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LECTURE PRESENTATIONS
For CAMPBELL BIOLOGY, NINTH EDITION
Jane B. Reece, Lisa A. Urry, Michael L. Cain, Steven A. Wasserman, Peter V. Minorsky, Robert B. Jackson
© 2011 Pearson Education, Inc.
Lectures by
Erin Barley
Kathleen Fitzpatrick
An Introduction to Ecology and
the Biosphere
Chapter 52
Overview: Discovering Ecology
• Ecology is the scientific study of the
interactions between organisms and the
environment
determine distribution of organisms and
their abundance
includes observation and experimentation
How is this different from the environment?
© 2011 Pearson Education, Inc.
Figure 52.1
Figure 52.2
Global ecology
Landscape ecology
Ecosystem ecology
Community ecology
Population ecology
Organismal ecology
Global Ecology
• The biosphere is the global ecosystem, the sum
of all the planet’s ecosystems
• Global ecology examines the influence of
energy and materials on organisms across the
biosphere
© 2011 Pearson Education, Inc.
• Landscape ecology - exchanges
of energy, materials, and
organisms across multiple
ecosystems
© 2011 Pearson Education, Inc.
Figure 52.2b
Landscape ecology
• Ecosystem ecology
emphasizes energy flow
and chemical cycling
among the various biotic
and abiotic components
© 2011 Pearson Education, Inc.
Figure 52.2c
Ecosystem ecology
• Community ecology deals
with the whole array of
interacting species in a
community
© 2011 Pearson Education, Inc.
Figure 52.2d
Community ecology
• Population ecology
focuses on factors
affecting population size
over time
© 2011 Pearson Education, Inc.
Figure 52.2e
Population ecology
• Organismal ecology includes
physiological, evolutionary, and
behavioral ecology
© 2011 Pearson Education, Inc.
Figure 52.2f
Organismal ecology
Concept 52.1:
• Climate - the long-term prevailing weather
conditions in an area: temperature, precipitation,
sunlight, wind
• Macroclimate - global, regional, and landscape
level
• Microclimate –small, area (community of
organisms underneath a fallen log)
© 2011 Pearson Education, Inc.
Global Climate Patterns
• determined by solar energy and the
planet’s movement in space
• causes temperature variations, which
drive evaporation and the circulation
of air and water
• This causes latitudinal variations in
climate
© 2011 Pearson Education, Inc.
Latitudinal Variation in Sunlight Intensity
What does this mean? Why does it happen?
© 2011 Pearson Education, Inc.
Figure 52.3a
Latitudinal variation in sunlight intensity
90°N (North Pole)
60°N
30°N
23.5°N (Tropic of
Cancer
60°S
90°S (South Pole)
0° (Equator)
23.5°S (Tropic of
Capricorn)
30°S
Low angle of incoming sunlight
Atmosphere
Sun overhead at equinoxes
Low angle of incoming sunlight
• Rising air masses release water and cause high
precipitation, especially in the tropics
• Dry, descending air masses create arid climates,
especially near 30°
• Air flowing close to Earth’s surface creates
predictable global wind patterns
• Cooling trade winds blow from east to west in the
tropics; prevailing westerlies blow from west to
east in the temperate zones
© 2011 Pearson Education, Inc.
Figure 52.3b
Global air circulation and precipitation patterns
Westerlies
Northeast trades
Southeast trades
Westerlies
30°N
0°
66.5°N (Arctic Circle)
30°N
0°
30°S
60°N
60°S
66.5°S (Antarctic Circle)
Descending
dry air
absorbs
moisture.
Ascending
moist air
releases
moisture.
Regional and Local Effects on Climate
• Climate is affected by
seasonality, large bodies of
water, and mountains
© 2011 Pearson Education, Inc.
Seasonality
• Seasonal variations of light and
temperature increase steadily toward
the poles (WHY?)
• Belts of wet and dry air straddling the
equator shift throughout the year with
the changing angle of the sun
• Changing wind patterns affect ocean
currents
© 2011 Pearson Education, Inc.
Figure 52.4
March equinox
December
solstice
September equinox
60°N
30°S
30°N
0° (equator)
Constant tilt
of 23.5°
June solstice
The trip around
• What do we mean by the equinox?
When do they occur?
• What do we mean by the solstices?
When do they occur?
Bodies of Water
• Oceans, their currents, and large
lakes moderate the climate of nearby
terrestrial environments
• The Gulf Stream carries warm water
from the equator to the North Atlantic
© 2011 Pearson Education, Inc.
Figure 52.5
Indian
Ocean
Subtropical
Gyre
California Current
30°N North Pacific
Subtropical Gyre
30°S
Equator
South Pacific
Subtropical Gyre
Labrador Current
Gulf Stream
North Atlantic
Subtropical
Gyre
South
Atlantic
Subtropical
Gyre
Antarctic Circumpolar Current
• Why is there a breeze at the ocean
shoreline?
© 2011 Pearson Education, Inc.
Figure 52.6
Air flow
Ocean
Mountain
range
Leeward side
of mountains
Mountains
• Rising air releases moisture on the windward
side of a peak and creates a “rain shadow” as it
absorbs moisture on the leeward side
• Mountains affect the amount of sunlight reaching
an area
• In the Northern Hemisphere, south-facing slopes
receive more sunlight than north-facing slopes
• Every 1,000 m increase in elevation produces a
temperature drop of approximately 6C
© 2011 Pearson Education, Inc.
Smith Rock - Oregon
Rocky Mountains
Microclimate
• fine-scale differences in the environment that
affect light and wind patterns
• characterized by differences in
– Abiotic factors, including nonliving attributes
such as temperature, light, water, and nutrients
– Biotic factors, including other organisms that
are part of an individual’s environment
© 2011 Pearson Education, Inc.
Global Climate Change
• Predictions for change can be
based on previous changes
• Glacier retreat 16,000 years ago
caused tree distribution patterns
to change
• Specie dispersal could determine
the range and climate changes
© 2011 Pearson Education, Inc.
Figure 52.7
Current
range
Predicted
range
Overlap
(a) 4.5°C warming over next
century
(b) 6.5°C warming over next
century
Figure 52.8
Sweden
Finland
Expanded range in 1997
Range in 1970
Concept 52.2:
• Biomes are major life zones characterized by
vegetation type (terrestrial biomes) or physical
environment (aquatic biomes)
• Climate is very important in determining why
terrestrial biomes are found in certain areas
© 2011 Pearson Education, Inc.
Tropic of
Cancer
30°N
30°S
Tropic of Capricorn
Equator
Tropical forest
Savanna
Desert
Chaparral
Temperate grassland
Temperate broadleaf forest
Northern coniferous forest
Tundra
High mountains
Polar ice
Figure 52.9
• A climograph plots the temperature and
precipitation in a region
• Biomes are affected not just by average
temperature and precipitation, but also by the
pattern of temperature and precipitation through
the year
© 2011 Pearson Education, Inc.
Temperate
broadleaf
forest
Arctic and
alpine
tundra
Northern
coniferous
forest
Annual
mean
temperature
(°C)
Temperate grassland Tropical forest
30
15
0
15
Desert
Annual mean precipitation (cm)
0 400
100 200 300
Figure 52.10
General Features of Terrestrial Biomes
• Terrestrial biomes are often named for major
physical or climatic factors and for vegetation
• Terrestrial biomes usually grade into each other,
without sharp boundaries
• The area of intergradation, called an ecotone,
may be wide or narrow
© 2011 Pearson Education, Inc.
• Vertical layering is an important feature of
terrestrial biomes, and in a forest it might consist
of an upper canopy, low-tree layer, shrub
understory, ground layer of herbaceous plants,
forest floor, and root layer
• Layering of vegetation in all biomes provides
diverse habitats for animals
• Biomes are dynamic and usually exhibit
extensive patchiness
© 2011 Pearson Education, Inc.
• Similar characteristic can arise in
distant biomes through
convergent evolution
–For example, cacti in North
America and euphorbs in
African deserts appear similar
but are from different
evolutionary lineages
© 2011 Pearson Education, Inc.
Figure 52.11
Cereus peruvianus
Euphorbia canariensis
• Disturbance is an event that changes a
community
– frequent fires can kill woody plants and maintain
the characteristic vegetation of a savanna
– fires and outbreaks of pests create gaps in
forest that allow different species to grow
• Fire suppression has changed the vegetation of
the Great Plains
© 2011 Pearson Education, Inc.
Disturbance and Terrestrial Biomes
Terrestrial Biomes
• Terrestrial biomes can be characterized by
distribution, precipitation, temperature, plants,
and animals
© 2011 Pearson Education, Inc.
Tropical Forest
• equatorial and subequatorial regions
• tropical rain forests, rainfall is relatively
constant,
• tropical dry forests precipitation is highly
seasonal
• (25–29C)
• Vertical layering
• Most diverse biome
© 2011 Pearson Education, Inc.
A tropical rain forest in Borneo
Figure 52.12a
Desert
• Deserts 30C N/S and in the interior of
continents
• < 30 cm per year
• Deserts may be hot or cold
• Plants adapted to climate (C4 and CAM)
• Many nocturnal animals
© 2011 Pearson Education, Inc.
A desert in the southwestern
United States
Figure 52.12b
Savanna
• Equatorial and subequatorial regions
• precipitation is seasonal
• warm year-round (24–29C) but more
seasonally variable than the tropics
• Fire adapted plants
• Grasses and forbs
• Zebras, lions, hyena
© 2011 Pearson Education, Inc.
A savanna in Kenya
Figure 52.12c
Chaparral
• midlatitude coastal regions on several continents
• rainy winters and dry summers
• Summer hot (30C+) while fall, winter, and spring
are cool (10–12C)
• Shrubs, grasses, herbs (fire-resistant)
• Insects and small mammals
© 2011 Pearson Education, Inc.
An area of chaparral
in California
Figure 52.12d
Temperate Grassland
• found on many continents
• Precipitation is highly seasonal
• Winters cold (often below –10C) and dry
summers hot (often near 30C) and wet
• Grasses and forbs adapted to fire and drought
• Bison and wild horses
© 2011 Pearson Education, Inc.
Grasslands National Park,
Saskatchewan
Figure 52.12e
Northern Coniferous Forest
• taiga, spans northern North America and
Eurasia and is the largest terrestrial biome on
Earth
• Precipitation varies
• Winters are cold and long while summers may
be hot (e.g., Siberia ranges from –50C to 20C)
• Pine, spruce, fir
• Cone shape protects breakage from snow
• Moose, bear, migratory birds, Siberian tigers
© 2011 Pearson Education, Inc.
A forest in Norway
Figure 52.12f
Temperate Broadleaf Forest
• midlatitudes in the Northern Hemisphere, with
smaller areas in Chile, South Africa, Australia,
and New Zealand
• Large amount of precip in all seasons
• Winters average 0C, while summers are hot
and humid (near 35C)
• Vertical layering
• Deciduous trees
• Mammals, birds, insects
© 2011 Pearson Education, Inc.
Great Smoky Mountains
National Park in
North Carolina, in autumn
Figure 52.12g
Tundra
• Arctic;
• Alpine tundra exists on high mountaintops at all
latitudes
• Precipitation is low in arctic tundra, and higher
in alpine tundra
• Winters are long and cold (below –30C) while
summers are relatively cool (less than 10C)
• Permafrost
• Reindeer, musk, oxen
• Mosses, grasses
© 2011 Pearson Education, Inc.
Denali National Park, Alaska,
in autumn
Figure 52.12h
Concept 52.3:
• largest part of the biosphere in terms of area
• They show less latitudinal variation than
terrestrial biomes
• Marine biomes have salt concentrations of
about 3%
• The largest marine biome (oceans) cover about
75% of Earth’s surface and have an enormous
impact on the biosphere
© 2011 Pearson Education, Inc.
• Freshwater biomes
have salt concentrations of less
than 0.1%
closely linked to soils and the
biotic components of the
surrounding terrestrial biome
© 2011 Pearson Education, Inc.
Figure 52.13
(a) Zonation in a lake
(b) Marine zonation
Littoral
zone Limnetic
zone
Photic
zone
Benthic
zone
Aphotic
zone
Pelagic
zone
0
200 m
Continental
shelf
2,000
6,000 m
Abyssal
zone
Benthic
zone
Photic
zone
Intertidal zone
Neritic
zone Oceanic zone
Aphotic
zone
Pelagic
zone
• Thermocline - temperature boundary called the
separating the warm upper layer from the cold
deeper water
• turnover – mixing of oxygenated water from
surface with nutrient rich water on the bottom of
a lake
© 2011 Pearson Education, Inc.
Figure 52.14
Winter Spring
Thermocline
Autumn
0°
2°
4°C 4°C
4°
4°C
4°
4°C
22°
18°
8°
Summer
• Communities vary with depth, light
penetration, distance from shore, and
position in the pelagic or benthic zone
• Most occur in the relatively shallow photic
zone
• The aphotic zone in oceans is extensive,
but harbors little life
© 2011 Pearson Education, Inc.
Tropic of
Cancer
Tropic of
Capricorn
30°N
30°S
Equator
Oceanic pelagic and benthic zones
Intertidal zones
Estuaries
Coral reefs
Rivers
Lakes
Figure 52.15
Aquatic Biomes characterized
by:
physical environment
chemical environment
geological features
photosynthetic organisms
heterotrophs
© 2011 Pearson Education, Inc.
Lakes
• Oligotrophic lakes are nutrient-
poor and generally oxygen-rich
• Eutrophic lakes are nutrient-rich
and often depleted of oxygen if
ice covered in winter
© 2011 Pearson Education, Inc.
• littoral zone close to shore
(shallow and well lit)
• limnetic zone – further from
shore small drifting animals called
zooplankton graze on the
phytoplankton
© 2011 Pearson Education, Inc.
An oligotrophic lake in Grand
Teton National Park, Wyoming
A eutrophic lake in the Okavango
Delta, Botswana
Figure 52.16a
Wetlands
• a habitat that is inundated by water at
least some of the time and that supports
plants adapted to water-saturated soil
• have high organic production and
decomposition and have
• low dissolved oxygen
• Wetlands can develop in shallow basins,
along flooded river banks, or on the
coasts of large lakes and seas
© 2011 Pearson Education, Inc.
• most productive biomes on Earth
• lilies, cattails, sedges, tamarack, and black
spruce
• invertebrates and birds, as well as otters, frogs,
and alligators
• Humans have destroyed up to 90% of wetlands;
wetlands purify water and reduce flooding
© 2011 Pearson Education, Inc.
Figure 52.16b
A basin wetland in the United Kingdom
Streams and Rivers
• prominent physical characteristic of is current
• Headwaters are generally cold, clear, turbulent,
swift, and oxygen rich; they are often narrow and
rocky
• Downstream waters form rivers and are
generally warmer, more turbid, and more
oxygenated; they are often wide, meandering,
and have silty bottoms
© 2011 Pearson Education, Inc.
Figure 52.16c
A headwater stream in the Great
Smoky Mountains
The Loire river (in France) far
from its headwaters
Estuaries
• transition area between river and sea
• Salinity varies with the rise and fall of the tides
• nutrient rich and highly productive
• include a complex network of tidal channels,
islands, natural levees, and mudflats
© 2011 Pearson Education, Inc.
• Saltmarsh grasses and algae are the major
producers
• An abundant supply of food attracts marine
invertebrates, fish, waterfowl, and marine
mammals
© 2011 Pearson Education, Inc.
Figure 52.16d
An estuary in the southeastern United States
Intertidal Zones
• periodically submerged and exposed by the
tides
• organisms are challenged by variations in
temperature and salinity and by the mechanical
forces of wave action
• Oxygen and nutrient levels are high
• Substrate varies from rocky to sandy
© 2011 Pearson Education, Inc.
Rocky intertidal zone on the Oregon coast
Figure 52.16e
Oceanic Pelagic Zone
• constantly mixed by wind-driven oceanic
currents
• Oxygen levels are high
• Turnover renews nutrients in the photic zones;
year-round stratification in tropical oceans leads
to lower nutrient concentrations
• 70% of Earth’s surface
© 2011 Pearson Education, Inc.
• Phytoplankton (diatoms) and
zooplankton are the dominant
organism; also found are free-
swimming animals
• Zooplankton (protists, worms,
copepods, krill, jellies, and
invertebrate larvae
• squids, fishes, sea turtles, and
marine mammals
© 2011 Pearson Education, Inc.
Figure 52.16f
Open ocean off the island of Hawaii
Fringing reef
Barrier Reef
Figure 52.16g
A coral reef in the Red Sea
Marine Benthic Zone
• The marine benthic zone consists of the
seafloor below the surface waters of the coastal,
or neritic, zone and the offshore pelagic zone
• Organisms in the very deep benthic (abyssal)
zone are adapted to continuous cold and
extremely high water pressure
• Substrate is mainly soft sediments; some areas
are rocky
© 2011 Pearson Education, Inc.
• Shallow areas contain seaweeds and filamentous
algae
• Deep-sea hydrothermal vents of volcanic origin on
mid-oceanic ridges are surrounded by unique
chemoautotrophic prokaryotes, as well as
echinoderms and arthropods
• Neritic benthic communities include invertebrates
and fishes
• Overfishing and dumping of waste have depleted
fish populations
© 2011 Pearson Education, Inc.
Figure 52.16h
A deep-sea hydrothermal vent community
Behavior and Habitat Selection
• Some organisms do not occupy all of their
potential range
• Species distribution may be limited by habitat
selection behavior
© 2011 Pearson Education, Inc.
Biotic Factors
• Biotic factors that affect the distribution of
organisms may include
– Predation
– Herbivory
• For example, sea urchins can limit the
distribution of seaweeds
– Competition
© 2011 Pearson Education, Inc.
Both limpets and urchins
removed
Only urchins
removed
RESULTS
Seaweed
cover
(%)
Only limpets removed
Control (both urchins
and limpets present)
Sea urchin
Limpet
100
80
60
40
20
0
February
1983
August
1983
August
1982
February
1984
Figure 52.20
Abiotic Factors
• Abiotic factors affecting distribution of organisms
include
– Temperature
– Water
– Sunlight
– Wind
– Rocks and soil
• Most abiotic factors vary in space and time
© 2011 Pearson Education, Inc.
Temperature
• Environmental temperature is an important factor
in distribution of organisms because of its effects
on biological processes
• Cells may freeze and rupture below 0°C, while
most proteins denature above 45°C
• Mammals and birds expend energy to regulate
their internal temperature
© 2011 Pearson Education, Inc.
Water and Oxygen
• Water availability in habitats is another important
factor in species distribution
• Desert organisms exhibit adaptations for water
conservation
• Water affects oxygen availability as oxygen
diffuses slowly in water
• Oxygen concentrations can be low in deep
oceans and deep lakes
© 2011 Pearson Education, Inc.
Salinity
• Salt concentration affects water balance of
organisms through osmosis
• Most aquatic organisms are restricted to
either freshwater or saltwater habitats
• Few terrestrial organisms are adapted to high-
salinity habitats
© 2011 Pearson Education, Inc.
Sunlight
• Light intensity and quality (wavelength) affect
photosynthesis
• Water absorbs light, thus in aquatic
environments most photosynthesis occurs near
the surface
• In deserts, high light levels increase temperature
and can stress plants and animals
© 2011 Pearson Education, Inc.
Figure 52.21
Rocks and Soil
• Many characteristics of soil limit distribution of
plants and thus the animals that feed on them
– Physical structure
– pH
– Mineral composition
© 2011 Pearson Education, Inc.
Why is species X absent from an area?
Does dispersal limit its distribution?
Does behavior limit its distribution?
Do biotic factors (other species)
limit its distribution?
Do abiotic factors limit
its distribution?
No
No
No
Yes
Yes
Yes
Physical
factors
Chemical
factors
Area inaccessible or
insufficient time
Habitat selection
Predation, parasitism,
competition, disease
Temperature, light, soil
structure, fire, moisture,
etc.
Water, oxygen, salinity,
pH, soil nutrients, etc.
Figure 52.UN01
Figure 52.UN02
100
Mean
height
(cm)
50
0
Altitude
(m)
1,000
3,000
2,000
0
Sierra Nevada
Great Basin
Plateau
Seed collection sites
Figure 52.UN03

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Introduction to Ecosystem and Ecology and the Biospehere

  • 1. LECTURE PRESENTATIONS For CAMPBELL BIOLOGY, NINTH EDITION Jane B. Reece, Lisa A. Urry, Michael L. Cain, Steven A. Wasserman, Peter V. Minorsky, Robert B. Jackson © 2011 Pearson Education, Inc. Lectures by Erin Barley Kathleen Fitzpatrick An Introduction to Ecology and the Biosphere Chapter 52
  • 2. Overview: Discovering Ecology • Ecology is the scientific study of the interactions between organisms and the environment determine distribution of organisms and their abundance includes observation and experimentation How is this different from the environment? © 2011 Pearson Education, Inc.
  • 4. Figure 52.2 Global ecology Landscape ecology Ecosystem ecology Community ecology Population ecology Organismal ecology
  • 5. Global Ecology • The biosphere is the global ecosystem, the sum of all the planet’s ecosystems • Global ecology examines the influence of energy and materials on organisms across the biosphere © 2011 Pearson Education, Inc.
  • 6. • Landscape ecology - exchanges of energy, materials, and organisms across multiple ecosystems © 2011 Pearson Education, Inc.
  • 8. • Ecosystem ecology emphasizes energy flow and chemical cycling among the various biotic and abiotic components © 2011 Pearson Education, Inc.
  • 10. • Community ecology deals with the whole array of interacting species in a community © 2011 Pearson Education, Inc.
  • 12. • Population ecology focuses on factors affecting population size over time © 2011 Pearson Education, Inc.
  • 14. • Organismal ecology includes physiological, evolutionary, and behavioral ecology © 2011 Pearson Education, Inc.
  • 16. Concept 52.1: • Climate - the long-term prevailing weather conditions in an area: temperature, precipitation, sunlight, wind • Macroclimate - global, regional, and landscape level • Microclimate –small, area (community of organisms underneath a fallen log) © 2011 Pearson Education, Inc.
  • 17. Global Climate Patterns • determined by solar energy and the planet’s movement in space • causes temperature variations, which drive evaporation and the circulation of air and water • This causes latitudinal variations in climate © 2011 Pearson Education, Inc.
  • 18. Latitudinal Variation in Sunlight Intensity What does this mean? Why does it happen? © 2011 Pearson Education, Inc.
  • 19. Figure 52.3a Latitudinal variation in sunlight intensity 90°N (North Pole) 60°N 30°N 23.5°N (Tropic of Cancer 60°S 90°S (South Pole) 0° (Equator) 23.5°S (Tropic of Capricorn) 30°S Low angle of incoming sunlight Atmosphere Sun overhead at equinoxes Low angle of incoming sunlight
  • 20. • Rising air masses release water and cause high precipitation, especially in the tropics • Dry, descending air masses create arid climates, especially near 30° • Air flowing close to Earth’s surface creates predictable global wind patterns • Cooling trade winds blow from east to west in the tropics; prevailing westerlies blow from west to east in the temperate zones © 2011 Pearson Education, Inc.
  • 21. Figure 52.3b Global air circulation and precipitation patterns Westerlies Northeast trades Southeast trades Westerlies 30°N 0° 66.5°N (Arctic Circle) 30°N 0° 30°S 60°N 60°S 66.5°S (Antarctic Circle) Descending dry air absorbs moisture. Ascending moist air releases moisture.
  • 22. Regional and Local Effects on Climate • Climate is affected by seasonality, large bodies of water, and mountains © 2011 Pearson Education, Inc.
  • 23. Seasonality • Seasonal variations of light and temperature increase steadily toward the poles (WHY?) • Belts of wet and dry air straddling the equator shift throughout the year with the changing angle of the sun • Changing wind patterns affect ocean currents © 2011 Pearson Education, Inc.
  • 24. Figure 52.4 March equinox December solstice September equinox 60°N 30°S 30°N 0° (equator) Constant tilt of 23.5° June solstice
  • 25. The trip around • What do we mean by the equinox? When do they occur? • What do we mean by the solstices? When do they occur?
  • 26. Bodies of Water • Oceans, their currents, and large lakes moderate the climate of nearby terrestrial environments • The Gulf Stream carries warm water from the equator to the North Atlantic © 2011 Pearson Education, Inc.
  • 27. Figure 52.5 Indian Ocean Subtropical Gyre California Current 30°N North Pacific Subtropical Gyre 30°S Equator South Pacific Subtropical Gyre Labrador Current Gulf Stream North Atlantic Subtropical Gyre South Atlantic Subtropical Gyre Antarctic Circumpolar Current
  • 28. • Why is there a breeze at the ocean shoreline? © 2011 Pearson Education, Inc.
  • 30. Mountains • Rising air releases moisture on the windward side of a peak and creates a “rain shadow” as it absorbs moisture on the leeward side • Mountains affect the amount of sunlight reaching an area • In the Northern Hemisphere, south-facing slopes receive more sunlight than north-facing slopes • Every 1,000 m increase in elevation produces a temperature drop of approximately 6C © 2011 Pearson Education, Inc.
  • 31. Smith Rock - Oregon
  • 33. Microclimate • fine-scale differences in the environment that affect light and wind patterns • characterized by differences in – Abiotic factors, including nonliving attributes such as temperature, light, water, and nutrients – Biotic factors, including other organisms that are part of an individual’s environment © 2011 Pearson Education, Inc.
  • 34.
  • 35.
  • 36.
  • 37. Global Climate Change • Predictions for change can be based on previous changes • Glacier retreat 16,000 years ago caused tree distribution patterns to change • Specie dispersal could determine the range and climate changes © 2011 Pearson Education, Inc.
  • 38. Figure 52.7 Current range Predicted range Overlap (a) 4.5°C warming over next century (b) 6.5°C warming over next century
  • 40. Concept 52.2: • Biomes are major life zones characterized by vegetation type (terrestrial biomes) or physical environment (aquatic biomes) • Climate is very important in determining why terrestrial biomes are found in certain areas © 2011 Pearson Education, Inc.
  • 41. Tropic of Cancer 30°N 30°S Tropic of Capricorn Equator Tropical forest Savanna Desert Chaparral Temperate grassland Temperate broadleaf forest Northern coniferous forest Tundra High mountains Polar ice Figure 52.9
  • 42. • A climograph plots the temperature and precipitation in a region • Biomes are affected not just by average temperature and precipitation, but also by the pattern of temperature and precipitation through the year © 2011 Pearson Education, Inc.
  • 43. Temperate broadleaf forest Arctic and alpine tundra Northern coniferous forest Annual mean temperature (°C) Temperate grassland Tropical forest 30 15 0 15 Desert Annual mean precipitation (cm) 0 400 100 200 300 Figure 52.10
  • 44. General Features of Terrestrial Biomes • Terrestrial biomes are often named for major physical or climatic factors and for vegetation • Terrestrial biomes usually grade into each other, without sharp boundaries • The area of intergradation, called an ecotone, may be wide or narrow © 2011 Pearson Education, Inc.
  • 45. • Vertical layering is an important feature of terrestrial biomes, and in a forest it might consist of an upper canopy, low-tree layer, shrub understory, ground layer of herbaceous plants, forest floor, and root layer • Layering of vegetation in all biomes provides diverse habitats for animals • Biomes are dynamic and usually exhibit extensive patchiness © 2011 Pearson Education, Inc.
  • 46. • Similar characteristic can arise in distant biomes through convergent evolution –For example, cacti in North America and euphorbs in African deserts appear similar but are from different evolutionary lineages © 2011 Pearson Education, Inc.
  • 48. • Disturbance is an event that changes a community – frequent fires can kill woody plants and maintain the characteristic vegetation of a savanna – fires and outbreaks of pests create gaps in forest that allow different species to grow • Fire suppression has changed the vegetation of the Great Plains © 2011 Pearson Education, Inc. Disturbance and Terrestrial Biomes
  • 49. Terrestrial Biomes • Terrestrial biomes can be characterized by distribution, precipitation, temperature, plants, and animals © 2011 Pearson Education, Inc.
  • 50. Tropical Forest • equatorial and subequatorial regions • tropical rain forests, rainfall is relatively constant, • tropical dry forests precipitation is highly seasonal • (25–29C) • Vertical layering • Most diverse biome © 2011 Pearson Education, Inc.
  • 51. A tropical rain forest in Borneo Figure 52.12a
  • 52. Desert • Deserts 30C N/S and in the interior of continents • < 30 cm per year • Deserts may be hot or cold • Plants adapted to climate (C4 and CAM) • Many nocturnal animals © 2011 Pearson Education, Inc.
  • 53. A desert in the southwestern United States Figure 52.12b
  • 54. Savanna • Equatorial and subequatorial regions • precipitation is seasonal • warm year-round (24–29C) but more seasonally variable than the tropics • Fire adapted plants • Grasses and forbs • Zebras, lions, hyena © 2011 Pearson Education, Inc.
  • 55. A savanna in Kenya Figure 52.12c
  • 56. Chaparral • midlatitude coastal regions on several continents • rainy winters and dry summers • Summer hot (30C+) while fall, winter, and spring are cool (10–12C) • Shrubs, grasses, herbs (fire-resistant) • Insects and small mammals © 2011 Pearson Education, Inc.
  • 57. An area of chaparral in California Figure 52.12d
  • 58. Temperate Grassland • found on many continents • Precipitation is highly seasonal • Winters cold (often below –10C) and dry summers hot (often near 30C) and wet • Grasses and forbs adapted to fire and drought • Bison and wild horses © 2011 Pearson Education, Inc.
  • 60. Northern Coniferous Forest • taiga, spans northern North America and Eurasia and is the largest terrestrial biome on Earth • Precipitation varies • Winters are cold and long while summers may be hot (e.g., Siberia ranges from –50C to 20C) • Pine, spruce, fir • Cone shape protects breakage from snow • Moose, bear, migratory birds, Siberian tigers © 2011 Pearson Education, Inc.
  • 61. A forest in Norway Figure 52.12f
  • 62. Temperate Broadleaf Forest • midlatitudes in the Northern Hemisphere, with smaller areas in Chile, South Africa, Australia, and New Zealand • Large amount of precip in all seasons • Winters average 0C, while summers are hot and humid (near 35C) • Vertical layering • Deciduous trees • Mammals, birds, insects © 2011 Pearson Education, Inc.
  • 63. Great Smoky Mountains National Park in North Carolina, in autumn Figure 52.12g
  • 64. Tundra • Arctic; • Alpine tundra exists on high mountaintops at all latitudes • Precipitation is low in arctic tundra, and higher in alpine tundra • Winters are long and cold (below –30C) while summers are relatively cool (less than 10C) • Permafrost • Reindeer, musk, oxen • Mosses, grasses © 2011 Pearson Education, Inc.
  • 65. Denali National Park, Alaska, in autumn Figure 52.12h
  • 66. Concept 52.3: • largest part of the biosphere in terms of area • They show less latitudinal variation than terrestrial biomes • Marine biomes have salt concentrations of about 3% • The largest marine biome (oceans) cover about 75% of Earth’s surface and have an enormous impact on the biosphere © 2011 Pearson Education, Inc.
  • 67. • Freshwater biomes have salt concentrations of less than 0.1% closely linked to soils and the biotic components of the surrounding terrestrial biome © 2011 Pearson Education, Inc.
  • 68. Figure 52.13 (a) Zonation in a lake (b) Marine zonation Littoral zone Limnetic zone Photic zone Benthic zone Aphotic zone Pelagic zone 0 200 m Continental shelf 2,000 6,000 m Abyssal zone Benthic zone Photic zone Intertidal zone Neritic zone Oceanic zone Aphotic zone Pelagic zone
  • 69. • Thermocline - temperature boundary called the separating the warm upper layer from the cold deeper water • turnover – mixing of oxygenated water from surface with nutrient rich water on the bottom of a lake © 2011 Pearson Education, Inc.
  • 70. Figure 52.14 Winter Spring Thermocline Autumn 0° 2° 4°C 4°C 4° 4°C 4° 4°C 22° 18° 8° Summer
  • 71. • Communities vary with depth, light penetration, distance from shore, and position in the pelagic or benthic zone • Most occur in the relatively shallow photic zone • The aphotic zone in oceans is extensive, but harbors little life © 2011 Pearson Education, Inc.
  • 72. Tropic of Cancer Tropic of Capricorn 30°N 30°S Equator Oceanic pelagic and benthic zones Intertidal zones Estuaries Coral reefs Rivers Lakes Figure 52.15
  • 73. Aquatic Biomes characterized by: physical environment chemical environment geological features photosynthetic organisms heterotrophs © 2011 Pearson Education, Inc.
  • 74. Lakes • Oligotrophic lakes are nutrient- poor and generally oxygen-rich • Eutrophic lakes are nutrient-rich and often depleted of oxygen if ice covered in winter © 2011 Pearson Education, Inc.
  • 75. • littoral zone close to shore (shallow and well lit) • limnetic zone – further from shore small drifting animals called zooplankton graze on the phytoplankton © 2011 Pearson Education, Inc.
  • 76. An oligotrophic lake in Grand Teton National Park, Wyoming A eutrophic lake in the Okavango Delta, Botswana Figure 52.16a
  • 77. Wetlands • a habitat that is inundated by water at least some of the time and that supports plants adapted to water-saturated soil • have high organic production and decomposition and have • low dissolved oxygen • Wetlands can develop in shallow basins, along flooded river banks, or on the coasts of large lakes and seas © 2011 Pearson Education, Inc.
  • 78. • most productive biomes on Earth • lilies, cattails, sedges, tamarack, and black spruce • invertebrates and birds, as well as otters, frogs, and alligators • Humans have destroyed up to 90% of wetlands; wetlands purify water and reduce flooding © 2011 Pearson Education, Inc.
  • 79. Figure 52.16b A basin wetland in the United Kingdom
  • 80. Streams and Rivers • prominent physical characteristic of is current • Headwaters are generally cold, clear, turbulent, swift, and oxygen rich; they are often narrow and rocky • Downstream waters form rivers and are generally warmer, more turbid, and more oxygenated; they are often wide, meandering, and have silty bottoms © 2011 Pearson Education, Inc.
  • 81. Figure 52.16c A headwater stream in the Great Smoky Mountains The Loire river (in France) far from its headwaters
  • 82. Estuaries • transition area between river and sea • Salinity varies with the rise and fall of the tides • nutrient rich and highly productive • include a complex network of tidal channels, islands, natural levees, and mudflats © 2011 Pearson Education, Inc.
  • 83. • Saltmarsh grasses and algae are the major producers • An abundant supply of food attracts marine invertebrates, fish, waterfowl, and marine mammals © 2011 Pearson Education, Inc.
  • 84. Figure 52.16d An estuary in the southeastern United States
  • 85. Intertidal Zones • periodically submerged and exposed by the tides • organisms are challenged by variations in temperature and salinity and by the mechanical forces of wave action • Oxygen and nutrient levels are high • Substrate varies from rocky to sandy © 2011 Pearson Education, Inc.
  • 86. Rocky intertidal zone on the Oregon coast Figure 52.16e
  • 87. Oceanic Pelagic Zone • constantly mixed by wind-driven oceanic currents • Oxygen levels are high • Turnover renews nutrients in the photic zones; year-round stratification in tropical oceans leads to lower nutrient concentrations • 70% of Earth’s surface © 2011 Pearson Education, Inc.
  • 88. • Phytoplankton (diatoms) and zooplankton are the dominant organism; also found are free- swimming animals • Zooplankton (protists, worms, copepods, krill, jellies, and invertebrate larvae • squids, fishes, sea turtles, and marine mammals © 2011 Pearson Education, Inc.
  • 89. Figure 52.16f Open ocean off the island of Hawaii
  • 92. Figure 52.16g A coral reef in the Red Sea
  • 93. Marine Benthic Zone • The marine benthic zone consists of the seafloor below the surface waters of the coastal, or neritic, zone and the offshore pelagic zone • Organisms in the very deep benthic (abyssal) zone are adapted to continuous cold and extremely high water pressure • Substrate is mainly soft sediments; some areas are rocky © 2011 Pearson Education, Inc.
  • 94. • Shallow areas contain seaweeds and filamentous algae • Deep-sea hydrothermal vents of volcanic origin on mid-oceanic ridges are surrounded by unique chemoautotrophic prokaryotes, as well as echinoderms and arthropods • Neritic benthic communities include invertebrates and fishes • Overfishing and dumping of waste have depleted fish populations © 2011 Pearson Education, Inc.
  • 95. Figure 52.16h A deep-sea hydrothermal vent community
  • 96. Behavior and Habitat Selection • Some organisms do not occupy all of their potential range • Species distribution may be limited by habitat selection behavior © 2011 Pearson Education, Inc.
  • 97. Biotic Factors • Biotic factors that affect the distribution of organisms may include – Predation – Herbivory • For example, sea urchins can limit the distribution of seaweeds – Competition © 2011 Pearson Education, Inc.
  • 98. Both limpets and urchins removed Only urchins removed RESULTS Seaweed cover (%) Only limpets removed Control (both urchins and limpets present) Sea urchin Limpet 100 80 60 40 20 0 February 1983 August 1983 August 1982 February 1984 Figure 52.20
  • 99. Abiotic Factors • Abiotic factors affecting distribution of organisms include – Temperature – Water – Sunlight – Wind – Rocks and soil • Most abiotic factors vary in space and time © 2011 Pearson Education, Inc.
  • 100. Temperature • Environmental temperature is an important factor in distribution of organisms because of its effects on biological processes • Cells may freeze and rupture below 0°C, while most proteins denature above 45°C • Mammals and birds expend energy to regulate their internal temperature © 2011 Pearson Education, Inc.
  • 101. Water and Oxygen • Water availability in habitats is another important factor in species distribution • Desert organisms exhibit adaptations for water conservation • Water affects oxygen availability as oxygen diffuses slowly in water • Oxygen concentrations can be low in deep oceans and deep lakes © 2011 Pearson Education, Inc.
  • 102. Salinity • Salt concentration affects water balance of organisms through osmosis • Most aquatic organisms are restricted to either freshwater or saltwater habitats • Few terrestrial organisms are adapted to high- salinity habitats © 2011 Pearson Education, Inc.
  • 103. Sunlight • Light intensity and quality (wavelength) affect photosynthesis • Water absorbs light, thus in aquatic environments most photosynthesis occurs near the surface • In deserts, high light levels increase temperature and can stress plants and animals © 2011 Pearson Education, Inc.
  • 105. Rocks and Soil • Many characteristics of soil limit distribution of plants and thus the animals that feed on them – Physical structure – pH – Mineral composition © 2011 Pearson Education, Inc.
  • 106. Why is species X absent from an area? Does dispersal limit its distribution? Does behavior limit its distribution? Do biotic factors (other species) limit its distribution? Do abiotic factors limit its distribution? No No No Yes Yes Yes Physical factors Chemical factors Area inaccessible or insufficient time Habitat selection Predation, parasitism, competition, disease Temperature, light, soil structure, fire, moisture, etc. Water, oxygen, salinity, pH, soil nutrients, etc. Figure 52.UN01