Gregor Mendel studied inheritance through pea plant experiments in the 1800s. He found that traits are determined by discrete factors, now called genes, which are passed from parents to offspring. Traits are dominant or recessive, with dominant traits expressed when either one or two copies are present and recessive traits only expressed with two recessive copies. During meiosis, alleles segregate so offspring receive one of each from parents randomly. Punnett squares can predict genotypic and phenotypic ratios by combining possible gametes.