RESPIRATION
• Why theneed for gas exchange (respiration)?
Gas exchange involves uptake of O2 and release of CO2
• What is oxygen used for?
Aerobic respiration uses energy of electrons to produce ATP
Energy-depleted electrons (and associated proton in H)
accepted by O2 to form H2O
• Where does CO2 come from?
CO2 released when glucose broken down to release
electrons in the Kreb’s Cycle
3.
• Respiration involvesdiffusion
RESPIRATION
• Respiration requires diffusion of O2 and CO2 across cell membranes
• Diffusion of O2 and CO2 is passive
• Occurs if different concentrations of O2 or CO2
on either side of membrane
4.
Aquatic Gas Exchange
•Advantage:
• Keeping surface moist is no problem
• Disadvantage:
• O2 concentrations in water are low,
especially in warmer and saltier
environments
5.
Terrestrial Gas Exchange
•Advantages:
• O2 diffuses faster in air
• Air contains much more O2 than
water
• Disadvantage:
• Surfaces must be internal to avoid
loss of water due to evaporation
6.
Respiratory Surface
• Mustaccommodate diffusion of CO2
out, O2 in
• Typically very thin
• Always moist or wet
• If they dry out, they cannot function
7.
Protists and LessComplex Animals
• Once again, less complex organisms
do not need complex organ systems
• Gas exchange takes place due to
diffusion
Skin
• For skinbreathers, respiratory surface = body
covering
• Must live in moist or wet places
• Must maintain high surface area to volume ratio
• Keep body small
• Keep body flat
• Or both
Insects
• Have aseries of openings along their sides called
SPIRACLES
• Air ducts in insects; Tubes called TRACHEA lead from
spiracles to all of the body tissues
• Smallest branches = tracheoles have liquid inside
• Extend directly to cells, Gas transfer directly to cells!
• Open circulatory system does not transport O2 and CO2
13.
Gills
• Respiratory surface= thin filaments
• Typically found in aquatic organisms
• Fish
• Crayfish, crabs, lobsters,
• Clams
14.
Fish
• Most fishbreathe with gills – feathery filaments that
contain capillaries and a large surface area for gas
exchange
• They breathe by pumping water through the mouth, over
gill filaments and out through slits in the sides of the
pharynx
• Double Pump System: by decreasing pressure in mouth,
water is forced in; by increasing pressure in mouth, water
is forced out through the opercula
15.
Advantages of Gillsto Fish
• Because the gills are so VASCULAR and have a large
surface area, gas exchange can happen adequately
• Increase surface area to volume ratio
• Overcome shape, size limitations associated with skin
breathers
16.
Counter Current Exchangein Fish
• “Counter-current” = two fluids moving in opposite
directions:
Water one way and Blood the other way
• “Exchange” = transfer of energy or materials
17.
d flow increasesthe efficiency of
ater
n from high to low concentration
ures gradient exists continually
an theoretically approach 100%
Why Counter Current?
19.
Lungs
• Biggest animalsEVER (blue whales) have lungs!
• This tells us that for very large animals, lungs are
better than gills, tracheae, skin
20.
Bird Lungs
• Havecounter-current exchange
• Can extract oxygen from air thin enough to kill a human!
• One-way flow of air through lung
• No “dead air space” left after exhaling
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– Cutaneous (skin) respiration
• How to maximize rate of diffusion?
– thin skin
• What constraints are associated
with this system?
– Increase surface area with parapodia
– cannot get too large
– No energetically expensive activities
– Use cilia on skin to move water
• Phyla Annelida
23.
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– Direct diffusion through cells
• How to maximize rate of diffusion?
– thin cell or tissue layers
• What constraints are associated
with this system?
– circulate water across outer surface
– cannot be large in size
• Phyla Cnidaria
– Cannot be involved in energy-
consuming activities
24.
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– External Gills
• Phylum Chordata
– thin gills
• What constraints are associated with this system?
– Increase surface area with feather-like arrangement of gills and by using skin
as additional respiratory surface
– Must be in constant motion or in water current
– Tissue delicate and cannot be protected
• larval fish and amphibians (also in some adult salamanders)
• How to maximize rate of diffusion?
– Relatively small body size
25.
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– Internal (covered) Gills
• Phylum Chordata
– Buccal cavity (mouth)
• Two cavities act together to
draw water across surface of
gills
– Opercular cavity (gill
chamber below gill
cover or operculum)
• Bony fish (Class Osteichthyes)
• Internal gills enclosed within body chambers that move water over gills
26.
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– Internal (covered) Gills
– thin gills (blood vessels close to surface of gill)
– Increase surface area of gill with feather-like arrangement
– Uses counter-current system in gills to increase P
• How to maximize rate of diffusion?
– Constantly ‘pump’ water across gills with buccal and opercular cavities
27.
• Basic typesof respiratory systems in animals
Respiratory Systems
How do organisms ‘solve’ common problems?
– Structure of Internal Gills