AllShepsPubs-1-18-16

Wayne D. Shepperd
Career Publications
(1) Shepperd, W. D. 1973. An instrument for measuring tree crown width. USDA
Forest Service Research Note RM-229, 3 p. Rocky Mountain Forest and Range
Experiment Station, Fort Collins, Colorado.
(2) Noble, D. L., and W. D. Shepperd. 1973. Gray-headed juncos important in first
season mortality of Engelmann spruce. Journal of Forestry 71(12):763-765. (refereed).
(3) Alexander, Robert R., and Wayne D. Shepperd. 1974. Ceratonia siliqua L. (carob).
In Seeds of Woody Plants of the United States (C. S. Schopmeyer, technical
coordinator). U.S. Department of Agriculture Handbook 450, p. 303-304.
(4) Shepperd, Wayne D. 1974. Growth of Engelmann spruce seedlings under selected
temperature environments. Masters report. Department of Forest and Wood Sciences,
College of Forestry and Natural Resources, Colorado State University, Fort Collins,
Colorado. 61 p.
(5) Alexander, Robert R., Wayne D. Shepperd, and Carleton B. Edminster. 1975. Yield
tables for managed stands of spruce-fir in the central Rocky Mountains. USDA Forest
Service Research Paper RM-134, 20 p. Rocky Mountain Forest and Range Experiment
Station, Fort Collins, Colorado.
(6) Shepperd, Wayne D., and Daniel L. Noble. 1976. Germination, survival, and growth
of lodgepole pine under simulated precipitation regimes: a greenhouse study. USDA
(7) Shepperd, Wayne D. 1977. Variation in growth of Engelmann spruce seedlings
under selected temperature environments. In Proceedings, Western Forest Genetics
Association Annual Meeting (Gilbert H. Fechner and Frederick J. Deneke, editors),
Colorado State University, Fort Collins, Colorado. August 2-4, 1977. p.5.
(8) Noble, Daniel L., Carleton B. Edminster, and Wayne D. Shepperd. 1978. Effects of
watering treatments on germination, survival, and growth of Douglas-fir: a greenhouse
study. USDA Forest Service Research Note RM-357, 6 p. Rocky Mountain Forest and
Range Experiment Station, Fort Collins, Colorado.
(9) Noble, Daniel L., Carleton B. Edminster, and Wayne D. Shepperd. 1978. Effects of
watering treatments on germination survival, and growth of ponderosa pine. USDA
(10) Shepperd, Wayne D. 1980. Hand-held calculator programs for the field forester.
USDA Forest Service General Technical Report RM-76, 17 p. Rocky Mountain Forest
and Range Experiment Station, Fort Collins, Colorado.

(11) Shepperd, Wayne D., Richard M. Jeffers, and Frank Ronco, Jr. 1981. An
Engelmann spruce seed source study in the central Rockies. USDA Forest Service
Research Paper RM-231, 5 p. Rocky Mountain Forest and Range Experiment Station,
Fort Collins, Colorado.
(12) Shepperd, Wayne D. 1981. Variation in growth of Engelmann spruce seedlings
under selected temperature environments. USDA Forest Service Research Note RM-404,
3 p. Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(13) Shepperd, Wayne D. 1982. Stand characteristics of Rocky Mountain aspen. In N.
E. DeByle, editor. Proceedings, Situation management of two Intermountain species:
aspen and coyotes, a symposium. Part I Aspen, April 23-24, 1981. Utah State
University, p 22-30. Logan, Utah.
(14) Shepperd, Wayne D., and Orville Engelby. 1983. Rocky Mountain aspen. In
Silvicultural systems for the major forest types of the United States. U.S. Department of
Agriculture Handbook No. 445, p 77-70. Washington, D.C.
(15) Shepperd, Wayne D., and Robert R. Alexander. 1983. Overview to silvicultural
systems in the central Rocky Mountains. USDA Forest Service Publication RMTT-1, 32
p. Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(16) Shepperd, Wayne D., and Robert R. Alexander. 1983. Silviculture of spruce-fir
forests in the central Rocky Mountains. USDA Forest Service Publication RMTT-2, 44
p. Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(17) Shepperd, Wayne D., and Robert R. Alexander. 1983. Silviculture of lodgepole
pine in the central Rocky Mountains. USDA Forest Service Publication RMTT-3, 36 p.
Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(18) Shepperd, Wayne D., Robert R. Alexander, and Frank Ronco, Jr. 1983. Silviculture
of ponderosa pine in the central and southern Rocky Mountains. USDA Forest Service
Publication RMTT-4, 36 p. Rocky Mountain Forest and Range Experiment Station, Fort
Collins, Colorado.
(19) Shepperd, Wayne D., and H. Todd Mowrer. 1984. Whole stand volume tables for
Rocky Mountain aspen. USDA Forest Service Research Note RM-440, 5 p. Rocky
Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(20) Alexander, Robert R., and Wayne D. Shepperd. 1984. Silvical characteristics of
Engelmann spruce. USDA Forest Service General Technical Report RM-114, 38 p.
Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(21) Alexander, Robert R., Raymond C. Shearer, and Wayne D. Shepperd. 1984.
Silvical characteristics of subalpine fir. USDA Forest Service General Technical Report

RM-115, 29 p. Rocky Mountain Forest and Range Experiment Station, Fort Collins,
Colorado.
(22) Shepperd, Wayne D., and John R. Jones. 1985. Nurse crop. In Norbert V. DeByle
and Robert P. Winokur, editors. Aspen: ecology and management in the western United
States. USDA Forest Service General Technical Report RM 119, p 181-184. Rocky
(23) Jones, John R., Wayne D. Shepperd, Norbert V. DeByle, and Robert P. Winokur.
1985. Management overview. In Norbert V. DeByle and Robert P. Winokur, editors.
Aspen: ecology and management in the western United States. USDA Forest Service
General Technical Report RM 119, p 193-195. Rocky Mountain Forest and Range
(23a) Schier, George A, Wayne D. Shepperd, and John R. Jones, Regeneration. In
Norbert V. DeByle and Robert P. Winokur, editors. Aspen: ecology and management in
the western United States. USDA Forest Service General Technical Report RM 119, p
197-208. Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
(24) Jones, John R., Wayne D. Shepperd, and Norbert V. DeByle. 1985. Intermediate
treatments. In Norbert V. DeByle and Robert P. Winokur, editors. Aspen: ecology and
management in the western United States. USDA Forest Service General Technical
Report RM 119, p 209-216. Rocky Mountain Forest and Range Experiment Station, Fort
Collins, Colorado.
(25) Jones, John R., and Wayne D. Shepperd. 1985. Rotations. In Norbert V. DeByle
(26) Jones, John R., and Wayne D. Shepperd. 1985. Harvesting. In Norbert V. DeByle
(27) Edminster, Carleton B., H. Todd Mowrer, and Wayne D. Shepperd. 1985. Site
index tables for aspen in Colorado and Wyoming. USDA Forest Service Research Note
RM-453, 4p. Rocky Mountain Forest and Range Experiment Station, Fort Collins,
Colorado.
(28) Alexander, Robert R., Charles A. Troendle, Merrill R. Kaufmann, Wayne D.
Shepperd, Glenn L. Crouch, and Ross K. Watkins. 1985. The Fraser Experimental
Forest, Colorado: Research program and published research 1937-1985. USDA Forest
Service General Technical Report RM-118, 46p. Rocky Mountain Forest and Range

(29) Hinds, T. E., W. D. Shepperd, and T. H. Laurent. 1985. Dothiora polyspora on
aspen. Plant Disease 69:1100. (refereed).
(30) Shepperd, Wayne D. and Sue E. McElderry. 1985. Open-pollinated progeny test of
ponderosa pine in the Black Hills: Ten-year results. Western Journal of Applied
Forestry 1(3):79-83. (refereed).
(31) Shepperd, Wayne D. 1985. Aspen ecology and management in the central and
southern Rocky Mountains. p. 233-236. In: Foresters' future: Leaders or followers?
Proceedings of the 1985 Society of American Foresters National Convention [Fort
Collins, Colo., July 28-31, 1985]. 455 p. Society of American Foresters, Bethesda, Md.
(32) Shepperd, Wayne D. 1986. Silviculture of aspen in the Central Rocky Mountains.
USDA Forest Service Publication RMTT-7, 38 p. Rocky Mountain Forest and Range
Experiment Station, Fort Collins, Colo.
(33) Shepperd, Wayne D. 1986. Aspen management in the Southern Rockies: The
silvicultural perspective. In: Proceedings, 36th Western Forest Insect Work Conference,
Boulder, Colorado.
(34) Shepperd, Wayne D. 1987. Silviculture research in Rocky Mountain aspen. p. 25-
29. In: Management of subalpine forests: Building on 50 years of research. Proceedings
of a technical conference [Silver Creek, Colo. July 6-9, 1987]. USDA Forest Service
General Technical Report RM-149 p 25-29. Rocky Mountain Forest and Range
Experiment Station, Fort Collins, Colo.
(35) Hinds, Thomas E. and Wayne D. Shepperd. 1987. Aspen sucker damage and defect
in Colorado clearcut areas. USDA Forest Service Research Paper RM-278, 12p. Rocky
Mountain Forest and Range Experiment Station, Fort Collins, Colo.
(36) Mowrer, H. Todd and Wayne D. Shepperd. 1987. Field measurement of age in
quaking aspen in the central Rocky Mountains. USDA Forest Service Research Note
RM-476, 4p. Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colo.
(37) Crouch, Glenn L. and Wayne D. Shepperd. 1987. Establishing a research base for
multifunctional management of aspen in the central Rocky Mountains. p. 191-200. In
Proceedings of the National Silviculture Workshop [Sacramento, CA, May 11-14, 1987].
322p. USDA Forest Service, Washington D. C.
(38) Shepperd, Wayne D. 1990. Management of aspen in the central Rocky Mountains:
An application of multiple use silviculture. p.73-78. In: Adams, Roy D., ed. Aspen
Symposium '89, proceedings [Duluth, MN July 25-27, 1989]. USDA Forest Service
General Technical Report NC-40 348p. North Central Forest Experiment Station, St.
Paul, MN.

(39) Shepperd, Wayne D. 1990. A classification of quaking aspen in the central Rocky
Mountains based on growth and stand characteristics. Western Journal of Applied
Forestry 5(3):69-75. (refereed).
(40) Alexander, Robert R. and Wayne D. Shepperd. 1990. Picea Engelmannii Parry ex
Engelm. Engelmann spruce. p. 187-203. In: Burns, Russell M. and Barbara H.
Honkala, tech. coords. Silvics of North America: 1. Conifers. Agriculture Handbook
654 Vol. 1, 675p. USDA Forest Service, Washington DC.
(41) Alexander, Robert R. , Raymond C. Shearer and Wayne D. Shepperd. 1990. Abies
lasiocarpa (Hook) Nutt. Subalpine fir. p60-70. In: Burns, Russell M. and Barbara H.
Honkala, tech. coords. Silvics of North America: 1. Conifers. Agriculture Handbook
654 vol 1, 675p. USDA Forest Service, Washington DC.
(42) Shepperd, Wayne D. 1991. Vegetative reproduction and early clonal growth of
aspen. Ph.D. dissertation, Colorado State University, Fort Collins, CO. 68p.
(43) Shepperd, Wayne D. 1991. Silviculture research for multiple resource management
of aspen in the central Rocky Mountains. In Proceedings of the National Silviculture
Workshop [Petersburg, AK, July 10-13, 1989]. 216p. USDA Forest Service, Washington
D. C.
(44) Jacobi, William R. and Wayne D. Shepperd. 1991. Fungi associated with sprout
mortality in aspen clearcuts in Colorado and Arizona. USDA Forest Service Research
Note RM-513, 5p. Rocky Mountain Forest and Range Experiment Station, Fort Collins,
Colo.
(45) Shepperd, W.D.; Troendle, C.E.; Edminster, C.E. 1992. Linking water and growth
and yield models to evaluate management alternatives in subalpine ecosystems. In:
Murphy, Dennis, comp. Getting to the future through silviculture - Workshop
proceedings; May 6-9, 1991; Cedar City UT. Gen. Tech. Rep. INT-291. Ogden UT: U.S.
Department of Agriculture, Forest Service, Intermountain Research Station. p42-48.
(46) Shepperd, Wayne D. 1992. A computer graphics technique for visualizing spatial
disturbance and succession in old-growth forests. pp. 195-200. In: Kaufmann, Merrill
R.; Moir, W. H.; Bassett, Richard L., Tech. Coords. Old-growth forests in the southwest
and Rocky Mountain regions: Proceedings of a workshop. Gen. Tech. Report RM-213.
Ft. Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest
and Range Experiment Station.
(47) Shepperd, Wayne D. 1993. The effect of commercial harvest activities on root
compaction and suckering of aspen. Western Journal of Applied Forestry 8(2): 62-66.
(refereed).
(48) Shepperd, Wayne D. 1993. Initial growth, development, and clonal dynamics of
regenerated aspen in the Rocky Mountains. Res. Pap. RM 312. Fort Collins, CO: U.S.

Department of Agriculture, Forest Service. Rocky Mountain Forest and Range
Experiment Station. 8p.
(49) Popovich, Steve J.; Shepperd, Wayne D.; Reichert, Donald W.; Cone, Michael A.
1993. Flora of the Fraser Experimental Forest, Colorado. Gen. Tech. Rep. RM-233. Fort
Collins, CO: U.S. Department of Agriculture, Forest Service 62p. Rocky Mountain
Forest and Range Experiment Station. 62p.
(50) Shepperd, W.D.; Smith, F.W. 1993. The role of near-surface lateral roots in the life
cycle of aspen in the central Rocky Mountains. Forest Ecol. Mgmt. 61:157-170.
(refereed).
(51) Haas, Timothy C.; Mowrer, H. Todd; Shepperd, Wayne D. 1994. Modeling aspen
stand growth with a temporal Bayes network. A.I. Applications 8(1): 15-28. (refereed).
(52) Shepperd, Wayne D. and M. L. Fairweather. 1994. Impact of large ungulates in
restoration of aspen communities in a southwestern ponderosa pine ecosystem. In:
Covington, W. Wallace; DeBano, Leonard F.; technical coordinators. Sustainable
ecological systems: implementing an ecological approach to land management.
Proceedings of a conference July 13-15, Flagstaff AZ. Gen. Tech. Rep. RM-247. Ft.
Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest
and Range Experiment Station. p 344-347.
(53) Shepperd, Wayne D. 1995. Seeing the trees: the GRAFGYM stand graphing
system. In: Foley, Louise H. Silviculture: From the cradle of forestry to Ecosystem
Management. Proceedings of the National Silvicultural Workshop, November 1-4, 1993,
Hendersonville, NC. Gen. Tech. Rep. SE-88. Ashville, NC: U.S. Department of
Agriculture, Forest Service, Southeastern Experiment Station. p 229-233.
(54) Brown, Peter M.; Shepperd, Wayne D.; Brown, Christopher C.; Mata, Stephen A.;
McClain, Douglas L. 1995. Oldest known Engelmann spruce, Fraser Experimental
Forest, Central Rocky Mountains. Res. Note RM-RN-534. Ft. Collins, CO: U.S.
Department of Agriculture, Forest Service, Rocky Mountain Forest and Range
Experiment Station. 6p.
(55) Brown, Peter M.; Shepperd, Wayne D. 1995. Engelmann spruce tree-ring
chronologies from Fraser Experimental Forest, Colorado: Potential for a long-term
temperature reconstruction in the Central Rocky Mountains. In: Tinus, Richard W., tech.
ed. Interior West global change workshop. April 25-27, 1995; Fort Collins, CO: U.S.
Department of Agriculture, Forest Service, Rocky Mountain forest and Range
Experiment Station. p 23-26.
(56) Troendle, C.A.; Shepperd, W.D.; Edminster, C.B. 1995. Linking water and growth
and yield models to evaluate management alternatives in subalpine ecosystems. In:
Ahuja, L.; Leppert, K.; Rojas, K.;Seely, E. eds. Workshop on computer applications in

water management, May 23-25, 1995. GPAC Pub. No. 154. Fort Collins, CO: Great
Plains Agricultural Council, Colorado State University. p. 179-183.
(57) Regan, Claudia M.; Shepperd, Wayne D.; Obedzinski, Robert A. 1995. Application
of the forest vegetation simulator in evaluating management for old-growth
characteristics in southwestern mixed conifer forests. In: Eskew, Lane G. comp. forest
health through silviculture: Proceedings of the 1995 National Silviculture Workshop,
Mescalero, New Mexico, May 8-11, 1995. Gen. Tech. Rep. RM-GTR-267. U.S.
Department of Agriculture, Forest Service, Rocky Mountain Forest and Range
Experiment Station. p. 183-194.
(58) Shepperd, Wayne D. 1996. Silviculture and management of aspen in the west. Nat.
Res. News, January 1996. p. 10-11.
(59) Shepperd, Wayne D. 1996. Response of aspen root suckers to regeneration methods
and post-harvest protection. Res. Pap. RM-RP-324. Fort Collins, CO: U.S. Department
of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station.
8p.
(60) Shepperd, Wayne D.; Edminster, Carleton B. 1997. The values of long-term
silvicultural research studies. In: Proceedings of National Silviculture Workshop, May
19-22, 1997, Warren, PA. Gen. Tech. Rep. GTR-NE-238. Radnor, PA. U.S. Department
of Agriculture, Forest Service, Northeastern Forest Experiment Station. p110-114.
(61) Ferguson,D.E., Applegate, V.J., Aune, P.S., Carlson, C.E., Geier-Hayes, K.,
Graham, R.T., Jacobsen, G.L., Jain, T.B., Powell, D.C., Shepperd, W.D., Sloan, J.P. and
A. Youngblood. 1997. Communicating the role of silviculture and Forest Service
silviculture research in the lnterior West. In: Communicating the role of silviculture in
managing the national forests: Proceedings of the National Silviculture Workshop. 1997
May 19-22; Warren, PA.: Gen. Tech. Rep. NE-238. Radnor, PA: U.S. Department of
Agriculture, Forest Service, Northeastern Forest Experiment Station. 104-109.
(62) Brown, Peter M.; Shepperd, Wayne D.; Mata, Stephen A.; McClain, Douglas L.
1998. Longevity of windthrown logs in a subalpine forest of central Colorado. Can. J.
For. Res. 28:1-5. (refereed).
(63) Graham, R.L.; Huff, D.D.; Kaufmann, M.R.; Shepperd, W.D.; Troendle, C.A.;
Lynch, D.; Sheehan, J. 1999. Bioenergy and watershed restoration in the mountainous
regions of the west: What are the environmental/community issues? Proceedings:
Bioenergy98 Conference. October 7, 1998, Madison WI (web published at:
http://www.esd.ornl.gov/bfdp/papers/bioen98/graham.html)
(64) Baker, Dan L.; Andelt, William F.; Burnham, Kenneth P.; Shepperd, Wayne D.
1999. Effectiveness of Hot Sauce and Deer Away repellents for deterring elk browsing of
aspen sprouts. J. Wild. Man. 64(4):1327-1336. (refereed).

(65) Brown, Peter M.; Kaufmann, Merrill R.; Shepperd, Wayne D. 1999. Long-term,
landscape patterns of past fire events in a montane ponderosa pine forest of central
Colorado. Landscape Ecology 14:513-532. (refereed).
(66) Kent, Brian; Shepperd, Wayne D.; Shields, Deborah J. 2000. The Colorado Front
Range Ecosystem Management Research Project: Accomplishments to date. In: Smith,
Helen Y., ed. 2000. The Bitterroot Ecosystem Management Research Project: What we
have learned: Symposium proceedings; 1999 May 18-20; Missoula, MT. Proceedings
RMRS-P-17. Ogden, UT. U.S. Department of Agriculture, Forest Service, Rocky
Mountain Research Station. p119-124.
(67) Shepperd, Wayne D; Binkley, Dan; Bartos, Dale L.; Stohlgren, Thomas J.; and
Eskew, Lane G., compilers. 2001. Sustaining aspen in western landscapes: Symposium
Proceedings; 13-15 June 2000; Grand Junction, CO. Proceedings RMRS-P-18. Fort
Collins, CO: U.S. Department of Agriculture, forest Service, Rocky Mountain Research
Station. 460 p.
(68) Shepperd, Wayne D. 2001. Manipulations to regenerate aspen ecosystems. In:
Shepperd, Wayne D; Binkley, Dan; Bartos, Dale L.; Stohlgren, Thomas J.; and Eskew,
Lane G., compilers. 2001. Sustaining aspen in western landscapes: Symposium
Proceedings; 13-15 June 2000; Grand Junction, CO. Proceedings RMRS-P-18. Fort
Station. p355-366. (refereed)
(69) Shepperd, Wayne D.; Bartos, Dale L.; Mata, Stephen A. 2001. Above- and below-
ground effects of aspen clonal regeneration and succession to conifers. Can. J. For. Res.
31: 739-745. (refereed).
(70) Hollenstein, Kurt; Graham, Robin L.; Shepperd, Wayne D. 2001. Biomass flow in
western forests: Simulating the effects of fuel reduction and pre-settlement restoration
treatments. J. Forestry 99(10): 12-19. (refereed).
(71) Negron, Jose F.; Shepperd, Wayne D.; Mata, Steve A.; Popp, John B.; Asherin,
Lance A.; Schoettle, Anna W.; Schmid, John M.; Leatherman, David A. 2001. Solar
treatments for reducing survival of mountain pine beetle in infested ponderosa pine logs.
Res. Pap. RMRS-RP-30. Ft. Collins, CO. U.S. Department of Agriculture, Forest
Service, Rocky Mountain Research Station. 11p.
(72) Brown, Peter M. and Shepperd, Wayne D. 2001. Fire history and fire climatology
along a 5º gradient in latitude in Colorado and Wyoming, USA. Paleobotanist 50
(2001):133-140. (referreed)
(73) Shepperd, Wayne D.; Battaglia, Michael A. 2002. Ecology, Management, and
Silviculture of Black Hills Ponderosa Pine. USDA Forest Service, Rocky Mountain
Research Station, General Technical Report RMRS-97. 110 p.

(74) Shepperd, W.D.; Asherin, L.A.; Edminster, C.B. 2003. Using Individual Tree
Selection silviculture to Restore Northern Goshawk Habitat: Lessons from a
Southwestern Study. In: Proceedings: U.S.F.S National Silviculture Workshop, Hood
River OR, May 7-10, 2000. Gen. Tech. Rept PNW-GTR-546. Portland, OR: U.S.
Department of Agriculture, Pacific Northwest Research Station. p. 1-9. (refereed)
(75) Rummer, B.; Prestemon, J.; May, D.; Vissage, J.; McRoberts, r.; Liknes, G.;
Shepperd, W.D.;Reutebuch, S.; Barbour, J.; Fried, J.; Stokes, B.; Bilek, E.; Skog, K.
2003. A strategic assessment of forest biomass and fuel reduction treatments in western
states. U.S. Department of Agriculture, Forest Service, WO. Report. Web published at
http://www.fs.fed.us/research/infocenter.html. 18p.
(76) Martinson, Erik; Omi, Phillip; Shepperd, Wayne. 2003. Part 3: Effects of fuel
treatments on fire severity. In: Graham, RussellT. Tech. Ed. 2003. Hayman fire case
study. Gen. Tech. Rep. RMRS-GTR-114. Ogden, UT: U.S. Department of Agriculture,
forest Service, Rocky Mountain Research Station. p. 96-126.
(77) Massman, W.J.; Shepperd, W.D.; Frank, J.M.; Platten, M.J. 2003. In situ soil
temperature and heat flux measurements during controlled surface burns at a southern
Colorado forest site. In: Omi, Philip N.; Joyce, Linda A. Tech.Coords. Fire, fuel
treatments, and ecological restoration, proper place, appropriate time: Symposium
Proceedings: April 16,18, Ft. Collins, CO. Proceedings RMRS-P-29. Fort Collins, CO:
U.S. Department of Agriculture, forest Service, Rocky Mountain Research Station. p 67-
85.
(78) Shepperd, W.D.; Gallup, Sarah. 2003. Fuel Treatment Performance-Fire Hazard
Reduction – Introduction, Session 1. In: Omi, Philip N.; Joyce, Linda A. Tech.Coords.
Fire, fuel treatments, and ecological restoration, proper place, appropriate time:
Symposium Proceedings: April 16,18, Ft. Collins, CO. Proceedings RMRS-P-29. Fort
Station. p 5.
(79) Massmann, W.J.; Frank, J.M.; Massman, S.M.; Shepperd, W.D. 2003. Performance
of high temperature heat flux plates and soil moisture probes during controlled surface
fires. Fifth Symposium on Fire and Forest Meteorology. Paper 1B.1. Orlando, FL.
November 16-20, 2003. Boston MS, American Meteorological Society. (Available at:
http://ams.confex.com/ams/pdfpapers/65302.pdf)
(80) Shepperd, Wayne D.; Eskew, Lane G., compilers. 2004. Silviculture in special
places: Proceedings of the National Silviculture Workshop; 2003 September 8-11;
Granby, CO. Proceedings RMRS-P-34. Fort Collins, CO: U.S. Department of
Agriculture, Forest Service, Rocky Mountain Research Station. 255 p.
(81) Bonnet, V. H.; Schoettle, A.W.; Shepperd, W.D. 2004. Spatial distribution of
ponderosa pine seedlings along environmental gradients within burned areas in the Black
Hills, south Dakota. In: Shepperd, Wayne D.; Eskew, Lane G., compilers. 2004.

Silviculture in special places: Proceedings of the National Silviculture Workshop; 2003
September 8-11; Granby, CO. Proceedings RMRS-P-34. Fort Collins, CO: U.S.
Department of Agriculture, Forest Service, Rocky Mountain Research Station. p. 93-101.
(82) Shepperd, Wayne D. Asherin, Lance A.; mata, Stephen A.; McClain, Douglas L.
2004. The cutting methods demonstration study at Fraser Experimental Forest. In:
Shepperd, Wayne D.; Eskew, Lane G., compilers. 2004. Silviculture in special places:
Proceedings of the National Silviculture Workshop; 2003 September 8-11; Granby, CO.
Proceedings RMRS-P-34. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Research Station. p. 102-109
(83) Miles, P.D.; McRoberts, R.E.; Skog, K.E.; Shepperd, W.D.; Vissage, J.S.; Stokes,
B.J. 2004 Accessing and extending the utility of United States Forest Inventory data.
Proceedings of the 15th
International Workshop on Database and Expert Systems
Applications pp 576-580. August 30- September 3, 2004, Zaragoza Spain, IEEE
Computer Society, Los Alamitos, CA 1014pp.
(84) Shepperd, W.D. 2004. Techniques to restore aspen forests in the Western U.S.
Transactions of the Western Section of the Wildlife Society. 40:52-60.
(85) Shepperd, W.D.; Reichert, D.L.; Mata, S.A. 2004. Overwinter storage of
carbohydrate in aspen. Transactions of the Western Section of the Wildlife Society.
40:45-48.
(86) Bonnet, V.H.; Schoettle, A.W.; Shepperd, W.D. 2005. Postfire environmental
conditions influence the spatial pattern of regeneration for Pinus ponderosa. Can. J.
For.Res. 35:37-47.
(87) USDA Forest Service. 2005. A strategic assessment of forest biomass and fuel
reduction treatments in Western States. Gen. Tech. Rep. RMRS-GTR-149. Fort Collins
CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station.
17p. (Shepperd was one of 17 contributors to this report).
(88) Battaglia, Michael A.; Dodson, Jonathan M.; Shepperd, Wayne D.; Platten, Mark J.;
Tallmadge, Owen M. 2005. Colorado Front Range fuel photo series. Gen. Tech. Rep.
RMRS-GTR-155WWW. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Research Station. 39 p.
(89) Shepperd, W.D.; Mata, S.A. 2005. Planting Aspen to Rehabilitate Riparian Areas: A
Pilot Study. Res Note RMRS-RN-26. Ft. Collins, CO, U.S. Department of Agriculture,
Forest Service, Rocky Mountain Research Station 5 p.
(90) Boyden, Suzanne; Binkley, Dan; Shepperd, Wayne 2005. Spatial and temporal
patterns in structure, regeneration, and mortality of an old-growth ponderosa pine forest
in the Colorado Front Range. Forest Ecology and Management 219:43-55.

(91) Shepperd, W., K. Abt, R.J. Barbour, R. Fight, R. Huggett, P. Miles, E. Reinhardt, K.
Skog, The fuel treatment evaluator - a silvicultural approach to reducing fire hazard. In:
Proceedings, Society of American Foresters National Convention, October 19-22, 2005,
Ft. Worth, TX. Bethesda MD. Society of American Foresters. (web published at:
http://store.safnet.org/merchant.mv)
(92) Keyser, Tara L.; Smith, Frederick W.; Shepperd, Wayne D. 2005. Trembling aspen
response to a mixed-severity wildfire in the Black Hills, South Dakota, USA. Canadian
Journal of Forest Research. 35: 2679-2684.
(93) Lentile, Leigh B.; Smith Frederick W.; Shepperd, Wayne D. 2005. Patch structure,
fire-scar formation, and tree regeneration in a large mixed-severity fire in the South
Dakota Black Hills, USA. Canadian Journal of Forest Research. 35: 2875-2885.
(94) Shepperd, W.D.; Edminster, C.B.; Mata, S.A. 2006. Long-Term Seedfall,
Establishment, Survival and Growth of Natural and Planted Ponderosa Pine in the
Colorado Front Range Western Journal of Applied Forestry. 21(1):19-26. (Referred).
(95) Massman, WJ, JM Frank, AE Jimenez Esquilin, ME Stromberger, and WD
Shepperd. 2006. Long term consequences of a controlled slash burn and slash mastication
to soil moisture and CO2 at a southern Colorado site. In: 27th Conference on Agricultural
and Forest Meteorology, AMS, Boston. (Manuscript available on CD, paper 2.2)
(96) Skog, Kenneth E.; Barbour, R. James; Abt, Karen L.; Bilek, E.M. (Ted); Burch,
Frank; Fight, Roger D.; Hugget, Robert J.; Miles, Patrick D.; Reinhardt, Elizabeth D.;
Shepperd, Wayne D. 2006. Evaluation of silvicultural treatments and biomass use for
reducing fire hazard in western states. Research Paper FPL-RP-634. Madison, WI: U.S.
Department of Agriculture, Forest Service, Forest Products Laboratory. 29 p.
(97) Miles, Patrick D.; Skog, Kenneth E.; Shepperd, Wayne D.; Reinhardt, Elizabeth D.;
Fight, Roger D. 2006. Broad-scale assessment of fuel treatment opportunities.
Proceedings of the sixth annual forest inventory and analysis symposium; Denver, Colo.,
September 21-24. 2004. Gen. Tech. Report WO 70. Washington, D.C.: U.S. Dept. of
Agriculture, Forest Service; p. 29-34. (http://www.treesearch.fs.fed.us/pubs/14241).
(98) Shepperd, Wayne D; Rogers, Paul C.; Burton, David; Bartos, Dale L. 2006.
Ecology, biodiversity, management, and restoration of aspen in the Sierra Nevada. Gen.
Tech. Rep. RMRS-GTR-178. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Research Station. 122p.
(99) Rowell, Eric; Seielstad, Carl; Vierling, Lee; Queen, Lloyd; Shepperd, Wayne. 2006.
Using laser altimetry-based segmentation to refine automated tree identification in
managed forests of the Black Hills, South Dakota. Photogrammetric Engineering &
Remote Sensing. 72 (12): 1379-1388.

(100) Keyser, T.L., Smith, F.W., Lentile, L.B., and Shepperd, W.D. 2006. Modeling
postfire mortality of ponderosa pine following a mixed-severity wildfire in the Black
Hills: The role of tree morphology and direct fire effects. Forest Science 52(5): 530-539.
(101) Lentile, L.B.; Smith, Frederick W.; Shepperd, Wayne D. 2006. The influence of
topography and forest structure on patterns of mixed-severity fire in ponderosa pine
forest of the South Dakota Black Hills, USA. International Journal of Wildland Fire. 15,
557–566.
(102) Shepperd, Wayne D. 2007. SDI-Flex: A New Technique of Allocating Growing
Stock for Developing Treatment Prescriptions in Uneven-Aged Forest Stands. In:
Powers, R. Ed. Proceedings of the 2005 National Silviculture Workshop, Tahoe City,
CA, June 6-10, 2005. PSW-GTR-203. Alburn, CA: USDA Forest Service, Pacific
Southwest Experiment Station. p 171-180.
(103) Jimnez-Esquilin, AE, ME Stromberger, WJ Massman, JM Frank, and WD
Shepperd. 2007. Microbial community structure and activity in a Colorado Rocky
Mountain forest soil scarred by slash pile burning. Soil Biology and Biochemistry 39
(2007) 1111–1120.
(104) Rogers, Paul C.; Shepperd, Wayne D.; Bartos, Dale L. 2007. Aspen in the Sierra
Nevada: Regional Conservation of a Continental Species. Natural Areas Journal 27 (2):
183-193.
(105) M.E. Hunter, W.D. Shepperd, L.B. Lentile, J.E. Lundquist, M.G. Andreu, J.L.
Butler, and F.W. Smith. 2007. A comprehensive guide to fuels treatment practices for
ponderosa pine in the Black Hills, Colorado Front Range and Southwest. Gen. Tech. Rep.
RMRS-GTR-198. Fort Collins, CO: U.S. Department of Agriculture, Forest Service,
Rocky Mountain Research Station 93p..
(106) Hood, Sharon M.; Miller, Melanie; Long, Donald; Ryan, Kevin C.; Battaglia,
Mike; Shepperd, Wayne D.; Bartos, Dale; Tausch, Robin J.; Kitchen, Stanley G.;
McArthur, E. Durant; Brooks, Matt; Esque, Todd; Duck, Tim. 2007. Fire ecology and
management of the major ecosystems of southern Utah. Gen. Tech. Rep. RMRS-GTR-
202. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain
Research Station. 110 p.
(107) Lezberg, Ann L.; Battaglia, Michael A.; Shepperd, Wayne D. Decades-old
silvicultural treatments influence surface wildfire severity and post-fire nitrogen
availability in a ponderosa pine forest. Forest Ecology and Management 255: 49-61.
(108) Worrall JJ, Egeland L, Eager T, Mask RA, Johnson EW, Kemp PA, Shepperd WD.
2008. Rapid mortality of Populus tremuloides in southwestern Colorado, USA. Forest
Ecology and Management 255(3-4): 686-696.

(109) Shepperd, Wayne D. 2008. Ginko biloba L. Ginko, maidenhair-tree, or Kew-tree.
In: Bonner,F.T.; Karrfalt, R.P. Eds. The Woody Plant Seed Manual. Agric. Handbook
No. 727. Washington, DC. U.S. Department of Agriculture, Forest Service. 1223p.
(110) Shepperd, Wayne D. 2008. Tamarix chinensis Lour. Saltcedar, or five-stamen
tamarisk. In: Bonner,F.T.; Karrfalt, R.P. Eds. The Woody Plant Seed Manual. Agric.
Handbook No. 727. Washington, DC. U.S. Department of Agriculture, Forest Service.
1223p.
(111) Shepperd, Wayne D. 2008. Ceratonia siliqua L. Carob. In: Bonner,F.T.; Karrfalt,
R.P. Eds. The Woody Plant Seed Manual. Agric. Handbook No. 727. Washington, DC.
U.S. Department of Agriculture, Forest Service. 1223p.
(112) Shepperd, Wayne D. 2008. Garrya Dougl. Silktassel. In: Bonner,F.T.; Karrfalt,
R.P. Eds. The Woody Plant Seed Manual. Agric. Handbook No. 727. Washington, DC.
U.S. Department of Agriculture, Forest Service. 1223p.
(113) Keyser, T.L., Lentile, L.B, Smith, F.W., and Shepperd, W.D. 2008. Changes in
forest structure following a mixed-severity wildfire in ponderosa pine forests of the Black
Hills, SD, USA. Forest Science 54(3): 328-338.
(114) Huggett, Robert J.; Abt, Karen L.; Shepperd, Wayne. 2008. Efficacy of mechanical
fuel treatments for reducing wildfire hazard. Forest Policy and Economics 10: 408-414.
(115) Battaglia, Mike A; Smith, Frederick W.; Shepperd, Wayne D. 2008. Can prescribed
fire be used to maintain fuel treatment effectiveness over time in Black Hills ponderosa
pine forests? Forest Ecology and Management 256 (2008) 2029–2038.
(116) Kaufmann M.R., G.H. Aplet, M. Babler, W.L. Baker, B. Bentz, M. Harrington, .C.
Hawkes, L. Stroh Huckaby, M.J. Jenkins, D.M. Kashian, R.E. Keane, D. Kulakowski, C.
McHugh, J. Negron, J. Popp, W.H. Romme, T. Schoennagel, W. Shepperd, F.W. Smith,
E. Kennedy Sutherland, D. Tinker, and T.T. Veblen. 2008. The status of our scientific
understanding of lodgepole pine and mountain pine beetles – a focus on forest ecology
and fire behavior. The Nature Conservancy, Arlington, VA. GFI technical report 2008-2.
(117) Keyser, Tara L.; Smith, Frederick W.; Shepperd, Wayne D. 2009. Short-term
impact of post-fire salvage logging on regeneration, hazardous fuel accumulation, and
understorey development in ponderosa pine forests of the Black Hills, SD, USA.
International Journal ofWildland Fire. 18, 451–458
(118) Musselman, R .C.; Smith, F. W.; Shepperd, W. D.; Asherin, L. A.; Gee, B. W.
2009. The use of landscape fabric and supplemental irrigation to enhance survival and
growth of woody perennials planted on reclaimed surface mine lands. In: Barnhisel, R. I.,
ed. Proceedings 2009 National Meeting of the American Society of Mining and
Reclamation: Revitalizing the Environment: Proven Solutions and Innovative

Approaches; Billings, MT, May 30-June 5, 2009. Lexington, KY: American Society of
Mining and Reclamation. 23 p.
(119) Keyser, Tara L.; Smith Frederick W. ; Shepperd, Wayne D. 2010. Growth response
of Pinus ponderosa following a mixed-severity wildfire in the Black Hills, South Dakota.
Western Journal of Applied Forestry. 25(2): 49-54.
(120) Bartos, Dale L.; Shepperd, Wayne D., comps. 2010. The aspen mortality summit;
December 18 and 19, 2006; Salt Lake City, UT. Proc. RMRS-P-60WWW. Fort Collins,
CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station.
21 p.
(121) Musselman, Robert C.; Shepperd, Wayne D.; Smith, Frederick W.; Asherin, Lance
A.; Gee, Brian W. 2012. Response of transplanted aspen to irrigation and weeding on a
Colorado reclaimed surface coal mine. Res. Pap. RMRS-RP-101. Fort Collins, CO: U.S.
Department of Agriculture, Forest Service, Rocky Mountain Research Station. 20 p.
(122)Musselman, R.C.; Shepperd, W.D.; Smith, F.W.; Asherin, L.A. 2014. Survival and
growth of aspen and serviceberry on reclaimed surface mine land with landscape fabric
and irrigation. J. Am. Soc. Mining Reclamation. 3-1:16-40.
(123) Shepperd, W.D., Smith, F.W., and Pelz, K. 2015. Group clearfell harvest can
promote regeneration of aspen forests affected by sudden aspen decline in western
Colorado. Forest Science 61. http://dx.doi.org/10.5849/forsci.14-101.
(124) Dudley, Megan; Negron, Jose F.; Tisserat, Ned A.; Shepperd, Wayne D.; Jacobi,
William R. 2915. Influence of climate on the growth of quaking aspen (Populus
tremuloides Michx.) in Colorado and southern Wyoming. Can. J. For. Res. 45: 1546–
1563. http://dx.doi.org/10.1139/cjfr-2015-009207/2015; DOI:10.1139/cjfr-2015-0092.
(125) Mackes,Kurt; Shepperd, Wayne; Jennings, Christopher. 2005. Evaluating the
bending properties of clear wood specimens produced from small-diameter ponderosa
pine trees. Forest Products J. 55(10): 72-80.
(126) Buechling, Arne; Martin, Patrick H.; Canham, Charles D.; Shepperd, Wayne D.;
and Battaglia, Mike A. 2016. Climate drivers of seed production in Picea engelmannii
and response to warming temperatures in the southern Rocky Mountains. J. Ecology
104:1051-1062.
Published Abstracts
Worrall JJ, Mask RA, Eager T, Egeland L, Shepperd WD. 2008. Sudden aspen decline in
southwest Colorado. Phytopathology 98(Supplement):S173

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