All reptiles breathe using lungs, though the method of lung ventilation differs between groups. In squamates, the largest reptile group, lung ventilation is accomplished almost exclusively by the axial musculature used for locomotion, forcing most squamates to hold their breath during intense runs. Other reptiles have evolved other adaptations for breathing, such as crocodiles using a diaphragm and turtles using different muscles to breathe with their rigid shells.