The anther is the male reproductive part of a flower that produces pollen grains. It consists of microsporangia located within each bilobed sac-like structure. Anther culture is a technique where immature pollen is cultured to divide and grow into tissue, either callus or embryos. This allows the production of haploid plants by interrupting the normal pollen development pathway. Several methods were discovered for generating haploids starting in the 1920s through anther and microspore culture.