I was born in New York City in 1935 and grew up in suburban New York, Connecticut, and western Pennsylvania, moving there in 1950. I attended the College of William and Mary from 1954-1956 and graduated from Stanford University in 1958 with a degree in Biological Sciences. In 1962 while studying at Tulane University, I became interested in Mexican fishes during an internship and focused my dissertation on the evolutionary problem of species of Chirostoma being restricted to one river system in Mexico. In 1963 I spent the summer in central Mexico collecting fishes with Salvador Contreras to fill gaps in previous collections and visited a limnological station to make collections from a lake.