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Improving Resiliency of 
Forests in Arkansas 
Andrew Nelson, Assistant Professor 
Arkansas Forest Resources Center, UA Division of 
Agriculture 
September 24, 2014
What is Forest Resiliency? 
Capacity of a forest to respond to a 
disturbance, by resisting damage and 
recovering quickly
Is this stand 
resilient? 
Slow growth 
Tree mortality
What about 
this stand?
Which 
stand is 
more 
resilient to 
wildfire?
Resilient Vs. 
Non-Resilient 
Forests 
Rist and Moen. 2013. FEM 310: 416-427 
Resilient Forest 
Non-Resilient Forest 
Disturbance 
Disturbance 
Resilient forests often 
rebound quicker 
following disturbance 
and have greater 
overall production 
than non-resilient 
forests
Some recent events have tested 
the resiliency of Arkansas forests 
Source: Delta Farm Press Source: Arkansas Democrat-Gazette
Ozark red oak decline was another 
event that tested forest resiliency
General strategies for enhancing 
forest resilience 
1. Increase composition of species tolerant of a 
particular disturbance/stress 
2. Reduce and maintain lower tree density 
3. Increase average tree size and vigor 
4. Plant genotypes adapted to conditions 
5. Re-introduce disturbances that mimic natural 
processes
Examples of improving resiliency in 
Arkansas 
• Mitigating losses of pine growth and yield from 
drought and wildfire 
• Conversion of bottomland sites from pine to 
hardwood 
• Restoring Ozark forests with prescribed burning 
and overstory disturbance
Consequences of drought for pine 
production 
• Drought = lower growth & greater mortality 
• RESULT is lower stand yields and possibly longer 
rotations 
• Loss of money
Drought effects on pine seedlings 
Slash pine seedling mortality during drought 
Credit: David Moorhead, UGA, bugwood.org
Drought effects on mature trees 
Pine shedding foliage in 
response to drought 
Credit: Robert Anderson, USFS-FS, bugwood.org 
• Water is necessary for trees to 
maintain physiological 
functioning 
• Required for 
photosynthesis 
• Required for wood 
production (cell wall 
expansion) 
• Less foliage = less growth
Extended droughts were common in 
Arkansas throughout the 20th century 
Source: http://www.ncdc.noaa.gov/cag/time-series 
61% of the last 
119 years in SE 
Arkansas were 
in mild to 
severe 
drought 
Dust 
Bowl
Recent droughts have been severe
All trees are susceptible to drought 
• Varies by age, site conditions, density, genetics 
• Smaller trees (seedlings and saplings) most likely to 
experience mortality from lack of water 
• Larger trees are more likely to exhibit declines in 
growth
Effects of precipitation on pine 
basal area growth 
Basal area growth in response to annual precipitation 
Amateis et al. 2013. SRS-GTR-175. pp.193-196 
Reducing 
precipitation by 
half can result 
in 33% declines 
in basal area 
growth for a 
given site index 
and latitude 
18 
16 
14 
12 
10 
8 
6 
4 
2 
0 
34.5 37 39.5 42 44.5 47 49.5 52 54.5 57 59.5 62 64.5 67 69.5 72 
Basal area growth (ft2/ac/yr) 
Annual precipitation (in) 
59.2 ft2/ac 
74.2 ft2/ac 
89.2 ft2/ac
Insect attacks increase with 
drought stress 
• Bark beetles tend to 
attack stressed trees 
• Insects can then 
spread to non-stressed 
trees causing an 
outbreak 
Ips engraver beetle outbreak in a loblolly pine 
stand in Texas during a drought 
Credit: Ronald Billings, TX Forest Service, Bugwood.org
Strategies for enhancing pine 
resiliency to drought 
• Site preparation 
• Early competition control 
• Selecting appropriate planting stock 
• Density management 
• Mid-rotation hardwood management
Site preparation to enhance 
resiliency 
Fracturing soil and 
bedding result is 
more water near 
tree roots 
Poorly-drained flatwood site in FL 
Burger & Pritchett. 1988. For Sci 34: 77-87 
Compared to no 
site prep
Early competition control 
• Seedling roots are primarily near the soil surface 
• More non-pine vegetation increases competition for 
water, increasing chance of mortality 
• Removal of non-pine vegetation allows pine seedlings 
to capture more water 
Pine seedlings 
Credit: James Miller, US Forest Service, Bugwood.org
Selecting appropriate planting stock 
• Container vs. bare-root 
seedlings 
• Container seedlings 
experience less initial 
planting shock after placed 
in the ground than bare-root 
• Planting seedlings selected 
for drought tolerance 
• Plant genotypes from 
appropriate geographical 
provinces 
Genetically similar loblolly pine 
populations 
Neale & Kremer. 2011. Nature Genetics. 12:111-122
Density management 
• Lower density 
equals less 
competition for 
water 
• Options include 
planting at lower 
densities and 
thinning to reduce 
competition 
1st commercial 
thin as row thin 
2nd commercial 
thin to target 
basal area
Mid-rotation vegetation control 
• Pines are already competing with each other for 
water. When other vegetation is present, 
competition may be even greater 
• Options may include herbicide or manual control 
timed with a commercial thinning operations
Drought can also increase wildfire 
frequency and severity 
Young pine stand where almost all tree 
died from wildfire
The number of wildfires & acres 
burned are related to summer 
drought 
2005 2006 2007 2008 2009 2010 2011 2012 2013 
Year 
Source: National Interagency Fire Center 
Number of Fires 
3000 
2500 
2000 
1500 
1000 
500 
0 
Thousand Acre Burned 
80 
70 
60 
50 
40 
30 
20 
10 
0 
Number of Fires 
Acres Burned 
Source: http://www.ncdc.noaa.gov/cag/time-series
Reasons drought is related to 
wildfire risk 
• Low precipitation creates more flammable fuels on 
the forest floor: drier fuels can accelerate spread 
• Drought causes trees to have less water in the 
foliage, increasing flammability & greater 
probability of spread from crown to crown
Older stands tend to be more 
resistant to wildfire
Thinning may reduce loss from 
wildfire 
Crown bases were scorched, but there is 
enough living foliage that the trees may not 
die
Knowledge gaps for enhancing 
pine resiliency to drought 
• What are the appropriate planting densities, 
genotypes, thinning intensities, and thinning 
schedules to maintain pine resiliency during 
drought? 
• Lower density stands will be more resilient, but may also 
produce less wood throughout the rotation 
• How many years of repeated droughts (and at what 
intensity) can forests experience before appreciable 
declines in resiliency (losses in growth and yield)?
Planting pine on hardwood sites 
Loblolly pine 
Bottomland Hardwoods
Planting pine on hardwood sites 
Loblolly pine 
Mississippi River
Issues with planting pine on 
hardwood sites 
• Soil conditions are not ideal for pine 
• Not matching the species to the site results in 
lower pine growth than on pine sites (Coastal Plain) 
• Lower yields at end or the rotation and/or longer 
rotation 
• Longer time to crown closure requiring greater initial 
initial vegetation control 
• Stressed trees are more susceptible to insects and 
pathogens
Deodor weevil can kill pine trees 
on bottomland hardwood sites
Management options for pine stands 
on bottomland hardwood sites 
• Convert stands back to species better suited to the 
site 
• Maintain low densities to decrease mortality from 
deodor weevil
Restoring Ozark forests with 
burning and overstory disturbance 
• Ozark forest frequently burned until fire 
suppression programs began in the 1920s 
• Fire suppression combined with increased timber 
extraction altered species composition and forest 
age structure 
• Many forests are now overstocked and even-aged, 
with high densities of shade tolerant, fire intolerant 
species
Re-introducing fire back into Ozark 
forests 
• Oak species in these stands are often low vigor and 
susceptible to drought and insect/pathogen attack
Prescribed burning can be used to 
reduce density of fire intolerant species 
Credit: Kyle Cunningham, UAEX
Research needs for upland 
hardwood restoration 
• Long-term forest changes that have occurred in the 
absence of fire and overstory disturbance 
• Effects of repeated prescribed burning in 
unharvested stands across a range of site quality 
• Combined effects of prescribed burning frequency 
and overstory disturbance intensity on forests
Sylamore Ranger District Indiana 
bat habitat restoration project 
• 67,151 acre project on the Sylamore RD in an area with 
numerous threatened and endangered (T&E) bat species 
• Approved in 2013, treatment implementation began in 2014 
• Goal is to enhance T&E species habitat with combinations of 
prescribed burning and overstory disturbance 
• Treatments will also enhance forest health and resiliency by 
reducing stem densities and enhancing tree vigor
Preliminary project on the 
Sylamore Experimental Forest 
• Established in 1934, shortly 
after fire suppression 
• Prescribed burned since 
1977 
• No intentional overstory 
disturbance 
• Project will examine long-term 
changes in forest 
vegetation following 
reintroduction of fire
Prescribed burning before 
overstory disturbance 
Credit: Chris Evans, IL Wildlife Action Plan, Bugwood.org 
• Reducing density of fire 
intolerant stems below 
the main canopy can 
prepare stands to 
increase chance of oak-hickory- 
pine regeneration 
following overstory 
disturbance
Summary 
• Numerous examples show that forest resiliency to 
disturbances is important in Arkansas for 
maintaining and increasing forest productivity and 
health 
• Silviculture plays a major role for enhancing forest 
resiliency by modifying practices to reduce losses 
from distrubances. Mainly focusing on improving 
forest vigor.

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Andrew Nelson's Presentation on Forest Resiliency

  • 1. Improving Resiliency of Forests in Arkansas Andrew Nelson, Assistant Professor Arkansas Forest Resources Center, UA Division of Agriculture September 24, 2014
  • 2. What is Forest Resiliency? Capacity of a forest to respond to a disturbance, by resisting damage and recovering quickly
  • 3. Is this stand resilient? Slow growth Tree mortality
  • 5. Which stand is more resilient to wildfire?
  • 6. Resilient Vs. Non-Resilient Forests Rist and Moen. 2013. FEM 310: 416-427 Resilient Forest Non-Resilient Forest Disturbance Disturbance Resilient forests often rebound quicker following disturbance and have greater overall production than non-resilient forests
  • 7. Some recent events have tested the resiliency of Arkansas forests Source: Delta Farm Press Source: Arkansas Democrat-Gazette
  • 8. Ozark red oak decline was another event that tested forest resiliency
  • 9. General strategies for enhancing forest resilience 1. Increase composition of species tolerant of a particular disturbance/stress 2. Reduce and maintain lower tree density 3. Increase average tree size and vigor 4. Plant genotypes adapted to conditions 5. Re-introduce disturbances that mimic natural processes
  • 10. Examples of improving resiliency in Arkansas • Mitigating losses of pine growth and yield from drought and wildfire • Conversion of bottomland sites from pine to hardwood • Restoring Ozark forests with prescribed burning and overstory disturbance
  • 11. Consequences of drought for pine production • Drought = lower growth & greater mortality • RESULT is lower stand yields and possibly longer rotations • Loss of money
  • 12. Drought effects on pine seedlings Slash pine seedling mortality during drought Credit: David Moorhead, UGA, bugwood.org
  • 13. Drought effects on mature trees Pine shedding foliage in response to drought Credit: Robert Anderson, USFS-FS, bugwood.org • Water is necessary for trees to maintain physiological functioning • Required for photosynthesis • Required for wood production (cell wall expansion) • Less foliage = less growth
  • 14. Extended droughts were common in Arkansas throughout the 20th century Source: http://www.ncdc.noaa.gov/cag/time-series 61% of the last 119 years in SE Arkansas were in mild to severe drought Dust Bowl
  • 15. Recent droughts have been severe
  • 16. All trees are susceptible to drought • Varies by age, site conditions, density, genetics • Smaller trees (seedlings and saplings) most likely to experience mortality from lack of water • Larger trees are more likely to exhibit declines in growth
  • 17. Effects of precipitation on pine basal area growth Basal area growth in response to annual precipitation Amateis et al. 2013. SRS-GTR-175. pp.193-196 Reducing precipitation by half can result in 33% declines in basal area growth for a given site index and latitude 18 16 14 12 10 8 6 4 2 0 34.5 37 39.5 42 44.5 47 49.5 52 54.5 57 59.5 62 64.5 67 69.5 72 Basal area growth (ft2/ac/yr) Annual precipitation (in) 59.2 ft2/ac 74.2 ft2/ac 89.2 ft2/ac
  • 18. Insect attacks increase with drought stress • Bark beetles tend to attack stressed trees • Insects can then spread to non-stressed trees causing an outbreak Ips engraver beetle outbreak in a loblolly pine stand in Texas during a drought Credit: Ronald Billings, TX Forest Service, Bugwood.org
  • 19. Strategies for enhancing pine resiliency to drought • Site preparation • Early competition control • Selecting appropriate planting stock • Density management • Mid-rotation hardwood management
  • 20. Site preparation to enhance resiliency Fracturing soil and bedding result is more water near tree roots Poorly-drained flatwood site in FL Burger & Pritchett. 1988. For Sci 34: 77-87 Compared to no site prep
  • 21. Early competition control • Seedling roots are primarily near the soil surface • More non-pine vegetation increases competition for water, increasing chance of mortality • Removal of non-pine vegetation allows pine seedlings to capture more water Pine seedlings Credit: James Miller, US Forest Service, Bugwood.org
  • 22. Selecting appropriate planting stock • Container vs. bare-root seedlings • Container seedlings experience less initial planting shock after placed in the ground than bare-root • Planting seedlings selected for drought tolerance • Plant genotypes from appropriate geographical provinces Genetically similar loblolly pine populations Neale & Kremer. 2011. Nature Genetics. 12:111-122
  • 23. Density management • Lower density equals less competition for water • Options include planting at lower densities and thinning to reduce competition 1st commercial thin as row thin 2nd commercial thin to target basal area
  • 24. Mid-rotation vegetation control • Pines are already competing with each other for water. When other vegetation is present, competition may be even greater • Options may include herbicide or manual control timed with a commercial thinning operations
  • 25. Drought can also increase wildfire frequency and severity Young pine stand where almost all tree died from wildfire
  • 26. The number of wildfires & acres burned are related to summer drought 2005 2006 2007 2008 2009 2010 2011 2012 2013 Year Source: National Interagency Fire Center Number of Fires 3000 2500 2000 1500 1000 500 0 Thousand Acre Burned 80 70 60 50 40 30 20 10 0 Number of Fires Acres Burned Source: http://www.ncdc.noaa.gov/cag/time-series
  • 27. Reasons drought is related to wildfire risk • Low precipitation creates more flammable fuels on the forest floor: drier fuels can accelerate spread • Drought causes trees to have less water in the foliage, increasing flammability & greater probability of spread from crown to crown
  • 28. Older stands tend to be more resistant to wildfire
  • 29. Thinning may reduce loss from wildfire Crown bases were scorched, but there is enough living foliage that the trees may not die
  • 30. Knowledge gaps for enhancing pine resiliency to drought • What are the appropriate planting densities, genotypes, thinning intensities, and thinning schedules to maintain pine resiliency during drought? • Lower density stands will be more resilient, but may also produce less wood throughout the rotation • How many years of repeated droughts (and at what intensity) can forests experience before appreciable declines in resiliency (losses in growth and yield)?
  • 31. Planting pine on hardwood sites Loblolly pine Bottomland Hardwoods
  • 32. Planting pine on hardwood sites Loblolly pine Mississippi River
  • 33. Issues with planting pine on hardwood sites • Soil conditions are not ideal for pine • Not matching the species to the site results in lower pine growth than on pine sites (Coastal Plain) • Lower yields at end or the rotation and/or longer rotation • Longer time to crown closure requiring greater initial initial vegetation control • Stressed trees are more susceptible to insects and pathogens
  • 34. Deodor weevil can kill pine trees on bottomland hardwood sites
  • 35. Management options for pine stands on bottomland hardwood sites • Convert stands back to species better suited to the site • Maintain low densities to decrease mortality from deodor weevil
  • 36. Restoring Ozark forests with burning and overstory disturbance • Ozark forest frequently burned until fire suppression programs began in the 1920s • Fire suppression combined with increased timber extraction altered species composition and forest age structure • Many forests are now overstocked and even-aged, with high densities of shade tolerant, fire intolerant species
  • 37. Re-introducing fire back into Ozark forests • Oak species in these stands are often low vigor and susceptible to drought and insect/pathogen attack
  • 38. Prescribed burning can be used to reduce density of fire intolerant species Credit: Kyle Cunningham, UAEX
  • 39. Research needs for upland hardwood restoration • Long-term forest changes that have occurred in the absence of fire and overstory disturbance • Effects of repeated prescribed burning in unharvested stands across a range of site quality • Combined effects of prescribed burning frequency and overstory disturbance intensity on forests
  • 40. Sylamore Ranger District Indiana bat habitat restoration project • 67,151 acre project on the Sylamore RD in an area with numerous threatened and endangered (T&E) bat species • Approved in 2013, treatment implementation began in 2014 • Goal is to enhance T&E species habitat with combinations of prescribed burning and overstory disturbance • Treatments will also enhance forest health and resiliency by reducing stem densities and enhancing tree vigor
  • 41. Preliminary project on the Sylamore Experimental Forest • Established in 1934, shortly after fire suppression • Prescribed burned since 1977 • No intentional overstory disturbance • Project will examine long-term changes in forest vegetation following reintroduction of fire
  • 42. Prescribed burning before overstory disturbance Credit: Chris Evans, IL Wildlife Action Plan, Bugwood.org • Reducing density of fire intolerant stems below the main canopy can prepare stands to increase chance of oak-hickory- pine regeneration following overstory disturbance
  • 43. Summary • Numerous examples show that forest resiliency to disturbances is important in Arkansas for maintaining and increasing forest productivity and health • Silviculture plays a major role for enhancing forest resiliency by modifying practices to reduce losses from distrubances. Mainly focusing on improving forest vigor.