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Transpiration
The Water Cycle
Definitions
The physiological process by which the
excess water is lost from the living
tissues of plants in the form of vapor is
called transpiration.
Transpiration is the process of water
movement through a plant and its
evaporation from aerial parts, such as
leaves, stems and flowers.
Transpiration is the process by which
water vapour leaves the living plants
body and enters the atmosphere.
(Michel,1978)
According to recent studies it has be revealed that about 10 % of the moisture
found in the atmosphere is released by plants through transpiration. The
remaining 90% is mainly supplied by evaporation from, ocean, seas, lakes, rivers,
streams etc.
During dry periods, transpiration increase the loss of water in the upper layer of
soil, which effect vegetation, horticultural and agronomic field crops.
Plant transpiration is pretty much an invisible process.
Transpiration is the evaporation of water from plants and trees.
Approximately 90% of all water taken up by a plant is lost to transpiration.
Characteristics
Transpiration is the final step in a continuous water pathway that starts
from the soil, into plant roots and ends as it passes into the atmosphere.
98% of a plants energy is used in the work of transpiration
Transpiration rates are highest in leaves.
Transpiration is an elegant, sustainable natural design that performs its
vital functions without electricity, without fossil fuels and without moving
parts
Many xeric plants have small leaves, silvery reflective leaves, hairy leaves
and/or produce essential oils which are all strategies to reduce
transpiration by reducing evaporation.
Characteristics
• Transpiration is a very useful procedure for plants-
oIt creates negative pressure gradient that helps draw water
and minerals up through the plant from its roots.
oHelps to keep the plant cool on hot weather- a method of
evaporating cooling.
oSupports photosynthesis and encourages the exchange of
gases, helping maintain levels of CO2 and O2 in the
atmosphere.
Importance of transpiration
• It also plays an significant part in Global Hydrological Cycle-
oReleases approx 10% of water back in to the environment.
oProduces 90-450 kg of water for each pound of solid material produced
by plants.
oCreates water vapor that forms into fog and clouds. Transpiration is also
the reason why there is higher humidity in places with lots of vegetation
cover
oTranspiration through plant stomata is the main pathway for water
entering the atmosphere over land.
oTo a water resources person, transpiration is considered a loss to the
watershed. If plants could somehow use less water, the amount saved
would remain on the ground to increase recharge
importance
• On the basis of the passages through which plants give out water
in the form of vapor transpiration is of three types:
Stomatal
transpiration
Cuticular
transpiration
Lenticular
transpiration
Types of transpiration
• Stomatal transpiration:
• Transpiration that occurs through
stomata called stomatal
transpiration. This type of
transpiration only occurs in its
presence of sunlight (in daytime).
Because stomata open in the
present of sunlight and close in
the darkness. In this method
plants give out 80-90% water in
the form of vapor.
Types of transpiration
Types of transpiration
• Cuticular transpiration:
Transpiration that occurs
through the cuticle or cracks of
thin cuticle layer of leaves and
stems is said to be cuticular
transpiration. This is a day-night
process. In this process, 5-10%
water is given out in the form of
vapor.
Types of transpiration
• Lenticular transpiration:
Sometimes transpiration occurs
through lenticels, the small
opening in the corky tissue
covering stems and twigs, and
this type of transpiration is said
to be the lenticular
transpiration. In this process,
only 0.1% water is given off of
the forms of vapor.
Types of transpiration
Aerial parts
of whole
young plant
Lenticels
(lenticular
transpiration)
0.1%
Cutin
(cuticular
transpiration)
3%~10%
Stomatum
(stomatal
transpiration)
~ 90%
Types of transpiration
Transpiration vs Evaporation
Transpiration vs Evaporation
Transpiration vs Evaporation
Water is passively
transported into the
roots and then into the
xylem.
The forces of cohesion
and adhesion cause the
water molecules to
form a column in the
xylem.
Water moves from the
xylem into the
mesophyll cells,
evaporates from their
surfaces and leaves the
plant by diffusion
through the stomata.
Mechanism of transpiration
Mechanism of transpiration
It involves three basic steps
• Absorption at the roots.
• Capillary action in the xylem vessels.
• Evaporation at the leaf.
Roots absorb water from the soil through
osmosis.
Water travels from high concentration
area to low concentration area via a
semi-permeable membrane called
osmosis.
Cell water concentration < soil water
concentration.
Absorption
why not soil molecules enter?
Active Transport Passive Transport
Requires cellular
energy.
Does not require
cellular energy.
It circulates from
lower concentrated
areas to the higher
concentrated areas
It circulates from the
higher concentrated
areas to the lower
concentrated areas
Absorption
Root hair----greater volume—greater surface area---more osmosis
Absorption
How does the water goes up against gravity?
Turgor Pressure exerted by fluid in a cell that
presses the cell membrane against the cell wall.
Turgor is what makes living plant tissue rigid.
Loss of turgor, resulting from the loss of water
from plant cells, causes flowers and leaves to wilt.
Turgor pressure
• The rate of loss of
water from the plant is
greater than the
absorption of water in
the plant
Wilting
Transpiration Pull
• Adhesion occurs when water
forms hydrogen bonds with
xylem cell walls.
• Cohesion occurs when water
molecules hydrogen bond
with each other.
• Cohesion between water
molecules creates a “water
chain” effect.
Capillary Action
Evaporation
• Molecules move from higher
concentration to lower
concentration, this is called
diffusion.
• Internal vapour>atmospheric vapour.
Evaporation
Total Process
• Water potential quantifies the tendency of free (not bound to
solutes) water to move from one area to another due to osmosis,
gravity, mechanical pressure, or matrix effects such as surface
tension.
Water potential
Distilled water
 P = 0
 S = 0
 = 0 MPa
Water potential
Water potential
Water potential
• Lack of water in soil
• Reduce the number of leaves
• Turn the leaves into spine
• Leaf---spine
stomata—less number of stomata—
reduce transpiration
Example: cactus
Adaptation & Distribution
Xerophyte plants
Mountain Plants
Atmospheric pressure decrease
Evaporation increase
Transpiration rate increase
OAK:40000 gallon/year
Adaptation & Distribution
Cold climate and Deciduous
• Shedding of leaves
Reduce transpiration
Deciduous tress shed twice a year
Adaptation & Distribution
Cold climate and Evergreen trees
• Reduce surface area of leaves
• Decrease surface area into needle like structure
• Reduce number of stomata
• Reduce transpiration
example-pine
Adaptation & Distribution
Equatorial Evergeen
• Bigger leaves,big surface,lot of stomata
• Thik waxy layer known as cuticle
• Cuticle cover the stomatal opening in epidermis
• Prevent excessive loss of water
• Help to limited amount of transpiration
• Ex-banana leaf
Adaptation & Distribution
Not having waxy layer
• Having sunken stomata
• Cover with lots of hair
• Fine fibrous structures covers the stomatal opening
• Traps the vapor that come out to stomata
• Humidity increase into it
• Reduce transpiration
• Ex: nerium oleander
Adaptation & Distribution
Stephen Hales (1972) devised three methods for measuring the rate of
transpiration:
1. By weighing potted plants
2. By potometer
3. By measuring the humidity of air
Measurement Of Transpiration
By weighing potted plants
• Record decrease
of weight over 3
hours
• Amount of water
loss from the
leaves
Measurement Of Transpiration
Potometer
Measurement Of Transpiration
Rate of absorption proportional to transpiration rate
Measurement Of Transpiration
Measurement Of Transpiration
By Measuring the Humidity of Air
This method can be used for measuring the rate respiration from single
leaf. The leaf is enclosed in plexiglass chamber and sealed. Air is passed
through the chamber at a definite flow rate. The humidity of the air
before entering the chamber and after leaving the chamber is
measure.
Amount of water transpired= Relative humidity ( after closing the
chamber) – Initial relative humidity of chamber
Measurement Of Transpiration
Thornthwaite then derived an equation to provide evaporation estimates based
on a series of observed evaporation measurement
The first part of the Thornthwaite estimation technique (Thornthwaite, 1944,
1954) derives a monthly heat index (i) for a region based on the average
temperature t (°C) for a month
Thornthwaite equation of evapo-transpiration
Thornthwaite equation of evapo-transpiration
Factors that affect
transpiration rate of
plants are smeared in
two ways-
Plants Parameters
Environmental
Conditions
Factors Affecting Transpiration
• Plants Parameters:
Stomata: When stomata are open, transpiration rates increase; when
they are closed, transpiration rates decrease.
Number of Stomata: More stomata will provide more pores for
transpiration.
Number of Leaves: More leaves mean a bigger surface area and
stomata for transpiration.
Factors Affecting Transpiration
Cuticle: The thicker the cuticle layer on a leaf surface, the slower the
transpiration rate.
Leaf Area (Transpiring Area): A plant with large leaf area will show
more transpiration than another plant with less leaf area
Root/Shoot Ratio: A low root/shoot ratio decreases the rate of
transpiration while a high ratio increases the rate of transpiration.
Factors Affecting Transpiration
Mesophyll: Compact mesophyll reduces transpiration while a loose
mesophyll increases transpiration.
Leaf Modifications: Formation of prickles, leaf spines, scaly leaves,
phyllodes, phylloclades, are all modifications found in xerophytes to
reduce transpiration.
Factors Affecting Transpiration
Environmental Conditions:
Light intensity:
• it warms the leaves up quicker.
• It also cause the stomata to open,
thus more transpiration occurs
during the day and the rate is higher
on a sunny day compared to a cold
dull day.
Factors Affecting Transpiration
Humid atmosphere
• At low humidity there is a lower
concentration of water molecules in
the air around the leaves. This
concentration gradient helps the
transport of water molecules from
the leaves by diffusion.
• High humidity means the air around
the leaves is already saturated and
has a higher concentration of water
molecules than inside the leaves.
Factors Affecting Transpiration
Temperature:
An increase in the air
temperature warms the water
inside the leaves more quickly
causing it to evaporate quicker. It
also increases the capacity of
the air to absorb more water.
ROT
Factors Affecting Transpiration
• Wind:
Transpiration relies on diffusion. Windy
conditions cause the air molecules to be
blown away from the leaves, preventing
the air around the leaves becoming
saturated with water molecules.
• Soil Water
Factors Affecting Transpiration
• Meteorology is a branch of the atmospheric sciences which includes
atmospheric chemistry and atmospheric physics, with a major focus
on weather forecasting.
Hydrometeorology is a branch of meteorology and hydrology that
studies the transfer of water and energy between the land surface and
the lower atmosphere.
Hydrometeorology is the science which deals with the movement of
water and water vapour in the atmosphere
Hydrometerology
Types
• Flood
• Tropical cyclone
• Drought
• Desertification
• Wildfire
• Hail
• Mudflow
Hydrometerology
Hydro-meteorological Instruments:
Maximum &
Minimum
Thermometer
Soil
Thermometer
Psychrometer Wind wanes
Anemometer Evaporimeters Rain Gauge
Fortin’s
Barometer
Barographs Pyranometer Radiometer
Sunshine
Recorder etc.
Hydrometerology
Soil thermometer, Psychrometer
(Top to bottom)
Thank you
Feel Free to ask any question
Plants' Water Loss Process Explained

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Plants' Water Loss Process Explained

  • 3. Definitions The physiological process by which the excess water is lost from the living tissues of plants in the form of vapor is called transpiration. Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Transpiration is the process by which water vapour leaves the living plants body and enters the atmosphere. (Michel,1978)
  • 4. According to recent studies it has be revealed that about 10 % of the moisture found in the atmosphere is released by plants through transpiration. The remaining 90% is mainly supplied by evaporation from, ocean, seas, lakes, rivers, streams etc. During dry periods, transpiration increase the loss of water in the upper layer of soil, which effect vegetation, horticultural and agronomic field crops. Plant transpiration is pretty much an invisible process. Transpiration is the evaporation of water from plants and trees. Approximately 90% of all water taken up by a plant is lost to transpiration. Characteristics
  • 5. Transpiration is the final step in a continuous water pathway that starts from the soil, into plant roots and ends as it passes into the atmosphere. 98% of a plants energy is used in the work of transpiration Transpiration rates are highest in leaves. Transpiration is an elegant, sustainable natural design that performs its vital functions without electricity, without fossil fuels and without moving parts Many xeric plants have small leaves, silvery reflective leaves, hairy leaves and/or produce essential oils which are all strategies to reduce transpiration by reducing evaporation. Characteristics
  • 6. • Transpiration is a very useful procedure for plants- oIt creates negative pressure gradient that helps draw water and minerals up through the plant from its roots. oHelps to keep the plant cool on hot weather- a method of evaporating cooling. oSupports photosynthesis and encourages the exchange of gases, helping maintain levels of CO2 and O2 in the atmosphere. Importance of transpiration
  • 7. • It also plays an significant part in Global Hydrological Cycle- oReleases approx 10% of water back in to the environment. oProduces 90-450 kg of water for each pound of solid material produced by plants. oCreates water vapor that forms into fog and clouds. Transpiration is also the reason why there is higher humidity in places with lots of vegetation cover oTranspiration through plant stomata is the main pathway for water entering the atmosphere over land. oTo a water resources person, transpiration is considered a loss to the watershed. If plants could somehow use less water, the amount saved would remain on the ground to increase recharge importance
  • 8. • On the basis of the passages through which plants give out water in the form of vapor transpiration is of three types: Stomatal transpiration Cuticular transpiration Lenticular transpiration Types of transpiration
  • 9. • Stomatal transpiration: • Transpiration that occurs through stomata called stomatal transpiration. This type of transpiration only occurs in its presence of sunlight (in daytime). Because stomata open in the present of sunlight and close in the darkness. In this method plants give out 80-90% water in the form of vapor. Types of transpiration
  • 11. • Cuticular transpiration: Transpiration that occurs through the cuticle or cracks of thin cuticle layer of leaves and stems is said to be cuticular transpiration. This is a day-night process. In this process, 5-10% water is given out in the form of vapor. Types of transpiration
  • 12. • Lenticular transpiration: Sometimes transpiration occurs through lenticels, the small opening in the corky tissue covering stems and twigs, and this type of transpiration is said to be the lenticular transpiration. In this process, only 0.1% water is given off of the forms of vapor. Types of transpiration
  • 13. Aerial parts of whole young plant Lenticels (lenticular transpiration) 0.1% Cutin (cuticular transpiration) 3%~10% Stomatum (stomatal transpiration) ~ 90% Types of transpiration
  • 17. Water is passively transported into the roots and then into the xylem. The forces of cohesion and adhesion cause the water molecules to form a column in the xylem. Water moves from the xylem into the mesophyll cells, evaporates from their surfaces and leaves the plant by diffusion through the stomata. Mechanism of transpiration
  • 18. Mechanism of transpiration It involves three basic steps • Absorption at the roots. • Capillary action in the xylem vessels. • Evaporation at the leaf.
  • 19. Roots absorb water from the soil through osmosis. Water travels from high concentration area to low concentration area via a semi-permeable membrane called osmosis. Cell water concentration < soil water concentration. Absorption
  • 20. why not soil molecules enter? Active Transport Passive Transport Requires cellular energy. Does not require cellular energy. It circulates from lower concentrated areas to the higher concentrated areas It circulates from the higher concentrated areas to the lower concentrated areas Absorption
  • 21. Root hair----greater volume—greater surface area---more osmosis Absorption
  • 22. How does the water goes up against gravity?
  • 23. Turgor Pressure exerted by fluid in a cell that presses the cell membrane against the cell wall. Turgor is what makes living plant tissue rigid. Loss of turgor, resulting from the loss of water from plant cells, causes flowers and leaves to wilt. Turgor pressure
  • 24. • The rate of loss of water from the plant is greater than the absorption of water in the plant Wilting
  • 26. • Adhesion occurs when water forms hydrogen bonds with xylem cell walls. • Cohesion occurs when water molecules hydrogen bond with each other. • Cohesion between water molecules creates a “water chain” effect. Capillary Action
  • 28. • Molecules move from higher concentration to lower concentration, this is called diffusion. • Internal vapour>atmospheric vapour. Evaporation
  • 30. • Water potential quantifies the tendency of free (not bound to solutes) water to move from one area to another due to osmosis, gravity, mechanical pressure, or matrix effects such as surface tension. Water potential
  • 31. Distilled water  P = 0  S = 0  = 0 MPa Water potential
  • 34. • Lack of water in soil • Reduce the number of leaves • Turn the leaves into spine • Leaf---spine stomata—less number of stomata— reduce transpiration Example: cactus Adaptation & Distribution Xerophyte plants
  • 35. Mountain Plants Atmospheric pressure decrease Evaporation increase Transpiration rate increase OAK:40000 gallon/year Adaptation & Distribution
  • 36. Cold climate and Deciduous • Shedding of leaves Reduce transpiration Deciduous tress shed twice a year Adaptation & Distribution
  • 37. Cold climate and Evergreen trees • Reduce surface area of leaves • Decrease surface area into needle like structure • Reduce number of stomata • Reduce transpiration example-pine Adaptation & Distribution
  • 38. Equatorial Evergeen • Bigger leaves,big surface,lot of stomata • Thik waxy layer known as cuticle • Cuticle cover the stomatal opening in epidermis • Prevent excessive loss of water • Help to limited amount of transpiration • Ex-banana leaf Adaptation & Distribution
  • 39. Not having waxy layer • Having sunken stomata • Cover with lots of hair • Fine fibrous structures covers the stomatal opening • Traps the vapor that come out to stomata • Humidity increase into it • Reduce transpiration • Ex: nerium oleander Adaptation & Distribution
  • 40. Stephen Hales (1972) devised three methods for measuring the rate of transpiration: 1. By weighing potted plants 2. By potometer 3. By measuring the humidity of air Measurement Of Transpiration
  • 41. By weighing potted plants • Record decrease of weight over 3 hours • Amount of water loss from the leaves Measurement Of Transpiration
  • 42. Potometer Measurement Of Transpiration Rate of absorption proportional to transpiration rate
  • 45. By Measuring the Humidity of Air This method can be used for measuring the rate respiration from single leaf. The leaf is enclosed in plexiglass chamber and sealed. Air is passed through the chamber at a definite flow rate. The humidity of the air before entering the chamber and after leaving the chamber is measure. Amount of water transpired= Relative humidity ( after closing the chamber) – Initial relative humidity of chamber Measurement Of Transpiration
  • 46. Thornthwaite then derived an equation to provide evaporation estimates based on a series of observed evaporation measurement The first part of the Thornthwaite estimation technique (Thornthwaite, 1944, 1954) derives a monthly heat index (i) for a region based on the average temperature t (°C) for a month Thornthwaite equation of evapo-transpiration
  • 47. Thornthwaite equation of evapo-transpiration
  • 48. Factors that affect transpiration rate of plants are smeared in two ways- Plants Parameters Environmental Conditions Factors Affecting Transpiration
  • 49. • Plants Parameters: Stomata: When stomata are open, transpiration rates increase; when they are closed, transpiration rates decrease. Number of Stomata: More stomata will provide more pores for transpiration. Number of Leaves: More leaves mean a bigger surface area and stomata for transpiration. Factors Affecting Transpiration
  • 50. Cuticle: The thicker the cuticle layer on a leaf surface, the slower the transpiration rate. Leaf Area (Transpiring Area): A plant with large leaf area will show more transpiration than another plant with less leaf area Root/Shoot Ratio: A low root/shoot ratio decreases the rate of transpiration while a high ratio increases the rate of transpiration. Factors Affecting Transpiration
  • 51. Mesophyll: Compact mesophyll reduces transpiration while a loose mesophyll increases transpiration. Leaf Modifications: Formation of prickles, leaf spines, scaly leaves, phyllodes, phylloclades, are all modifications found in xerophytes to reduce transpiration. Factors Affecting Transpiration
  • 52. Environmental Conditions: Light intensity: • it warms the leaves up quicker. • It also cause the stomata to open, thus more transpiration occurs during the day and the rate is higher on a sunny day compared to a cold dull day. Factors Affecting Transpiration
  • 53. Humid atmosphere • At low humidity there is a lower concentration of water molecules in the air around the leaves. This concentration gradient helps the transport of water molecules from the leaves by diffusion. • High humidity means the air around the leaves is already saturated and has a higher concentration of water molecules than inside the leaves. Factors Affecting Transpiration
  • 54. Temperature: An increase in the air temperature warms the water inside the leaves more quickly causing it to evaporate quicker. It also increases the capacity of the air to absorb more water. ROT Factors Affecting Transpiration
  • 55. • Wind: Transpiration relies on diffusion. Windy conditions cause the air molecules to be blown away from the leaves, preventing the air around the leaves becoming saturated with water molecules. • Soil Water Factors Affecting Transpiration
  • 56. • Meteorology is a branch of the atmospheric sciences which includes atmospheric chemistry and atmospheric physics, with a major focus on weather forecasting. Hydrometeorology is a branch of meteorology and hydrology that studies the transfer of water and energy between the land surface and the lower atmosphere. Hydrometeorology is the science which deals with the movement of water and water vapour in the atmosphere Hydrometerology
  • 57. Types • Flood • Tropical cyclone • Drought • Desertification • Wildfire • Hail • Mudflow Hydrometerology
  • 58. Hydro-meteorological Instruments: Maximum & Minimum Thermometer Soil Thermometer Psychrometer Wind wanes Anemometer Evaporimeters Rain Gauge Fortin’s Barometer Barographs Pyranometer Radiometer Sunshine Recorder etc. Hydrometerology
  • 60. Thank you Feel Free to ask any question