The 1UZ-FE engine is a 4.0-liter V8 with 32 valves and DOHC design. It produces 250 horsepower at 5,600 rpm and 260 ft-lbs of torque at 4,400 rpm. Key features include a compact design achieved through a scissors gear mechanism and lightweight materials, low noise and vibration, and high reliability ensured by features like plastic tightening bolts. The engine is controlled by an ECU to maintain peak performance and efficiency.