Using the following list of hominin species, summarize the known trajectory of human biological and cultural evolution involving the five key traits that distinguish Homo sapiens from the other living Great Apes: Ardipithecus Ramidus, Australopithecus afarensis, Homo ergaster, and Homo heidelbergensis. Approximately when did each of these traits appear? What kinds of behaviors are we able to infer from the fossil and archaeological evidence, even if there is not direct evidence of it?.