Charles Darwin published his theory of evolution by natural selection in 1859 in his book On the Origin of Species. Darwin observed that species vary in traits and the individuals with traits better suited to their environment tended to survive and reproduce more, passing on those favorable traits. This gradual process of natural selection leads to evolution of species over generations as they adapt to their environment. Darwin's observations of finches and tortoises on the Galapagos Islands supported this theory by showing related but distinct species adapted to each local environment. The document provides examples of natural selection, extinction, and other evolutionary patterns like adaptive radiation and convergent evolution.