundle sheath cells Stomata (singular = “stoma”) • openings for gas exchange, transpiration • regulated by guard cells
  • 93. All gymnosperms and most eudicots undergo growth in diameter or width – 2o growth. • most monocots undergo primary growth only VASCULAR CAMBIUM • a single-celled ring of meristem between primary xylem and phloem • produces new (secondary) xylem toward the inside and new (secondary) phloem toward the outside CORK CAMBIUM • produces cork cells periderm in place of the original epidermis to produce a protective outer layer