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Change is Constant!
Fixity of Species ??
Photo credit: Dennis Sandgathe
Extinct Species?
4
Variation within species
Incredible
Biological
Variation
John Ray
(1627–1705)
Species
concept
and Genus
Linnaeus
(1707–1778)
Taxonomy and
Binomial
Nomenclature
Kingdom
Phylum
Class
Order
Family
Genus
Species
George Buffon
(1707–1788)
“Species adapt slightly
to new conditions.”
François-Hubert Drouais
Vestigial organs and limbs
Lamarck
(1744–1829)
“Inheritance of
acquired characteristics”
Evidence
for
Evolution
Charles Darwin
(1809–1882)
Voyage of the HMS Beagle 1832–1836
15
Geographic correlation between the
fossil record and living organisms
Megatherium:
extinct species
of sloth
Three-toed sloth
of Central and
South America
16
Glyptodon:
an extinct relative
of the armadillo
Armadillo
Galápagos
Islands
Galápagos Finches
Galápagos Tortoises
2.2 Natural Selection Theory
Evolution is
not a theory
It is a
demonstrable
fact
The theory of
Photo credit: GorillazFanAdam
Natural Selection
6 Fundamental Premises of Natural Selection
1. More offspring are produced by a species than can be
supported by naturally available food sources.
Thomas Malthus
(1766–1834)
1798
An Essay on the
Principles of
Population
“Animal populations tend to
increase exponentially while
food supplies tend to
remain stable.”
2020
me
most
of you
World population
400 BC to 2050 AD
in billions
2. Within any one species there exists a significant
amount of biological/genetic variability.
3. Individuals with advantageous traits will have an edge over
other members of their species in terms of their ability to
survive longer.
4. Those individuals with traits that will increase their chance
of living longer are more likely to reach the age at which
they will begin reproducing and so are more likely to
contribute offspring to subsequent generations ...
… and therefore, those traits that gave these individuals
this advantage will be passed on to their offspring giving
them the same advantage.
5. Those traits that are advantageous under current
environmental conditions will tend to be passed on.
An individual’s reproductive success is generally dictated
by their reproductive potential or fitness: products of their
ability to adapt (or not) to specific environmental conditions.
6. Over long periods of time, accumulated changes within a
species will eventually result in new species. This is called
speciation.
Plains bison
Bison bison bison
Wood bison
Bison bison athabascae
Starting with subspecies …
32
Genus species subspecies Genus species subspecies
European bison
Bison bonasus
North American bison
Bison bison
… eventually separate species
Genus species Genus species
… eventually different genera (plural of genus)
Bison
North America
Bison bison
Aurochs
Eurasia
Bos primigenius
34
The process of speciation is often aided
by geographic isolation.
35
Example: Trinidad Guppies Upstream
habitats
Downstream habitats
Other Fundamental Components
of the Evolutionary Process
1. Of the inherent variability in
traits in a species:
36
Most represents neither potential
advantage or disadvantage to
their survival so are not subject
to Natural Selection …
…they are Neutral Traits.
… and this depends almost entirely on the environment they face.
Of the variability, a very small percentage has the potential to be
disadvantageous, and a very small percentage has the potential to
be advantageous …
2. Natural selection only acts on the variation that already exists
in a species.
37
Now that`s what
I`m talkin aboot!
Natural selection selects from what’s already there.
Natural selection does not grant organisms what they “need”.
A common misconception about evolution is that it is moving in a
specific, purposeful direction.
38
3. Natural selection can only act on genetic variability.
39
40
4. Natural selection can only play a role in the
selection of traits that affect the reproductive
success of an individual.
41
But …
What about female
menopause in humans?
42
Grandmother Hypothesis
Microevolution
Micro- vs. Macro-evolution
Chimpanzee Bonobo
Eastern Gray FoxWestern Gray Fox
Macroevolution
44
Traditional view: species spread into a variety of environments
and, either through geographic barriers or just sheer distance,
different populations fail to continue to interbreed and so very
slowly and gradually diverge into new species as they adapt to
the new conditions they face.
How Does Speciation Actually Occur?
Phyletic Gradualism
45
The Fossil Record
… an obvious lack of
intermediate forms 46
Punctuated Equilibrium
47
48
1,000,000 years ago500,000 years ago
Replacement Evolution
desertification?
parent species goes extinct
A B
vs Divergent Evolution
Also called Anagenesis and quite rare.
Occurs when all members of species
are together in a contained environment
– caves for example:
Divergent Evolution
Both species
continue to exist
A B
Also called Cladogenesis.
It results from species occurring as separate populations that
can be subject to different selective pressures and is the most
common way that evolution occurs.

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History and fundamentals of evolutionary theory

Editor's Notes

  1. 1-1: http://commons.wikimedia.org/wiki/Image:Huxley_-_Mans_Place_in_Nature.png
  2. 2-1: http://commons.wikimedia.org/wiki/Image:Jan_Snellinck_II-Paradiesische_Landschaft_und_Erbs%C3%BCnde.jpg
  3. 3-1: Sandgathe, Dennis
  4. 4-1: http://commons.wikimedia.org/wiki/Image:Charrolaise.jpg 4-2: http://commons.wikimedia.org/wiki/Image:Belmont_Red.jpg 4-3: http://commons.wikimedia.org/wiki/Image:Texas_longhorn.jpg 4-4: http://commons.wikimedia.org/wiki/Image:Hereford_bull_large.jpg
  5. 5-1: http://commons.wikimedia.org/wiki/Image:Anoa_(Bubalus_spec.)2.jpg 5-2: http://commons.wikimedia.org/wiki/Image:Wisente_Bison_bonasus-cc.jpg 5-3: http://commons.wikimedia.org/wiki/Image:NCElk-27527-2.jpg 5-4: http://commons.wikimedia.org/wiki/Image:Dortmund-Zoo-IMG_5518-a.jpg
  6. 6-1: http://commons.wikimedia.org/wiki/Image:Erebia1Seitz.jpg 6-2: http://commons.wikimedia.org/wiki/Image:CatagrammaPerisama.jpg
  7. 7-1: http://commons.wikimedia.org/wiki/Image:SS-ray.jpg
  8. 8-1: http://commons.wikimedia.org/wiki/Image:Carl_von_Linn%C3%A9.jpg (Also Unit 1.2, Slide: 14-1)
  9. 9-1: http://commons.wikimedia.org/wiki/Image:Buffon_1707-1788.jpg
  10. 10-1: http://commons.wikimedia.org/wiki/Image:Jean-Baptiste_de_Lamarck.gif (Also Unit 1.1, Slide 24-1)
  11. Reused image 13-1: http://commons.wikimedia.org/wiki/Image:Charles_Darwin_by_G._Richmond.jpg (Also Unit 1.1, Slide 27-1) 13-2: http://commons.wikimedia.org/wiki/Image:Darwin_as_monkey_on_La_Petite_Lune.jpg
  12. 14-1: http://commons.wikimedia.org/wiki/Image:Voyage_of_the_Beagle-en.svg
  13. 15-1: http://commons.wikimedia.org/wiki/Image:Galapagos_Islands_-_Overview.PNG
  14. 16-3: Sandgathe, Dennis (Bottom image is my own construction)
  15. 17-1: http://commons.wikimedia.org/wiki/Image:Aldabra_Tortoise_Image_002.jpg 17-2: Jack Haverty, 3kitty.org. WS20080126-153854-800x600 (I have direct permission from the photographer) 17-3: http://commons.wikimedia.org/wiki/Image:Lonesome_George_-Pinta_giant_tortoise_-Santa_Cruz.jpg
  16. 12-1: http://commons.wikimedia.org/wiki/File:Barylambda.jpg 12-2: http://commons.wikimedia.org/wiki/File:Euoplocephalus_BW.jpg 12-3: http://commons.wikimedia.org/wiki/File:Ragfish.png 12-4: http://commons.wikimedia.org/wiki/File:Trilobite_Mivart.png 12-5: http://commons.wikimedia.org/wiki/File:Clam_(PSF).jpg 12-6: http://commons.wikimedia.org/wiki/File:Coral_(PSF).png 12-7: http://commons.wikimedia.org/wiki/File:GrandNP4.jpg
  17. 2-1: http://commons.wikimedia.org/wiki/Image:Felis_Chaus.jpg
  18. 3-1: http://commons.wikimedia.org/wiki/Image:Mouse_litter.jpg
  19. 4-1: http://commons.wikimedia.org/wiki/Image:Thomas_Malthus.jpg
  20. 5-1: Credit: Dennis Sandgathe My own image Population in 1964: 3.3 billion
  21. 6-1: http://commons.wikimedia.org/wiki/Image:LA2-NSRW-1-0039.jpg
  22. 7-1: http://commons.wikimedia.org/wiki/Image:Connochaetes_taurinus_-Wildebeest_crossing_river_-East_Africa.jpg
  23. 8-1: http://commons.wikimedia.org/wiki/Image:Gazella_thomsoni_in_Masai_Mara.jpg 8-2: http://commons.wikimedia.org/wiki/Image:Vincent.limshowchen_-_Bee_and_flower_(by-sa).jpg 8-3: http://commons.wikimedia.org/wiki/Image:Kampfadler_01_klein.jpg 8-4: http://commons.wikimedia.org/wiki/Image:Tawny_Frogmouth_camouflage.jpg
  24. 9-1: http://commons.wikimedia.org/wiki/Image:Ocelot.jpg
  25. 10-1: http://commons.wikimedia.org/wiki/Image:TigerWoods_Prowl.jpg 10-2: http://commons.wikimedia.org/wiki/Image:Sumatran_Tiger_006.jpg 10-3: http://commons.wikimedia.org/wiki/Image:White_tigers_drinking.jpg
  26. 26-1: http://commons.wikimedia.org/wiki/Image:Sus_scrofa_LC0074.jpg
  27. 11-1: http://commons.wikimedia.org/wiki/Image:Wood-Buffalo-NP_Waldbison_2_98-07-02.jpg 11-2: http://commons.wikimedia.org/wiki/Image:Bull_Bison.jpg
  28. 11-1: http://commons.wikimedia.org/wiki/Image:Wood-Buffalo-NP_Waldbison_2_98-07-02.jpg 11-2: http://commons.wikimedia.org/wiki/Image:Bull_Bison.jpg
  29. 12-1: http://commons.wikimedia.org/wiki/Image:Anoa_(Bubalus_spec.)2.jpg 12-2: http://commons.wikimedia.org/wiki/Image:Wisente_Bison_bonasus-cc.jpg
  30. Reused image ?? 17-1: http://commons.wikimedia.org/wiki/Image:Gazella_thomsoni_in_Masai_Mara.jpg
  31. 19-1: http://evolution.berkeley.edu Credit: University of California Museum of Paleontology. Understanding Evolution. Berkeley, CA: Author. Retrieved October 30, 2009, from http://evolution.berkeley.edu
  32. 20-1: http://commons.wikimedia.org/wiki/Image:Human_evolution.svg
  33. 21-1: http://img.photobucket.com/albums/v224/fun05/fun/lion.jpg Photobucket does not claim any ownership rights in the text, files, images, photos, video, sounds, musical works, works of authorship, applications, or any other materials (collectively, "Content") that you post on or through the Photobucket Services. By displaying or publishing ("posting") any Content on or through the Photobucket Services, you hereby grant to Photobucket and other users a non-exclusive, fully paid and royalty-free, worldwide, limited license to use, modify, delete from, add to, publicly perform, publicly display, reproduce and translate such Content, including without limitation distributing part or all of the Site in any media formats through any media channels, except Content marked "private" will not be distributed outside the Photobucket Services. Photobucket and/or other Users may copy, print or display publicly available Content outside of the Photobucket Services, including without limitation, via the Site or third party websites or applications (for example, services allowing Users to order prints of Content or t-shirts and similar items containing Content).
  34. 22-1: http://commons.wikimedia.org/wiki/Image:Gazella_thomsonii_in_flight.jpg
  35. 23-1: http://www.3kitty.org/travelrama/africa%2084/3599-0045.jpg I have an email from the photographer, Jack Haverty, giving me permission to use any photos from the 3kitty.com website as long as I include the appropriate credit for him.
  36. 24-1: http://commons.wikimedia.org/wiki/Image:Edward_S._Curtis_Collection_People_008.jpg
  37. Western and Eastern Gray Fox diverged around 500 kya 27-1: http://commons.wikimedia.org/wiki/Image:Eastern_Grey_Squirrel_in_St_James%27s_Park,_London_-_Nov_2006.jpg 27-2: http://commons.wikimedia.org/wiki/Image:Red_Squirrel_-_Lazienki.JPG 27-3: http://commons.wikimedia.org/wiki/Image:Chimpanzee-Head.jpg 27-4: http://commons.wikimedia.org/wiki/Image:Bonobo.jpg
  38. 28-1: http://commons.wikimedia.org/wiki/Image:NCElk-27527-2.jpg 28-2: http://commons.wikimedia.org/wiki/Image:Gros_Morne_NP_bullmoose.jpg 28-3: http://commons.wikimedia.org/wiki/Image:Eastern_Grey_Squirrel_in_St_James%27s_Park,_London_-_Nov_2006.jpg 28-4: http://commons.wikimedia.org/wiki/Image:Beaver_pho34.jpg 28-5: http://commons.wikimedia.org/wiki/Image:Chimpanzee-Head.jpg 28-6: http://commons.wikimedia.org/wiki/Image:Male_silverback_Gorilla.JPG
  39. 29-1: Source: Dennis Sandgathe MY OWN IMAGE
  40. 30-1: http://commons.wikimedia.org/wiki/Image:Dead_horse_in_desert_close.jpg http://commons.wikimedia.org/wiki/Image:2003-08-19_Skeleton_in_Yellowstone.jpg (?) 30-2: Middle: http://commons.wikimedia.org/wiki/Category:Fossil_Chalicotherium 30-3: http://commons.wikimedia.org/wiki/Image:Moropus.jpg
  41. 31-1: Source: Dennis Sandgathe MY OWN IMAGE