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8. The segment polarity genes are unique among the other Drosophila segmentation genes for a
variety of reasons. a. Explain the embryonic patterning that these genes control, explain this in
terms of the expression pattern of these genes and the types of phenotypes they produce when
absent. b. The segment polarity genes set up two distinct gradients that control cell identity.
What are these gradients? c. The expression of engrailed in each segment marks a compartment
boundary. Explain how this boundary effects cell behavior. d. The gap and pair-rule genes
function during the syncytial blastoderm but the segment polarity genes act after cells are
formed. Why is this more relevant to understanding mechanisms of regulation of vertebrate
development?

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