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Sheep Breeding and
Reproduction
Sheep Genetics
• 27 pairs of Genes
• Two genes that form a gene pair may be the same
(Homozygous)
• If they are different = Heterozygous
• If Heterozygous, then one allele of the gene pair
may express itself over another = dominance
• This dominance can vary from complete to co-
dominance
• A hidden gene expression is called recessive
Sheep Genetics
• Some are sex linked
• Examples
• Genotype vs Phenotype
• Estimated phenotypic variance of the flock
(pg BRD 19)
• Heritability
Heritability
• Proportion of the total phenotypic variation that is
due to the variation in additive gene effects
• In other words, the proportion of differences due
to genetic effects and is important in the prediction
of response rates from selection.
• The square root of the variance is the standard
deviation, which is the ave. deviation of each
indiv. in the pop. from the pop. Average
• Heritability est. – handout BRD 33
Correlation
• Quantifies a relationship between two
variables
• Measured between zero and one
• Positive versus negative correlations
• Multiple effects (Regression models)
Animal Identification
• Accurate identification
• Written records
• Accurate measurements of economic
importance
• ??? Registered or not
• Two ID nos. , one for flock or indiv. ID and
the other for registration ID (if reg.)
Record Keeping
• Determine clear goals
• Allow for records such as pedigree, birth
weight, singles or twins, weaning weight,
etc.
• Five categories of records:
– REPRODUCTION, MATERNAL , GROWTH,
WOOL AND CARCASS
Estimated Breeding Value (EBV)
• EBV = b (P- ave.P), where b is the
heritability of a particular trait
• Example; Ram A has a grease fleece weight
of 15 # and the ave. flock grease fleece
weight is 11 #, then 15-11 = selection
differential [4]
• 4 x .4 (h) = 1.6 # of grease fleece advantage
for breeding Ram A
EPD’s (Expected Progeny
Differences)
• Simply ½ of the EBV
• The ave. EPD in a population is + or – from
the average of the population. A comparison
• Ex. If a ewe has a +.3 for no. of lambs born,
then one would expect the progeny to
produce .3 more lambs per lambing than the
progeny of average ewes.
EPD’s
• Another ex. is two rams A and B
• +.5 and +1.1 for fleece weight, respectively
• Ram B progeny would be expected to have
.6 pounds heavier fleeces than Ram A
• Example 3; Ram has +1.0 and Ewe has +.5
for weaning weight, then the progeny will
be 1.5 # more than the pop. ave. for WW
EPD’s
• Accuracy
• Gives an idea of reliability of the estimat
• Measured from 0-1
• An accuracy of .45 is not very reliable,
whereas > .9 is considered reliable
• Low accuracies are a result of limited
information known about the parents or low
numbers of progeny occur
Generation Interval
• The time lapse between birth of an animal
and the birth of its replacement – usually 3-
4 years for sheep
• A measurement of progress or rate of
improvement
• Genetic Improvement per year = heritability
x selection differential divided by the
generation interval
Methods of Selection for Single
Traits
• Individual selection-selection on their own
performance
• Family selection-selection based on
bloodlines; useful when (h) is low
• Pedigree selection-similar to family and is
dependent upon how closely related the
ancestors. Useful when considering same
sex
Methods of Selection for Single
Traits
• Progeny Test- observing the performance of
the offspring. Must be mated to several
ewes and then look at the offspring. Best
when looking at carcass traits
• Combined Selection- uses more than one of
the above mentioned methods
Methods of Selection for
Multiple Traits
• Tandem Selection- focuses on multiple
traits, yet one at a time. After the
performance of one is achieved, then move
to the next trait
• Independent Culling- Set minimum
standards for more than one trait at a time
for the indiv. Cull any that does not meet
the min. stds for any trait
Methods of Selection for
Multiple Traits
• Selection Index- rank indiv. animals for two
or more traits based on a combination effect
• One the farm testing:
• Primary focus is on the ewe flock
• Therefore, traits such as prolificacy,
weaning weight and fleece weight
National Sheep Improvement
Program
• Oversees the promotion, funding, development
and implementation of a national genetic
evaluation program for sheep
• Minimum criteria for record keeping
• Lamb records: ID #, sire and dam ID, date of
birth, sex, type of birth, & type of rearing.
• Additional records are reproductive, growth and
fleece traits
NSIP
• Focuses on:
• Number of lambs born per ewe lambing
• Body weight at 30,60,90,120 & 240 days
• Grease &/or clean fleece wt.
• Staple length
• Fleece grade
• Pounds of lamb weaned per ewe exposed per year
Adj. Factors for No. of lambs born per
lambing to a common ewe
Age of Dam Adj. Factor
1 1.45
2 1.15
3 1.05
4 1.00
5, 6, & 7 .96, .96 & .95
8 .98
9+ 1.00
Major Economically Important
Traits
• Reproductive Efficiency
• Carcass Merit
• Milk Production
• Total Ewe Production- # lamb/ewe exposed
• Mature Size
• Hardiness & Adaptability
• Wool Production
• Growth Rate
Selection for Growth
• Growth is a very important trait, esp. for
meat breeds
• Adjusted 90 day weaning weight
• If birth weight is known:
• [Actual wt. - birth wt. x adj. Days (90)/
actual age ]+ birth wt.
• If birth wt. is not known:
• Actual wt. X adj. Age in days/age
Selection for Fleece Traits
• Grease and Clean Fleece Wts
• Yield
• Staple length- at least 3”/yr.
• Fiber diameter
• Crimp
• Color
• Density
• Belly Wool-undesirable on sides
Importance of Genetic
Improvement in Seedstock
Flocks
• Most produce their own replacements
• Therefore, genetic improvement from outside is
by the Ram purchased
• Theoretically, Genetic merit increases at the same
rate as the genetic merit of the rams, yet because
of generation intervals and replacement
production usually improvement lags two
generations behind
Mating Systems
• Purebreeding- common genetic group
• Outbreeding- unrelated within a breed
• Inbreeding-closely related with one or more
common ancestors (more than 50% related)
• Linebreeding-common ancesters, but not 50%
related
• Crossbreeding- mating of different breeds
• Grading Up-enhanced concentration of
crossbreeding to rams of a single breed
Inbreeding Coefficients
• Full brother mated to sister = .25
• Sire on daughter = .25
• Half brother to half sister = .125
• Sire on Mother = .5
• Therefore; > .5 has to be compounded over
time and added generation to generation
Crossbreeding
• Heterosis or Hybrid Vigor
• Superiority of the crossbred indiv. Relative
to the average performance of the parents
• Maximized when parents have no breed
ancestry in common (3 breed terminal X)
Crossbreeding Systems
• Two Breed Terminal(50% Heterosis)
• Three Breed Terminal (100%)
• Three Breed Rotational (86 %)
• Four Breed Rotational (93 %)
• Two Breed Rotational
• Roto-Terminal- combination of rotational and
terminal systems. The poor ewes would still be
used in a terminal crossing manner
Inherited Defects
• Many defects are recdessive in nature
• Selection pressure is exercised against certain
traits, esp. lethals and fleece defects
• Dwarfism, Spider Syndrome, Jaw Defects,
Rectal Prolapse, Inverted Eyelids,
Cryptorchidism, Horns or Scurs, Face covering,
Color, Skin Folds, Silky, Britch fibers, grey color,
paralyzed limbs, earless, etc.
Sheep
Breeding and Reproduction
• Improved lamb production
• More lambs per lambing
• More frequent lambing
• Increased percent of total sheep nos.
• Reducing death losses
Sheep
Breeding and Reproduction
• breed dependent - seasonal breeding
• Puberty-weight and age (70-100 # or 5-9 months)
• Breed to lamb as 2 year olds, yet can breed as a
yearling, but with less efficiency
• Actually, greater production per lifetime from
yearlings; Ramb. is later maturing and less effic.
• gestation - 144-152 days (med wool less)
• heat periods - 20 to 42 hours (no signs)
Sheep
Breeding and Reproduction
• ovulation occurs late in heat cycle
• 14- 20 day cycle ave. 16-17 days
• Prolifacy - > 100% lamb crop
• flushing ewes works in sheep, too
• Factors affecting reprod.
– daylight (<14 hrs/da), temp. ( <74 F or > 100),
nutrition
Hormonal Control
• Progesterone (Progestins)- produced by the corpus
luteum
• Prostaglandins- induces the regression of the
corpus luteum (ineffective in normally cycling
ewes); lutalyse
• Estrogen
• Gonadotropins- GnRH, HCG, FSH & LH
• Melatonin- regulatory of the seasonal breeder;
produced by the pineal gland
• ACTH
Sheep
Breeding and Reproduction
• Lambing
– > 4 sq. ft. pen for ewes, clean & dry
– presentation of front legs
– orphan lambs is not uncommon
– 25% death loss is common
– Feed ewes small amounts of water at first with
oats, wheat bran and hay (small amounts)
Sheep
Breeding and Reproduction
• Ram management
• mating guide
– ram lamb hand (20-25) pasture (25-35)
– yearling or > hand (50-75) pasture (35-60)
– use marker rams to detect bred ewes both at
breeding and 60 days after breeding
Replacement Ewe Selection
Item 140 % Prolif 170 % Prolif
No. ewes 100 100
Ewe lambs 70 85
Twin lambs 40 70
Tw lambs >
ave. in WW
20 35
Ewe lambs
exposed
20 30
Preg. Ewes 20
Physiological changes of
reproduction in the ewes
• REPRO – 4 chart
• Ewes in the south have longer breeding
patterns
• Sheep near the equator are less likely to
have seasonal breeding
Factors affecting reproduction in
the ewe
• Heredity
• Age
• Photoperiod (seasonal)
• Temperature and humidity
• Nutrition and Exercise
• Parturition and lactation
• Disease and parasites
• Fertility of & assoc. with the ram
Pregnancy Testing
• Breeding Marks
• Ultrasonic Scanning- best detected between
70 & 120 days
• Bagging or Udder Palpation
• Blood Progesterone- at the time of the next
expected heat
• PSPB- a protein called pregnancy-specific
protein B after day 21 of breeding.
Factors affecting the reproduction
in the Ram
• Breeding soundness exam
• Palpation of the testicles, epididymis, and
penis and visual appraisal of feet, legs, eyes
and jaws.
• Semen evaluation
• Disease prevention
• Heat stress
Semen Collection
• Components- seminal fluid and sperm
• Quality of sperm – morphology and
viability (percent live)
• Methods- artificial vagina and
electroejaculation
• Semen handling-dilution characteristics:
glucose or fructose, egg yolk, citrate or
phosphate, antibitotics, glycerol
Insemination
• Natural- 3-5 billion sperm inseminated
• AI – vaginal approach – 200 million “
• AI – cervical approach- 100 million “
• AI - Intruterine insemination via lapraroscopic
surgery- 20 million
• Time of insemination – vaginal or cervical = 12 to
18 hrs after onset of estrus
• Synchronizing with progesterone sponge should
be 48-58 hrs after removal
Accelerated and Out-of-Season
Lambing
• Day length control (natural or artificial)
• Considerations
• Produce lambs when feeding conditions are
favorable (growth rate and cost of feed)
• Market lambs when lamb supply is low
• Fertility and prolificacy
Desirable traits for accelerated
lambing
• Ewes can breed year round
• Ewes that can mate while lactacting
• Ewes that have a good lambing rate (ie
twinning)
• Sires that produce a desirable market lamb
and have the libido and fertility for
conception year round

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5. Sheep Breeding and Reproduction.ppt

  • 2. Sheep Genetics • 27 pairs of Genes • Two genes that form a gene pair may be the same (Homozygous) • If they are different = Heterozygous • If Heterozygous, then one allele of the gene pair may express itself over another = dominance • This dominance can vary from complete to co- dominance • A hidden gene expression is called recessive
  • 3. Sheep Genetics • Some are sex linked • Examples • Genotype vs Phenotype • Estimated phenotypic variance of the flock (pg BRD 19) • Heritability
  • 4. Heritability • Proportion of the total phenotypic variation that is due to the variation in additive gene effects • In other words, the proportion of differences due to genetic effects and is important in the prediction of response rates from selection. • The square root of the variance is the standard deviation, which is the ave. deviation of each indiv. in the pop. from the pop. Average • Heritability est. – handout BRD 33
  • 5. Correlation • Quantifies a relationship between two variables • Measured between zero and one • Positive versus negative correlations • Multiple effects (Regression models)
  • 6. Animal Identification • Accurate identification • Written records • Accurate measurements of economic importance • ??? Registered or not • Two ID nos. , one for flock or indiv. ID and the other for registration ID (if reg.)
  • 7. Record Keeping • Determine clear goals • Allow for records such as pedigree, birth weight, singles or twins, weaning weight, etc. • Five categories of records: – REPRODUCTION, MATERNAL , GROWTH, WOOL AND CARCASS
  • 8. Estimated Breeding Value (EBV) • EBV = b (P- ave.P), where b is the heritability of a particular trait • Example; Ram A has a grease fleece weight of 15 # and the ave. flock grease fleece weight is 11 #, then 15-11 = selection differential [4] • 4 x .4 (h) = 1.6 # of grease fleece advantage for breeding Ram A
  • 9. EPD’s (Expected Progeny Differences) • Simply ½ of the EBV • The ave. EPD in a population is + or – from the average of the population. A comparison • Ex. If a ewe has a +.3 for no. of lambs born, then one would expect the progeny to produce .3 more lambs per lambing than the progeny of average ewes.
  • 10. EPD’s • Another ex. is two rams A and B • +.5 and +1.1 for fleece weight, respectively • Ram B progeny would be expected to have .6 pounds heavier fleeces than Ram A • Example 3; Ram has +1.0 and Ewe has +.5 for weaning weight, then the progeny will be 1.5 # more than the pop. ave. for WW
  • 11. EPD’s • Accuracy • Gives an idea of reliability of the estimat • Measured from 0-1 • An accuracy of .45 is not very reliable, whereas > .9 is considered reliable • Low accuracies are a result of limited information known about the parents or low numbers of progeny occur
  • 12. Generation Interval • The time lapse between birth of an animal and the birth of its replacement – usually 3- 4 years for sheep • A measurement of progress or rate of improvement • Genetic Improvement per year = heritability x selection differential divided by the generation interval
  • 13. Methods of Selection for Single Traits • Individual selection-selection on their own performance • Family selection-selection based on bloodlines; useful when (h) is low • Pedigree selection-similar to family and is dependent upon how closely related the ancestors. Useful when considering same sex
  • 14. Methods of Selection for Single Traits • Progeny Test- observing the performance of the offspring. Must be mated to several ewes and then look at the offspring. Best when looking at carcass traits • Combined Selection- uses more than one of the above mentioned methods
  • 15. Methods of Selection for Multiple Traits • Tandem Selection- focuses on multiple traits, yet one at a time. After the performance of one is achieved, then move to the next trait • Independent Culling- Set minimum standards for more than one trait at a time for the indiv. Cull any that does not meet the min. stds for any trait
  • 16. Methods of Selection for Multiple Traits • Selection Index- rank indiv. animals for two or more traits based on a combination effect • One the farm testing: • Primary focus is on the ewe flock • Therefore, traits such as prolificacy, weaning weight and fleece weight
  • 17. National Sheep Improvement Program • Oversees the promotion, funding, development and implementation of a national genetic evaluation program for sheep • Minimum criteria for record keeping • Lamb records: ID #, sire and dam ID, date of birth, sex, type of birth, & type of rearing. • Additional records are reproductive, growth and fleece traits
  • 18. NSIP • Focuses on: • Number of lambs born per ewe lambing • Body weight at 30,60,90,120 & 240 days • Grease &/or clean fleece wt. • Staple length • Fleece grade • Pounds of lamb weaned per ewe exposed per year
  • 19. Adj. Factors for No. of lambs born per lambing to a common ewe Age of Dam Adj. Factor 1 1.45 2 1.15 3 1.05 4 1.00 5, 6, & 7 .96, .96 & .95 8 .98 9+ 1.00
  • 20. Major Economically Important Traits • Reproductive Efficiency • Carcass Merit • Milk Production • Total Ewe Production- # lamb/ewe exposed • Mature Size • Hardiness & Adaptability • Wool Production • Growth Rate
  • 21. Selection for Growth • Growth is a very important trait, esp. for meat breeds • Adjusted 90 day weaning weight • If birth weight is known: • [Actual wt. - birth wt. x adj. Days (90)/ actual age ]+ birth wt. • If birth wt. is not known: • Actual wt. X adj. Age in days/age
  • 22. Selection for Fleece Traits • Grease and Clean Fleece Wts • Yield • Staple length- at least 3”/yr. • Fiber diameter • Crimp • Color • Density • Belly Wool-undesirable on sides
  • 23. Importance of Genetic Improvement in Seedstock Flocks • Most produce their own replacements • Therefore, genetic improvement from outside is by the Ram purchased • Theoretically, Genetic merit increases at the same rate as the genetic merit of the rams, yet because of generation intervals and replacement production usually improvement lags two generations behind
  • 24. Mating Systems • Purebreeding- common genetic group • Outbreeding- unrelated within a breed • Inbreeding-closely related with one or more common ancestors (more than 50% related) • Linebreeding-common ancesters, but not 50% related • Crossbreeding- mating of different breeds • Grading Up-enhanced concentration of crossbreeding to rams of a single breed
  • 25. Inbreeding Coefficients • Full brother mated to sister = .25 • Sire on daughter = .25 • Half brother to half sister = .125 • Sire on Mother = .5 • Therefore; > .5 has to be compounded over time and added generation to generation
  • 26. Crossbreeding • Heterosis or Hybrid Vigor • Superiority of the crossbred indiv. Relative to the average performance of the parents • Maximized when parents have no breed ancestry in common (3 breed terminal X)
  • 27. Crossbreeding Systems • Two Breed Terminal(50% Heterosis) • Three Breed Terminal (100%) • Three Breed Rotational (86 %) • Four Breed Rotational (93 %) • Two Breed Rotational • Roto-Terminal- combination of rotational and terminal systems. The poor ewes would still be used in a terminal crossing manner
  • 28. Inherited Defects • Many defects are recdessive in nature • Selection pressure is exercised against certain traits, esp. lethals and fleece defects • Dwarfism, Spider Syndrome, Jaw Defects, Rectal Prolapse, Inverted Eyelids, Cryptorchidism, Horns or Scurs, Face covering, Color, Skin Folds, Silky, Britch fibers, grey color, paralyzed limbs, earless, etc.
  • 29. Sheep Breeding and Reproduction • Improved lamb production • More lambs per lambing • More frequent lambing • Increased percent of total sheep nos. • Reducing death losses
  • 30. Sheep Breeding and Reproduction • breed dependent - seasonal breeding • Puberty-weight and age (70-100 # or 5-9 months) • Breed to lamb as 2 year olds, yet can breed as a yearling, but with less efficiency • Actually, greater production per lifetime from yearlings; Ramb. is later maturing and less effic. • gestation - 144-152 days (med wool less) • heat periods - 20 to 42 hours (no signs)
  • 31. Sheep Breeding and Reproduction • ovulation occurs late in heat cycle • 14- 20 day cycle ave. 16-17 days • Prolifacy - > 100% lamb crop • flushing ewes works in sheep, too • Factors affecting reprod. – daylight (<14 hrs/da), temp. ( <74 F or > 100), nutrition
  • 32. Hormonal Control • Progesterone (Progestins)- produced by the corpus luteum • Prostaglandins- induces the regression of the corpus luteum (ineffective in normally cycling ewes); lutalyse • Estrogen • Gonadotropins- GnRH, HCG, FSH & LH • Melatonin- regulatory of the seasonal breeder; produced by the pineal gland • ACTH
  • 33. Sheep Breeding and Reproduction • Lambing – > 4 sq. ft. pen for ewes, clean & dry – presentation of front legs – orphan lambs is not uncommon – 25% death loss is common – Feed ewes small amounts of water at first with oats, wheat bran and hay (small amounts)
  • 34. Sheep Breeding and Reproduction • Ram management • mating guide – ram lamb hand (20-25) pasture (25-35) – yearling or > hand (50-75) pasture (35-60) – use marker rams to detect bred ewes both at breeding and 60 days after breeding
  • 35. Replacement Ewe Selection Item 140 % Prolif 170 % Prolif No. ewes 100 100 Ewe lambs 70 85 Twin lambs 40 70 Tw lambs > ave. in WW 20 35 Ewe lambs exposed 20 30 Preg. Ewes 20
  • 36. Physiological changes of reproduction in the ewes • REPRO – 4 chart • Ewes in the south have longer breeding patterns • Sheep near the equator are less likely to have seasonal breeding
  • 37. Factors affecting reproduction in the ewe • Heredity • Age • Photoperiod (seasonal) • Temperature and humidity • Nutrition and Exercise • Parturition and lactation • Disease and parasites • Fertility of & assoc. with the ram
  • 38. Pregnancy Testing • Breeding Marks • Ultrasonic Scanning- best detected between 70 & 120 days • Bagging or Udder Palpation • Blood Progesterone- at the time of the next expected heat • PSPB- a protein called pregnancy-specific protein B after day 21 of breeding.
  • 39. Factors affecting the reproduction in the Ram • Breeding soundness exam • Palpation of the testicles, epididymis, and penis and visual appraisal of feet, legs, eyes and jaws. • Semen evaluation • Disease prevention • Heat stress
  • 40. Semen Collection • Components- seminal fluid and sperm • Quality of sperm – morphology and viability (percent live) • Methods- artificial vagina and electroejaculation • Semen handling-dilution characteristics: glucose or fructose, egg yolk, citrate or phosphate, antibitotics, glycerol
  • 41. Insemination • Natural- 3-5 billion sperm inseminated • AI – vaginal approach – 200 million “ • AI – cervical approach- 100 million “ • AI - Intruterine insemination via lapraroscopic surgery- 20 million • Time of insemination – vaginal or cervical = 12 to 18 hrs after onset of estrus • Synchronizing with progesterone sponge should be 48-58 hrs after removal
  • 42. Accelerated and Out-of-Season Lambing • Day length control (natural or artificial) • Considerations • Produce lambs when feeding conditions are favorable (growth rate and cost of feed) • Market lambs when lamb supply is low • Fertility and prolificacy
  • 43. Desirable traits for accelerated lambing • Ewes can breed year round • Ewes that can mate while lactacting • Ewes that have a good lambing rate (ie twinning) • Sires that produce a desirable market lamb and have the libido and fertility for conception year round