This document provides an annotated catalogue of the carabid beetles of Albania. It summarizes all available literature records and material from various European museums and private collections that was revised by the author. In total, the catalogue includes 543 species, 103 genera, and 31 tribes of Carabidae found in Albania. Several dozen taxa are added for the first time to the fauna of either the Balkan Peninsula or Albania. The catalogue also presents some taxonomic changes based on a study of types and other specimens. It is intended to improve knowledge of the systematics and distribution of carabid beetles in the Balkans.

1. ANNOTATED CATALOGUE
of the
Carabid Beetles
of Albania
(Coleoptera: Carabidae)
by
B.V.Guéorguiev
T
his catalogue summarizes all available literature records
and a lot of new material revised or determined by the
author in various European museums and private collec-
tions. At present the carabid fauna of Albania can be considered
as well-studied. The catalogue includes references to or data for
543 species, 103 genera and 31 tribes of Carabidae. Several dozen
taxa of specific or supraspecific rank are added for the first time
to the fauna of either the Balkan Peninsula or Albania. A moderate
taxonomic contribution based on a study of types, topotypes or
series of previously problematic species or subspecies from
Albania and the neighbouring countries is also presented; as a
result, 38 various taxonomic changes (new synonyms, revalida-
tions, new statuses etc.) are introduced. All species are listed with
detailed localities and references. The book is addressed to spe-
cialists in coleopterology and entomology, biogeographers and
nature conservationists, collectors and all ground beetle lovers.
Without doubt, this excellent catalogue, compiled with dili-
gence and knowledge, will be an important tool for anyone,
not only specialists on Balkan Carabidae, and will improve our
insights into the systematics and distribution of carabid beetles.
The hope of the author to place Albania with this catalogue side
by side to Bulgaria (certainly the best-known regional fauna of
Carabidae in the Balkan Peninsula, catalogued in 1995 by the
same author and his father) is surely fulfilled. David Wrase
Pensoft Series Faunistica 64
” AnnotatedCatalogueoftheCarabidBeetlesofAlbania(Coleoptera:Carabidae)
“

2. Annotated Catalogue of the
Carabid Beetles of Albania
(Coleoptera: Carabidae)
Borislav V. Guéorguiev

3. 2 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
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4. SOFIA–MOSCOW
2007
Annotated Catalogue of the
Carabid Beetles of Albania
(Coleoptera: Carabidae)
BORISLAV V. GUÉORGUIEV

5. 4 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Pensoft Publishers
Geo Milev Str. 13a, Sofia 1111, Bulgaria
pensoft@mbox.infotel.bg
www.pensoft.net
ANNOTATED CATALOGUE OF THE CARABID
BEETLES OF ALBANIA (COLEOPTERA: CARABIDAE)
Borislav V. Guéorguiev
First published 2007
ISBN 978-954-642-295-8
Pensoft Series Faunistica No 64
ISSN 1312-0174
© PENSOFT Publishers
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or
transmitted in any form by any means, electronic, mechanical, photocopying, recording or
otherwise, without the prior written permission of the copyright owner.
Printed in Bulgaria, February 2007
Editor: Ivailo Stoyanov
Scientific readers: Augusto Vigna Taglianti & David W. Wrase

6. Contents
Introduction 7
Background of the study 7
Aims of the study 8
Short description of the studied area 8
Treatment of the taxa 10
Material and depositories of the collections 10
Acknowledgments 11
Abbreviations 13
Topographic treatment of some localities 14
Systematic part 16
Results 187
Conclusions 188
Taxonomic structure of the Albanian Carabidae (Table 1) 189
Appendix 193
References 205
Index 224

7. 6 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
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8. Introduction
The ground beetle fauna of Albania is not very well known taking into account the
fact that an overall investigation based upon studied material and critical survey on
the literature has never been made. So far, our knowledge of this fauna is still largely
due to Apfelbeck’s well-known monograph (Apfelbeck, 1904) and other works of the
same author published in the period 1901-1930 (Apfelbeck, 1901a; 1901b; 1902a;
1903; 1904; 1905; 1906; 1907a; 1907b; 1909; 1911; 1916; 1917; 1918a; 1918b; 1918c;
1919a; 1919b; 1930). Later, important faunistic contributions were made also by Csiki
(1940), Schatzmayr (1943b), and Hieke et al. (1986). Dealing with the Bulgarian ground
beetles, Hieke & Wrase (1988) mentioned 296 species (Nebriinae – 14, Cicindelinae –
5, Carabinae – 10, Omophroninae – 1, Brachininae – 10, Siagoninae – 1, Scaritinae –
12, Trechinae – 58, Harpalinae – 185) for Albania without detailed records. For polit-
ical reasons, in the period 1945-1990 not many faunistic records were published, and
relatively few materials from the country were available in the collections. Since 1990
collecting ground beetles there became possible again. Recently, 363 species (Nebrii-
nae – 10, Cicindelinae – 8, Carabinae – 18, Omophroninae – 1, Brachininae – 11,
Siagoninae – 1, Scaritinae – 20, Apotominae – 1, Trechinae – 55, Harpalinae – 238)
have been mentioned for Albania in the most recent catalogue of Palaearctic Co-
leoptera by Löbl & Smetana (2003).
Examination of the available collections added information about 55 new species
for the country plus 119 species omitted in the last work as well as many new faunistic
data for other species. The subsequent critical survey of the literature data brought to
the conclusion that 543 species of Carabidae live in Albania. Moreover, some taxo-
nomic changes and a lot of comments for many taxa have been made.
Background of the study
My interest in the Albanian Carabidae has appeared under the stimulus of my late
father, the well-known Bulgarian coleopterologist Vassil Borissov Guéorguiev (*1935
– †1996), immediately after the printing of the catalogue of the Bulgarian Carabidae
(Guéorguiev & Guéorguiev, 1995). The insufficient level of exploration of the cara-
bid fauna of the country at this time (because of the strong and prolonged political
isolation and lack of regional specialists), along with relatively diverse and fresh mate-
rial collected by Bulgarian zoologists in Albania, were the principal reasons for start-
ing this investigation. The complicated orography of this country, particularly the
complex relief, determines a great diversity of zonal (seacoast, lowland, hilly, forest,
subalpine, alpine) and intrazonal (riparian, stony, limestone) habitats, that could favor
INTRODUCTION 7

9. 8 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
a high diversity of the focused animal groups – the ground beetles. Finally, it was
necessary to sort out some interpretation of toponyms in the region, which were
incorrectly treated in the past (for example, some authors have erroneously cited
records or have referenced old collection material as being from Albania).
Aims of the study
The present study pursues two objectives, and the first one is the accomplishment of
faunistic inventory of the Carabidae from Albania. This process includes discovery
of all faunistic publications for Albania and identification or partial revision of the
collections (for unpublished and published specimens, respectively) and their regis-
tration. The result of these activities is the improvement of the faunistic knowledge
(species diversity and distribution) about the ground beetles not only on the territory
of Albania, but also on the Balkan Peninsula, and in Europe on a large scale. The
second objective of the study is taxonomical – examination of type material and
revision of the status of some doubtful taxa from Albania and its adjacent regions
(Montenegro, the Kosovo Region of Serbia, Macedonia and Northwestern Greece),
which will contribute to the systematics of the group.
Short description of the studied area
Albania (indigenous name Shqipëri) is situated in the southwestern part of the Bal-
kan Peninsula and is meridionally 300-400 km long. The country occupies about 28,748
km2
, and its maximal width is about 150 km. Albania is bordered in northwest with
Montenegro, in northeast and east with the Kosovo Region of Serbia and Mace-
donia, and in south with Greece. In the west, Ionic Sea divides Albania from the
Apennine Peninsula where the strait Otranto is the close water area between both
lands. One third of the country, mainly located in the western part, lies lower than
300 m a.s.l., the territory at altitude between 300 and 1000 m is almost 40 % of the
country, while that above 1000 m a.s.l. is about 30 %. The mountains situated in the
east of Albania are part of the Dinaric mountainous system and they represent rela-
tively young and high massifs. The highest top is Korabi Peak (2764 m) located on the
east state border. The main Albanian rivers, Drin, Shkumbin, Semani, Vijosë, flow in
direction northwest - southeast.
The situation of the country in the Mediterranean area and in a zone of subtropical
climate determines the principal features of the flora and fauna. The summer is dry and
warm with average temperatures in July between 24 and 25 C°, the autumn and winter
are mild and rainy. In Albania falls one of the most abundant rainfalls in Europe –
between 1000 and 1800 mm per year in the western lowlands, and to 2500 mm in the
mountains. The seasonal distribution of the precipitations is very uneven – in the sum-

10. Map of Albania
MAP OF ALBANIA 9

11. 10 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
mer the rainfall is just about 1/10 from the annual sum, and most plains and hilly parts
of the country are characterized with heavy summer droughts. The days with average
temperature more than 10 C° are about 300 (in the seacoast plains) and favor the devel-
opment of Mediterranean flora and fauna. The evergreen sclerophilous Mediterranean
forests are developed in the western part of Albania up to altitude of 400-500 m and in
some other regions – higher. These forests consist of maquises (mosaically distributed
in the country) where Olea sp., Quercus coccifera, Q. ilex, other Quercus spp., Laurus nobilis,
Phyllirea angustifolia, Pistacia lentiskus, Juniperus oxicedrus and other dominate. The following
higher zone is occupied by xeric oak-hornbeam forests. The two zones are ones mostly
suffering from human activities – in many places those forests are destroyed and, in are
replaced by farmlands or secondary xeric formations including mostly Paliurus spina-
cristi. The belt of the mesophilous oak-hornbeam and beech forests is developed at
altitude from 900-1000 up to 1600-1800 m a.s.l. This vegetation is more abundant in the
northern parts of Albania, where the mesophilous forests form the upper forest bor-
der. Normally in Northern Albania the coniferous communities are above the beech
forests, but elsewhere inversions in the distribution of the deciduous and coniferous
forests, owing to local humidity conditions, may occur. The orophytic (subalpine and
alpine) grass belt starts from 1900-2000 m a.s.l. It may be noted that the intrazonal
riverside communities consist mainly of Platanus orientalis.
Treatment of taxa
The arrangement of the taxa followed the order used in the recent Palaearctic Cata-
logue (Löbl & Smetana, 2003), with only two exceptions: genus Sinechostictus (Bembi-
diini) is regarded as separate genus, according to the recent studies on the phylogeny
of Trechitae (Grebennikov, 1997; Grebennikov & Maddison, 2005), and the arrange-
ment of Tachyini follows Sciaky & Vigna Taglianti (2003). Synonyms are included if
they consider taxa from Albania.
Material and depositories of the collections
So far, faunal records for Albania were found in 279 publications, not all of them
including original or detailed data. The main part of the studied materials are kept in
three large Central European museums, which preserve the most diverse collections of
ground beetle species from the Balkan Peninsula, as well as type material from the
region. Common feature of these collections is that the principal part of the Albanian
material had been collected before 1942. Both the most diverse and most numerous
materials collected from the country are these in the Museum für Naturkunde der Hum-
boldt Universität in Berlin. There, the collection of A. Bischoff is the most outstanding
one; other notable collections are these of E. v. Oertzen and H. Hopp, V. Apfelbeck

12. and A. Winneguth, A. Penther and H. Zerny (most part), H. Priesner (most part), L.
Mader, C. Lona, J. Müller. F. Hieke, G. Müller-Motzfeld and L. Behne (Hieke et al.,
1986), as well as the materials collected in the second half of XX century by H. Peks
and S. Peks. The second important depository is that in the Naturhistorisches Museum
Wien, Vienna, Austria. From aspect of the species diversity the materials from Albania
there are comparable with these in Berlin. The most significant ground beetle collec-
tions in Vienna are these of V. Apfelbeck and his laboratory assistant A. Winneguth, E.
Csiki, A. Penther and H. Zerny, H. Priesner, L. Mader (most part), A. Bischoff and A.
Winkler (most part). Additionally, small parts of the collections of Matzenauer, C. Ravas-
ini and C. Lona, J. Müller, L. Weirather and J. Meschnigg are also housed in Vienna.
Another place of important carabid collection is the Hungarian Natural History
Museum, Budapest. The most important collections here are these of V. Apfelbeck
and A. Winneguth, E. Csiki (most part) and Xh. Murraj. Other small collections housed
in Budapest are those of M. Grabowski, L. Mader, C. Ravasini and C. Lona, A. Win-
kler and A. Bischoff. Recently, Fehér et al. (2004) also collected material of Carabidae
from Albania that is currently in process of preparation.
An important place as depository of type material from the Balkans is the collec-
tion of the Deutsches Entomologisches Institut in Müncheberg, Germany; some of
the types deposited there have been reviewed. Materials from one American and two
Balkan museums have been examined too. The most substantial collection among
three is that housed in the National Museum of Natural History in Sofia. It has been
collected by Bulgarian expeditions to Albania after 1992 and at present it is contain-
ing about 160 species. The other two collections are: i/ the one in the Museum of
Natural Sciences, Tiranë (Albania), which in part was identified and published by
Murraj (1968), and ii/ that in the Carnegie Museum of Natural History, Pittsburgh
(USA), consisting of recently collected material of 57 species.1
Acknowledgments
The most important part of the material was examined in the course of the author’s
work under the European Union-funded Integrated Infrastructure Initiative “Syn-
thesys”, which is actually the prime cause for the completion of this work (Applica-
tions DE-TAF-725, AT-TAF-758 and HU-TAF-817). For the work with the collec-
1
Except the aforementioned collections, there are other four European museums that may house more
or less large number of Carabidae from Albania. At the present time a significant part of the former
collection of V. Apfelbeck is preserving in the Zemaljski muzej Bosne i Hercegovine in Sarajevo
(Bosnia Herzegovina). The Museo Civico di Storia Naturale in Trieste (Italy) and the Museo Civico di
Storia Naturale in Milano (Italy) house the most part of the materials collected by the Italian expeditions
in Albania before the Second World War. The first one is remarkable with the collections of C. Ravasini,
C. Lona and J. Müller, and the second one - with that of A. Schatzmayr. Most probably, the collection
of the Narodni Museum in Prague (Czech Republic) also contains materials from Albania.
ACKNOWLEDGEMENTS 11

13. 12 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
tions and the materials on loan I am indebted to Prof. V. Andoni (MNST), Dr. P.
Beron (NMNHS), Dr. R. Booth (BMNH), Mr. R. Davidson (CMNH), Mr. B. Jaeger
(ZMHU), Dr. A. Popov (NMNHS), Dr. H. Schönmann (NMW), Dr. Gy. Szel
(HNHM), Dr. M. Uglih (ZMHU) and Dr. L. Zerche (DEI).
With pleasure, I record my deep appreciation to the colleagues who contributed
this study giving their unpublished data (marked with the paragraphs “Material in-
cluded” or “Additional material included”): in the first place Prof. Dr. A. Vigna Taglianti
(Università di Roma”La Sapienza”, Roma, Italy) as well as Mr. P. Bulirsch (Prague,
Czech Republic), Dr. F. Hieke (ZMHU), Mr. S. Hristovski (Institute of Biology, Sko-
pje, Republic of Macedonia), Mr. B. Jaeger (ZMHU), Prof. Xh. Murraj (Tiranë, Alba-
nia) and Mr. D.W. Wrase (Berlin, Germany). It is worth mentioning that, Prof. Dr. A.
Vigna Taglianti told me that several years ago he began the preparation of a checklist
of the Carabidae of Albania, being still not completed. Generously, he entrusted to
me the draft of this work which included a moderate number of records collected
from the Italian expeditions to Albania carried out during the last 15 years. Dr. F.
Hieke made valuable comments and improvements in the part about genus Amara, as
well as kindly provided me the data of both unavailable papers and papers in press.
The following colleagues also revised and improved different parts of the catalogue:
Dr. P. Bonavita (Bembidiini; Università di Roma”La Sapienza”, Roma, Italy), Mr. P.
Bulirsch (Scaritinae), Mr. J. Gebert (Cicindelinae; Rohne, Germany), Dr. P.M. Giachi-
no (Speluncarius, Tapinopterus; Settore Fitosanitario Regionale, Torino, Italy), Mr. B.
Jaeger (Stenolophina), Mr. P. Neri (Bembidiini; Forlí, Italy), Dr. B. Kataev (Harpalini;
ZISP), Mr. L. Picciau (Speluncarius, Tapinopterus; Museo Regionale de Scienze Naturali,
Torino), Dr. L. Toledano (Bembidiini; Museo Civico di Storia Naturale, Verona, Ita-
ly), Prof. Dr. A. Vigna Taglianti (whole draft, and especially Bembidiini) and Mr. D.W.
Wrase (whole draft, and especially Harpalini).
For the determination of material, I thank Dr. D. Fedorenko (Dyschirius; Moscow
Pedagogical State University, Moscow, Russia), Dr. F. Hieke (Amara), Dr. B. Kataev
(Anisodactylus, Harpalus, Ophonus, Trichotichnus, Acupalpus, Bradycellus), both Mr. G. Ledoux
and Mr. Ph. Roux (Nebria; Paris, France), Mr. P. Neri (Bembidion), Mr. M. Pavesi (Syn-
tomus, Olisthopus; MSNM, Italy) and Mr. D.W. Wrase (Notiophilus, Harpalus, Ophonus,
Parophonus, Microlestes).
Special thanks are due to Prof. Dr. A. Vigna Taglianti, Mr. D.W. Wrase, and Mr. I.
Stoyanov (Central Laboratory of General Ecology, Sofia, Bulgaria), who made care-
ful reviews and proposed important changes of the earlier versions of the manu-
script on which this paper is based. Willingly, again Mr. I. Stoyanov helped me with
the revision of the English text. Prof. Dr. A. Casale (Università di Sassari, Sassari)
kindly allowed me the reproduction of his drawing of Abanotrechus beroni, and Dr. P.
Stoev (NMNHS) gave me the basis of the map.
I owe deep gratitude to my late farther for his strong and enthusiastic support in the
initial stages of the work, as well as to my mother and my wife for their tolerance.

14. Abbreviations
I. Abbreviations of collections studied upon the European Union-funded In-
tegrated Infrastructure Initiative “Synthesys” by the author
DEI Deutsches Entomologisches Institut, Müncheberg, Germany
HNHM Hungarian Natural History Museum (Magyar Természettudományi Múzeum),
Budapest, Hungary
MNHUB Museum für Naturkunde der Humboldt Universität zu Berlin, Bereich Zoo-
logisches Museum, Berlin, Germany
NMW Naturhistorisches Museum Wien, Vienna, Austria
II. Abbrevuiations of other collections studied by the author
BMNH The Natural History Museum, London, United Kingdom
CMNH Carnegie Museum of Natural History, Pittsburgh, USA
MNST Museum of Natural Sciences, Tirana, Albania
NMNHS National Museum of Natural History, Sofia, Bulgaria
III. Abbreviations of collections studied by other colleagues
MCG Museo Civico di Storia Naturale “Giacomo Doria”, Genoa, Italy
MHNG Muséum d’Histoire Naturelle, Geneva, Switzerland
MNHP Muséum National d’Histoire Naturelle, Paris, France
MSNM Museo Civico di Storia Naturale di Milano, Milano, Italy
MTKD Museum für Tierkunde, Dresden, Germany
NMPC National Museum Prague (Národni muzeum v Praze), Prague, Czech Re-
public
ZISP Zoological Institute, St. Petersburg, Russia
ZSM Zoologische Staatssammlung, Munich, Germany
cAVT private collection A. Vigna Taglianti, Roma, Italy
cBJ private collection B. Jaeger, Berlin, Germany
cFC private collection F. Cassola, Rome, Italy
cLR private collection G. Ledoux & P. Roux, Paris, France
cPB private collection P. Bulirsch, Prague, Czech Republic
cPN private collection P. Neri, Forlí, Italy
cSH private collection S. Hristovski, Skopje, Macedonia
cWH private collection W. Heinz, Schwanfeld, Germany
cWR private collection D.W. Wrase, Berlin, Germany
cXM private collection Xh. Murraj, Tiranë, Albania
ABBREVIATIONS 13

15. 14 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
IV. Other abbreviations and symbols used in the systematic part of the catalogue
! taxa still not well defined
? taxa with doubtful or questionable occurrence in Albania
[ ] taxa that most probably inhabit Albania; they were recorded from neighbouring
region
# taxa incorrectly indicated for Albania, and therefore excluded from the catalogue
f. female specimen/s
m. male specimens/s
s. specimen/s
Topographic treatment of some localities
(first are the correct treatments of toponyms, then their most frequent variants in the
examined references and collections)
Buenë River = Bojana River
Çukall Mt. = Cukali; Cucale
Dukati = Fush-Dukati; Fushes Dukati; Dukat-Fshat
Durrës = Durazzo
Gjalicë Mt. or Gjalicë e Lumes = Montes Gjallica Ljums; Djalica Ljums
Gjirokastër = Argirokastro
Gostilë = Köstil
Ibë (? Ibë e Postmë; ? Ibë e Sipërme) = Iba
Karbunara e Madhe = Karbunara
Kukës = Kukësi
Lezhë = Alessio
Liqen i Shkodrës = Shkodër Lake
Llogora = Logora; Lungara
Lura Mt. = Kunora e Lurës
Lurë e Vjetër = Lurja
Lushnje = Lushnia
Maja e Rrabës = “Raba Massiv”
Mal i Dajtit Mt. = Dajti Mt.; Dajt Massif
Mal i Shenjt Mt. = Mal i Shêit
Maranajt Mt. = Maranaj
Nangë = Nangat
North Albanian Alps = Prokletija
Peshtrik Mt. = Paštrik; Pashtrik
Rapshë = Rapša; Rapsa
Shengjin = San Giovanni di Medua; Medua
Shëndelli Mt. = Shendeli; Shen Ili

16. Shkëlzen Mt. = Shkölsen; Skölsen
Shkodër = Scutari
Stan-Karbunarë = Star Karbunara
Tejas = Radomir (village in East Albania, Korab Mt.)
Tepelenë = Tepelene
Tërpan = Terpan
Theth = Thethi
Vermosh = Vermosa
Vijosë River = Vijosa River
Vlorë = Valona; Avlona
Zepë Mt. = Zebia; Zebja
TOPOGRAPHIC TREATMENT OF SOME LOCALITIESEDITORIAL 15

17. 16 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Family CARABIDAE Latreille, 1802
Subfamily NEBRIINAE Laporte, 1834
Tribe NEBRIINI Laporte, 1834
1. Genus Eurynebria Ganglbauer, 1891
Notes. New genus for the fauna of Albania.
1.1. Eurynebria complanata (Linnaeus, 1767)
Materialstudied.“Apfelb.Albanien”,4s.(MNHUB);“Albania:Valona.”/“complanataorientalisTypus!
Apfb.”, 1 s. (HNHM); “Valona. Albania” / “Typus! Apf.”, 1 s. (HNHM); “Valona” / “v. albanica
Cotypus! Apfb”, 4 s. (HNHM); “Südalbanien Kopstein” / “Eurynebria complanata var. albanica Apf.”, 2
s. (NMW); “Alban. Wght. Valona” / “Eurynebria complanata var. albanica Apf.”, 8 s. (NMW); “Alban.
Wght. Valona”, 4 s. (MNHUB); “Albanien Avlona 08” / “E. complanata albanica Coll. Apfelbeck”, 4 s.
(HNHM); “Valona Albanien V.1908 H. Hopp”, 2 s. (MNHUB); “Ruskuli Alb., Mader” / “Eurynebria
complanata var. albanica Apf.”, 2 s. (NMW); “Shëng… 25.X.1965 kol. Murraj”, 2 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Shën Pjeter, Durrës District,
29.V.2004, 1 f., A. Paparistos leg. (cAVT).
Notes. Eurynebria complanata orientalis Apfelbeck, E. complanata albanica and E. complanata var.
albanica Apfelbeck are nomen nudum. The species was recorded hitherto only for European
Turkey on the Balkans (Farkač & Janata, 2003: 78). New species for the fauna of Albania.
2. Genus Leistus Frölich, 1799
Subgenus Leistus s.str.
2.1. Leistus fulvibarbis Dejean, 1826
2.1.1. Leistus fulvibarbis fulvibarbis Dejean, 1826
References. Leistus fulvibarbis: Apfelbeck, 1904: 50 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);
Schatzmayr, 1943b: 94 (Stan-Karbunarë); Perrault, 1986: 58, fig. 3; Hieke & Wrase, 1988: 21;
Perrault, 1991: 198, fig. 22.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Skutari Bojana”, 6 s. (NMW;
MNHUB); “Apfelb. Alban. spt. IV.-V.’05”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Logara
Juni”, 1 s. (MNHUB); “leg. Bischoff Albania 1934” / “Logara”, 1 s. (MNHUB); “Shen Ili” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”,
1 s. (MNHUB); Tiranë Park, 1-9.IX.1997, 7 s., R. Ward leg. (CMNH).
Notes. Farkač & Janata (2003: 82) have omitted L. fulvibarbis for Albania. Here, the species is
confirmed for the country.
SYSTEMATIC PART

18. COLEOPTERA: CARABIDAE: NEBRIINAE 17
2.2. Leistus nitidus (Duftschmid, 1812)
References. Leistus nitidus: Perrault, 1986: 58, fig. 3; Perrault, 1991: 198, fig. 23.
Material studied. “Albania leg. Bischoff 1936” / “Shkëlzen”, 1 s. (MNHUB).
Notes. Perrault displayed the species range as including North Albania (Perrault, 1986;
Perrault, 1991). In the second work he noted also a single locality on the map. First detailed
record for Albania.
Subgenus Pogonophorus Latreille, 1802
2.3. Leistus apfelbecki Ganglbauer, 1891
Material included (studied A. Vigna Taglianti). Valbona, Lugu i Cirkut Mt., 2000 m,
16.VII.1993, 1 m., L. Latella leg. (cAVT).
Notes. New species for the fauna of Albania.
2.4. Leistus magnicollis Motschulsky, 1866
References. Leistus magnicollis: Reitter, 1884: 36; Apfelbeck, 1904: 50; Apfelbeck, 1907a: 498,
501 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 21; Perrault, 1992: 21, fig. 11.
Material studied. “Oroši (Ptr.) Alban. spt.”, 2 s. (NMW); “Merdita Oroschi”, 2 s. (HNHM);
“Alban. Exp.’18 Pashtrik 4-14.7.”, 1 s. (NMW); “Albania leg. Bischoff 1935” / “Koritnik”, 1 s.
(MNHUB); “Albania leg. Bischoff 1935” / “Llogora”, 3 s. (MNHUB); “Giallica” / “Albania
leg. Bischoff 1936”, 9 s. (MNHUB); “Albania leg. Bischoff 1936” / “Shkalla Haruse” / “Korab”,
1 s. (MNHUB); “Cafa e Sharit Iballe” / “Albania leg. Mueller 1939”, 7 s. (MNHUB); “Maja e
Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë,
1000 m, 9.V.1995, 2 s., L. Penev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 1
s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).
Notes. Farkač & Janata (2003: 83) have omitted L. magnicollis for Albania. Here the species is
confirmed for the country.
2.5. Leistus parvicollis Chaudoir, 1869
References. Leistus parvicollis: Perrault, 1992: 18-19, 22, fig. 12; Farkač & Janata, 2003: 83.
Material studied. “Oroši (Ptr.) Alban. spt.”, 1 s. (NMW); “Alban. spt. Maranai”, 1 s. (HNHM);
“Alban. Exped. Kula Ljums 15-20.VIII.’18”, 1 s. (NMW); “Albania leg. Bischoff 1935” /
“Cukale”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB); “Bashtrik”
/ “Albania leg. Bischoff 1936”, 7 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 7 s.
(MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania leg. Bischoff
1936” / “Mal i Krujes”, 2 s. (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Mal i
Dajtit”, 1 s. (MNHUB); “Mali i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB);
Gjirokastër, 6.V.1994, 1 s., P. Stoev leg. (NMNHS).
Notes. For the present, the differentiation of L. parvicollis from L. spinibarbis s.l. bases only on
the ratio of the maximal width of head and the width of pronotum base (Perrault, 1992: 19).
First detailed records for Albania.

19. 18 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
2.6. Leistus rufomarginatus (Duftschmid, 1812)
References. Leistus rufomarginatus: Apfelbeck, 1907a: 498 (Oroshi); Hieke & Wrase, 1988: 21;
Farkač & Janata, 2003: 84.
Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.-V.’05”, 5 s. (NMW); “Albania
leg. Bischoff 1932” / “Mal i Dajit”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “Tomor-
Buranj Juli 1932”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB).
2.7. Leistus spinibarbis (Fabricius, 1775)
2.7.1. Leistus spinibarbis rufipes Chaudoir, 1843 [= punctatus Reitter, 1885]
References. Leistus spinibarbis rufipes: Apfelbeck, 1904: 49 (Vlorë; Berat); Perrault, 1992: 18, 22
(fig. 12); Farkač & Janata, 2003: 84. Leistus spinibarbis: Anonymous, 1907: 279; Hieke & Wrase,
1988: 21. Leistus spinibarbis punctatus: Apfelbeck, 1909: 297 (Vlorë).
Material studied. “Latif Skutari”, 2 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt. IV.-V.’05”,
12 s. (NMW); “Merdita Munela”, 4 s. (NMW); “Alban. spt. Maranai”, 9 s. (HNHM; MNHUB);
“Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 9 s. (NMW); “Albania leg. Bischoff 1932” / “Ige-
Ezren”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “Kjori”, 2 s. (MNHUB); “leg. Bischoff
Albania 1934” / “Mal i Krues”, 10 s. (MNHUB); “Giallica” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 24 s. (MNHUB); “Mal i Dajit” / “Albania
leg. Bischoff 1936”, 11 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 23 s. (MNHUB);
“Maja e Policanit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “23.IV.1958
leg. Xh. Murraj”, 10 s. (HNHM); Mal i Dajtit Mt., 10-11.V.1965, 4 s. (MNST).
Notes. Except the material studied above I have seen more specimens in MNHUB, which
remained unregistered.
3. Genus Nebria Latreille, 1802
Subgenus Alpaeus Bonelli, 1810
Notes. The systematic of the Balkan species from this group is complex and still not well
settled. Comparison carried out between specimens from all the subspecies of Nebria aetolica
Apfelbeck, 1901 s.l. in HNHM revealed that some of them are more closely related each other
than to others. The subspecies have vicarious distribution in the alpine zones of the high
mountains of South Albania, Southwestern Macedonia, North and Central Greece. Details in
the pronotum form manifested that N. aetolica galicica, N. aetolica macedonica, N. aetolica papingutica
and N. aetolica rambouseki are more similar each other than any of them to the remaining
subspecies. For the reason to display more natural relations in this group the following
nomenclatorial acts are proposed:
Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. of Nebria (Alpaeus) aetolica macedonica Mařan, 1938
Nebria (Alpaeus) macedonica galicica Horvatovich, 1974, new assessment of Nebria (Alpaeus)
aetolica galicica Horvatovich, 1974
Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974, new assessment of Nebria (Alpaeus)
aetolica papingutica Horvatovich, 1974
Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assessment of Nebria (Alpaeus) aetolica
rambouseki Mařan, 1938

20. COLEOPTERA: CARABIDAE: NEBRIINAE 19
3.1. Nebria attemsi Apfelbeck, 1908
References. Nebria attemsi: Bänninger, 1925: 272; Mařan, 1938: 51 (Korab Mt.); Csiki, 1940:
210 (Korab Mt.); Ledoux & Roux, 1992: 258; Farkač & Janata, 2003: 85.
Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 40 s. (HNHM); “Akad. Balk.
Exp. Csiki 1918” / “Albania Mts. Korab”, more than 70 s. (HNHM); “Alban. Exped. Korab, 1918,
24-31.VII.”, more than 80 s. (NMW); “Albania leg. Bischoff 1932” / “Tomor”, 1 s. (MNHUB);
“Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” /
“Derzë”, 2 s. (MNHUB); “Albania leg. Bischoff 1935” / “Korab”, more than 50 s. (MNHUB).
Notes. Apfelbeck published three descriptions of this species (Apfelbeck, 1908; Apfelbeck,
1917; Apfelbeck, 1930), the oldest one is correctly treated as valid.
3.2. Nebria bosnica Ganglbauer, 1889
3.2.1. Nebria bosnica jezercensis Horvatovich, 1974
References. Nebria bosnica jezercensis: Horvatovich, 1974b: 90-91 (Maja e Jezercë Mt., loc. typ.);
Huber, 2001: 78 (Maja e Jezercë Mt.); Farkač & Janata, 2003: 85 (Maja e Jezercë Mt.).
3.2.2. Nebria bosnica kulti Mařan, 1939
References. Nebria apfelbecki kulti: Mařan, 1939a: 20-21 (Shkëlzen Mt., loc. typ.). Nebria bosnica:
Csiki, 1940: 210 (Shkëlzen Mt.). Nebria bosnica kulti: Horvatovich, 1974b: 90 (Shkëlzen Mt.);
Huber, 2001: 78 (Shkëlzen Mt.); Farkač & Janata, 2003: 85 (Shkëlzen Mt.).
Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 5 s. (HNHM); “Shkëlzën”
/ “Albania leg. Bischoff 1936”, 1 syntype (HNHM); “Albania leg. Bischoff 1936” / “Shkëlzën”,
15 s. (MNHUB).
3.2.3. Nebria bosnica prokletiensis Horvatovich, 1974
References. Nebria apfelbecki: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near
Vermosh); Mařan, 1939a: 20 (North Albanian Alps: Grebeni near Vermosh). Nebria bosnica
prokletiensis: Horvatovich, 1974b: 91 (North Albanian Alps, loc. typ., state attribution
unspecified); Farkač & Janata, 2003: 85.
Material studied. “Prokletija Gbt. Dr. Penther”, 1 paratype (HNHM).
Notes. N. bosnica apfelbecki Ganglbauer, 1891 inhabits only mountains in Eastern Bosnia
Herzegovina and Western Montenegro.
3.2.4. Nebria bosnica sturanyi Apfelbeck, 1906
References. Nebria sturanyi: Apfelbeck, 1906: 1662-1663 (Maranajt Mt., loc. typ.); Apfelbeck,
1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1918c: 160 (Maranajt Mt.);
Bänninger, 1925: 270; Mařan, 1939a: 20 (Maranajt Mt.); Pretner, 1978: 146, 166. Nebria bosnica
sturanyi: Horvatovich, 1974b: 89 (Maranajt Mt.); Huber, 2001: 77-78 (Maranajt Mt.); Farkač &
Janata, 2003: 85 (Maranajt Mt.).
Material studied. “Latif Alban spt.”, 1 s. (HNHM); “Alban. spt. Maranai”, 4 s. (MNHUB);
“Maranai Alban.-occ.” / “Coll. Apfelbeck”, 1 s. (HNHM); “Maranai G.” / “Apfelb. Alban. spt.
11.V.05”, 3 s. (HNHM); “Apfelb. Alban. spt. IV.-V.05” / “Maranai” / “sturanyi Apfelb.”, 15 s.
(NMW); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 6 s., P. Beron & B.
Petrov leg. (cLR; NMNHS).

21. 20 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
? Nebria bosnica tresnjevikensis Horvatovich, 1974
Notes. This geographical race inhabits the Montenegrin part of the North Albanian Alps and
may live in the northernmost alpine spot in Albania too. Huber (2001: 81, Abb. 3) showed
map where the subspecies is shown to live in the country in spite of lack of exact locality.
? Nebria dahli (Duftschmid, 1812)
? Nebria dahli montenegrina Apfelbeck, 1904
References. Nebria dahli velebitica: Apfelbeck, 1904: 56 (“Sv. Troice bei Prisren”). Nebria dahli:
Horion, 1941: 81.
Notes. The locality “Sv. Troice bei Prisren” (Apfelbeck, 1904) is situated on the northern slopes of the
Shar Planina Mountains, in the Kosovo Region of Serbia. Probably Horion (1941) followed the same
record. N. dahli may live in Albania since it occurs in the massif of Durmitor (Apfelbeck, 1904 sub N.
dahli montenegrina), and N. dahli velebitica Heyden, 1884 inhabits only in Croatia (Farkač& Janata, 2003: 85).
3.3. Nebria ganglbaueri Apfelbeck, 1905
3.3.1. Nebria ganglbaueri ganglbaueri Apfelbeck, 1905
References. Nebria ganglbaueri: Apfelbeck, 1905: 240 (Albania / Kosovo border: Koritnik Mt.,
alpine belt, loc. typ.); Apfelbeck, 1907b: 636 (Koritnik Mt., alpine belt, loc. typ.); Bänninger,
1925: 270. Nebria ganglbaueri ganglbaueri: Horvatovich, 1975: 272 (Koritnik Mt.); Farkač & Janata,
2003: 86 (Koritnik Mt.).
Material studied. “Latif Koritnik” / “ganglbaueri Apfelb.”, 2 syntypes (NMW).
Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b),
the older one is correctly treated as valid.
3.3.2. Nebria ganglbaueri gyalicaensis Horvatovich, 1975
References. Nebria ganglbaueri: Csiki, 1940: 210 (Gjalicë Mt.). Nebria ganglbaueri gyalicaensis:
Horvatovich, 1975: 275-276 (Gjalicë Mt., loc. typ.); Farkač & Janata, 2003: 86 (Gjalicë Mt.).
Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype
and 18 paratypes (HNHM); “Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 1 paratype (NMW);
“Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 16 s. (NMW).
3.3.3. Nebria ganglbaueri korabensis Horvatovich, 1975
References. Nebria ganglbaueri: Csiki, 1940: 210 (Korab Mt.). Nebria ganglbaueri korabensis:
Horvatovich, 1975: 274-275 (Korab Mt., loc. typ.); Farkač & Janata, 2003: 86.
Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype and 22
paratypes (HNHM); “Alban. Exped. Korab 1918 24-31.VII.”, 23 s. (NMW); “Albania leg.
Bischoff 1935” / “Korab”, more than 140 s., incl. 27 paratypes (MNHUB).
Notes.Farkač&Janata(2003:86)notedthissubspeciesforex-Yugoslavia(SerbiaandMontenegro).
In fact its type locality is the Korab Mt. lying on the border between Albania and Macedonia.
3.3.4. Nebria ganglbaueri merditana Apfelbeck, 1906
References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Munella Mt., loc. typ.);
Apfelbeck, 1907a: 500, 503 (Munella Mt.); Bänninger, 1925: 270; Mandl, 1964: 33-36;
Horvatovich, 1974a: 359. Nebria ganglbaueri merditana: Horvatovich, 1975: 272-273 (Munella
Mt.); Farkač & Janata, 2003: 86 (Munella Mt.).

22. COLEOPTERA: CARABIDAE: NEBRIINAE 21
Material studied. “Albania”, 1 s. (DEI); “Albania Merdita”, 1 s. designated incorrectly as
syntype of Nebria ganglbaueri ditana Apfelbeck (DEI); “Merdita Munela” / “Coll. Apfelbeck”,
5 s. (HNHM); “Merdita Munela”, 1 s. designated incorrectly as syntype of Nebria ganglbaueri
ditana Apfelbeck (DEI); “Merdita Munela”, 9 s. (MNHUB); “Merdita Munela” / “Vgth. 1906
Merdita”, 14 s. (NMW).
Notes. Apfelbeck, 1906 described this taxon from two closely situated mountains (Munella
Mt. and Zepë Mt.) situated in the area of Merdita. Also, he gave no exact “loc. typ.”. Later on,
Horvatovich (1975) attributed the specimens from Zepë Mt. to the ssp. zebiaensis, so that “loc.
typ.” of the ssp. merditana should be Munella Mt. Nebria ganglbaueri ditana Apfelbeck is either
nomen nudum or incorrect spelling of the name merditana. In the second case the two specimens
in DEI may be syntypes.
3.3.5. Nebria ganglbaueri zebiaensis Horvatovich, 1975
References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Zepë Mt.); Apfelbeck,
1907a: 500, 503 (Zepë Mt.). Nebria ganglbaueri zebiaensis: Horvatovich, 1975: 273-274 (Zepë
Mt., loc. typ.); Farkač & Janata, 2003: 86 (Zepë Mt.).
Material studied. “Merdita Zebia” / “Coll.Apfelbeck”, 1 holotype and 4 paratypes (HNHM);
“Merdita Zebia” / “Vgth. 1906 Merdita”, 2 s. (DEI).
Notes. Further study on the validity of this race (with respect to the previous one) is needed.
3.4. Nebria germarii Heer, 1837
! Nebria germarii ssp.
Material studied. “Albania leg. Bischoff 1935” / “Maja e Jezeroe” / “Nebria!! wahrscheinlich”
/ “N. germari ssp. n. det. Dr. Horvatovich”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania. For the present the subspecies status of the
local population is not defined, but it probably tends to ssp. durmitorensis Apfelbeck, 1904.
3.5. Nebria macedonica Mařan, 1938
3.5.1. Nebria macedonica galicica Horvatovich, 1974 [= korabica Horvatovich, 1974]
References. Nebria aetolica galicica: Horvatovich, 1974a: 360-361 (loc. typ. in Macedonian part
of Galičica Mt.); Ledoux & Roux, 1992: 257 (Korab Mt.); Farkač & Janata, 2003: 84. Nebria
aetolica korabica: Horvatovich, 1974a: 361 (loc. typ. in Macedonian part of Korab Mt.).
Material studied. “Korab”, 1 paratype of Nebria aetolica korabica (HNHM).
3.5.2. Nebria macedonica papingutica Horvatovich, 1974 [= tomoricensis Horvatovich, 1974]
References. Nebria aetolica tomoricensis: Mařan, 1945: 56 (Tomor Mt.); Horvatovich, 1974a:
364-365 (Tomor Mt., loc. typ. of tomoricensis; Tomor Mt.: Abbas Ali). Nebria aetolica papingutica:
Horvatovich, 1974a: 363-364 (Maja e Papingut Mt., loc. typ. of ssp. papingutica; Maja e Policanit
Mt.); Ledoux & Roux, 1992: 257 (Maja e Papingut Mt.; Tomor Mt.); Farkač & Janata, 2003: 84.
Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s., including 3
paratypes of Nebria aetolica tomoricensis (HNHM; NMW); “Tomor Monte Toik” / “Albania
1922 Ravasini, Lona”, 1 paratype of Nebria aetolica tomoricensis (HNHM); “Tomor: Tomor Alban.
mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 holotype and 4 paratypes of Nebria aetolica
tomoricensis (HNHM); “Tomori Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 5

23. 22 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
paratypes of Nebria aetolica tomoricensis (NMW); “Tomori Toik Alban. mer.” / “Winkler, Lona,
Bischoff, V.1931”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Tojk Tomor”, 10 paratypes of
Nebria aetolica tomoricensis (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e
Papingut”, 1 holotype and 26 paratypes of Nebria aetolica papingutica (HNHM; MNHUB);
“Albania leg. Bischoff 1937” / “Maja e Policanit”, 2 paratypes of Nebria aetolica papingutica
(HNHM); “Albania leg. Bischoff 1937” / “Mal i That”, 2 s. (HNHM).
[Nebria macedonica rambouseki Mařan, 1938]
Notes. This geographical race was described from the Macedonian part of the Jablanitsa Mt.
(Mařan, 1938: 57; Mařan, 1945: 56). So far there is no sure record from Albania.
Subgenus Boreonebria Jeannel, 1937
3.6. Nebria rufescens (Strøm, 1768)
3.6.1. Nebria rufescens rufescens (Strøm, 1768) [= gyllenhali (Schönherr, 1806)]
References. Nebria gyllenhali: Hieke & Wrase, 1988: 22. 12. Nebria rufescens rufescens: Farkač &
Janata, 2003: 88.
Material studied. “Albania leg. Bischoff 1935” / “Shkëlzen”, 4 s. (MNHUB).
Notes. First detailed record for Albania.
Subgenus Eunebria Jeannel, 1937
3.7. Nebria jockischi Sturm, 1815
3.7.1. Nebria jockischi jockischi Sturm, 1815
References. Nebria jockischi: Hieke & Wrase, 1988: 22.
Material studied. “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania leg.
Bischoff 1936” / “Korab”, 8 s. (MNHUB).
Notes. Farkač & Janata (2003: 90) have omitted N. jockischi for Albania. The species is here
confirmed for the country with first detailed records.
Subgenus Nebria s.str.
3.8. Nebria brevicollis (Fabricius, 1792)
References. Nebria brevicollis: Apfelbeck, 1904: 54 (Velipoja; Vlorë); Apfelbeck, 1909: 297
(Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes; Lushnje; Stan-Karbunarë; Berat); Hieke & Wrase,
1988: 22; Farkač & Janata, 2003: 92. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë).
Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s. (HNHM); “Albania
Sauk” / “2.V.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Dajti” / “2.6.1959 leg. Xh. Murraj”,
1 s. (HNHM); Tiranë, 10.V.1961, 1 s. (MNST); Mal i Dajtit Mt., 10-11.V.1965, 3 s. (MNST);
Tiranë Park, 8.III.1998, 54 s., shaded lake edge under leaves and wet, rotten wood, R. Ward
leg. (CMNH; NMNHS); “Albania Tomor Mts. Kajala e Tomorit, 26.V.2004, K. Harmos & D.
Muránya”, 1 s. (HNHM).

24. COLEOPTERA: CARABIDAE: NEBRIINAE 23
Additional material included (studied A. Vigna Taglianti). Tiranë, Mal i Dajtit Mt.,
7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley
of Vijosë River, 300-360 m, 18.IV.2003, 3 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT);
Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 m., 2 f., P. Crucitti & C.
Cavalier leg. (cAVT).
Notes. Except the material studied above I have seen additionally more than 20 s. from different
localities in MNHUB.
3.9. Nebria kratteri Dejean, 1831
3.9.1. Nebria kratteri kratteri Dejean, 1831 [= valonensis Apfelbeck, 1904]
References. Nebria kratteri: Apfelbeck, 1904: 52 (South Albania); Apfelbeck, 1907a: 498, 501,
503 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1928: 46; Holdhaus, 1956: 7 (Mal i
Dajtit Mt.); Magistretti, 1965: 79; Ledoux & Roux, 1990: 94. Nebria kratteri valonensis: Apfelbeck,
1904: 55 (Vlorë); Bänninger, 1925: 274; Mařan, 1939a: 19 (Vlorë); Jeanne, 1974: 66. Nebria
kratteri ssp.: Schatzmayr, 1943b: 94 (cave “Stere Gropa e Vasos” near Dukati). Nebria kratteri
kratteri: Farkač & Janata, 2003: 92.
Material studied. “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);
“Mal-i-Dajtit Tiranë” / “Albania Bischoff 1931”, 2 s. (DEI); “Tomor: Kulmak Albania mer.”
/ “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara juni” / “Albania Bischoff 1932”, 1
s. (DEI); “Tomor-Buranj juli 1932” / “Albania Bischoff 1932”, 1 s. (DEI).
Notes. The record “Chani Driskos bei Jannina” (Apfelbeck, 1904: 55) refers to Northwestern
Greece. The locality at Mal i Shenit Mt. (Apfelbeck, 1907a), representing the northernmost
known spot in the species range, needs verification. In MNHUB, I have seen more than 100
specimens from different localities, which remained unregistered.
# Nebria salina Fairmaire & Laboulbène, 1854
References. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë).
Notes. This Atlantic species does not inhabit the Balkan Peninsula, so that the record for it
referred to N. brevicollis.
Tribe NOTIOPHILINI Motschulsky, 1850
4. Genus Notiophilus Dumeril, 1806
4.1. Notiophilus aestuans Dejean, 1826 [= pusillus C.R. Waterhouse, 1833]
References. Notiophilus aestuans: Csiki, 1940: 212 (Gjalicë Mt.). Notiophilus pusillus: Hieke &
Wrase, 1988: 24.
Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania,
Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918
29.V.-4.VI.”, 2 s. (NMW); “Mal i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” /
“Albanialeg.Bischoff 1936”,2s.(MNHUB);“Shkëlzën”/“Albanialeg.Bischoff 1936”,4s.(MNHUB).
Notes. Bousquet & Barševskis (2003: 96) have omitted N. aestuans for Albania. Here it is confirmed
for the country. The record “Peristeri-Gebirge bei Jannina” (Apfelbeck, 1904: 61 sub N. pusillus)
refers to Greece.

25. 24 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
4.2. Notiophilus aquaticus (Linnaeus, 1758)
References. Notiophilus aquaticus: Hieke & Wrase, 1988: 24.
Material studied. “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Korab”
/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Notes. Bousquet & Barševskis (2003: 96) have omitted the species for Albania. Here it is
confirmed for the country with first detailed records.
4.3. Notiophilus biguttatus (Fabricius, 1779)
References. Notiophilus biguttatus: Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 25.
Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Albania leg. Bischoff 1935” / “Korab”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” / “Maja
e Jezercoë”, 3 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania
leg. Bischoff 1936” / “Shkëlzën”, 3 s. (MNHUB); “Albania leg. Mueller 1939” / “Kunora
Drunit Cucale”, 8 s. (MNHUB); “Albania leg. Mueller 1939” / “Maja e Shpatit Iballe”, 1 s.
(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov
leg. (NMNHS); Lushnje, Divjaka National Park, 10.V.1995, 1 s., S. Golovach, B. Petrov, P.
Stoev leg. (NMNHS).
Notes. Bousquet & Barševskis (2003: 96) have omitted the species for Albania. Here it is
confirmed for the country.
4.4. Notiophilus danieli Reitter, 1897
References. Notiophilus danieli: Apfelbeck, 1904: 64 (Vlorë); Ganglbauer, 1904: 648; Apfelbeck,
1909: 297 (Vlorë); Magistretti, 1965: 84; Hieke & Wrase, 1988: 25.
Material studied. Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is
confirmed for the country.
4.5. Notiophilus germinyi Fauvel, 1863 [= hypocrita Putzeys, 1866]
References. Notiophilus hypocrita: Hieke & Wrase, 1988: 24.
Material studied. “Alban. Exp.’ 18 Pashtrik 4-14.7.”, 3 s. (NMW); “Koritnik” / “Albania leg.
Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB);
“Gjallica” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).
Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is
confirmed for the country with first detailed records.
4.6. Notiophilus palustris (Duftschmid, 1812)
References. Notiophilus palustris: Csiki, 1940: 212 (Gjalicë Mt.).
Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM).
Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is
confirmed for the country.

26. COLEOPTERA: CARABIDAE: CICINDELINAE 25
4.7. Notiophilus rufipes Curtis, 1829
References. Notiophilus rufipes: Apfelbeck, 1904: 64 (Velipoja near Lezhë); Hieke & Wrase,
1988: 25.
Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomor-
Buranj Juli 1932” / “Albania Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936
leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania leg. Mueller
1939”, 2 s. (MNHUB); “Maja e Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB);
Tiranë Park, 1-9.IX.1997, 2 s., R. Ward leg. (CMNH).
Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is
confirmed for the country.
4.8. Notiophilus substriatus C.R. Waterhouse, 1833
References. Notiophilus substriatus: Csiki, 1940: 212 (Kula Ljums; foot of Gjalicë Mt.); Hieke
& Wrase, 1988: 25.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Logara Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Murrsi” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal
i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Tiranë Park, 22-31.X.1997, 1 s., R.
Ward leg. (CMNH); Tiranë Park, 1-9.IX.1997, 1 s., R. Ward leg. (CMNH); Tiranë Park, 12-
18.XII.1997, 1 s., R. Ward leg. (NMNHS).
Notes. Bousquet & Barševskis (2003: 98) have omitted the species for Albania. Here it is
confirmed for the country.
Subfamily CICINDELINAE Latreille, 1802
Tribe CICINDELINI Latreille, 1802
5. Genus Calomera Motschulsky, 1862
5.1. Calomera fischeri (M. F. Adams, 1817)
5.1.1. Calomera fischeri fischeri (M. F. Adams, 1817)
References. Cicindela fischeri: Apfelbeck, 1907a: 496, 501 (Kiri River near Mesi); Csiki, 1940:
209 (Kula Ljums); Hieke & Wrase, 1988: 11.
Material studied. “Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
Shëngjin, 16.V.1963, 1 m. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev
leg. (NMNHS).
Notes. Puchkov & Matalin (2003: 100) have omitted the species for Albania. Here it is
confirmed for the country.

27. 26 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
5.2. Calomera littoralis (Fabricius, 1787)
5.2.1. Calomera littoralis nemoralis (Olivier, 1790)
References. Cicindela lunulata var. nemoralis: Apfelbeck, 1894: 546. Cicindela lunulata: Murraj,
1968: 33 (Golem; Suhagora; Theth; Tomor Mt.). Lophyridia nemoralis nemoralis: Mandl, 1981:
150; 154. Calomera littoralis nemoralis: Puchkov & Matalin, 2003: 100.
Material studied. Golem, 19.V.1958, 2 m. (MNST); Golem, 24.III. 1961, 1 m., 1 f. (MNST);
Mal i Dajtit Mt.: Shkall Priskë, 30.VI.1960, 1 m. (MNST); Shëngjik, 20.VII.1961, 1 m. (MNST);
Shëngjik, 16.V.1963, 2 m., 2 f. (MNST); Laguna Patog, 18.IX.1993, 4 s., S. Beshkov leg.
(NMNHS); Velipoja, 19.IX.1993, 5 s., S. Beshkov leg. (NMNHS).
Notes. Part of the material cited by Murraj (1968) was found and revised. Calomera lunulata
(Fabricius, 1781) is a valid species with primarily North African range, in Europe it lives only
in Sicily (Puchkov & Matalin, 2003: 101).
[genus Cephalota Dokhtouroff, 1883]
[subgenus Taenidia Rivalier, 1950]
[Cephalota circumdata (Dejean, 1822)]
[Cephalota circumdata circumdata (Dejean, 1822)]
Notes. Most probably, the species inhabits the southern seacoast of Albania (see also Gebert,
1999: 30, Abb. 1.2.).
6. Genus Cicindela Linnaeus, 1758
Subgenus Cicindela s.str.
6.1. Cicindela campestris Linnaeus, 1758
6.1.2. Cicindela campestris olivieria Brullé, 1832
References. Cicindela campestris: Apfelbeck, 1904: 6 (Velipoja near Lezhë; Vlorë; Berat; Durrës);
Apfelbeck, 1907a: 496 (Kiri River near Mesi); Apfelbeck, 1909: 297 (Vlorë; Berat); Csiki, 1940:
209 (Gjalicë Mt.; Korab Mt.); Murraj, 1968: 33 (Tiranë; Lurë; Sauk; Pashaliman; Kulla e Lumës);
Hieke & Wrase, 1988: 9. Cicindela campestris pontica: Schatzmayr, 1943b: 93 (Karbunara e Madhe;
Lushnje; Tërpan; “Poggio Boschetto”). Cicindela campestris olivieria: Mandl, 1944: 7; 12; 13 (Tiranë;
Korab Mt.); Puchkov & Matalin, 2003: 104. Cicindela campestris campestris: Puchkov & Matalin,
2003: 103.
Material studied. Dukati, 7.V.1958, 1 m. (MNST); Golem, 19.V.1958, 1 f. (MNST); Golem,
8.VII.1965, 1 f. (MNST); Mali-me Gropa, 3.VII.1958, 1 f. (MNST); Tiranë, 4.V.1959, 1 m. (MNST);
Tiranë, 10.IV.1965, 1 f. (MNST); Sauk, 9.VII.1961, 1 f. (MNST); Sauk, 11.IV.1964, 2 m. (MNST);
Sauk, 30.10.1963, 1 f. (MNST); Lurë, 18.VII.1961, 1 m. (MNST); Ibë, 24.IV.1962, 1 f. (MNST);
Mal i Dajtit Mt., 10.V.1965, 3 f. (MNST); North Albanian Alps, Maj’e Redohimes, 2400-2550 m,
29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m,
1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m,
20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); Vlorë, 1.V.1994, 1 s., P. Stoev leg.
(NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS); North
Albanian Alps, near cave B 52, 1600 m, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS); Petrela 15
km NE Tiranë, 300 m, 9.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).

28. COLEOPTERA: CARABIDAE: CICINDELINAE 27
Additional material included (studied F. Cassola). Rapsh, 15.V.2002, 1 s., A. Paparistos leg.
(cFC); Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003,
1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cFC); Vasjar (Tepelenë District), 18.IV.2003, 1 m., P.
Crucitti & C. Cavalieri leg. (cFC).
Notes. By the lack of modern study, the intraspecific relations of C. campestris s.l. in the Balkan
Peninsula are obscure. For this reason all the data about it from Albania are referred to ssp.
olivieria provisionally. The ssp. pontica Fischer von Waldheim, 1828 inhabits only the Black Sea
Coast, so that the record for this race is referred to ssp. olivieria. Except the material studied
above I have also seen additional specimens from Gjallica Lums and the Korab Mt. in HNHM.
# Cicindela hybrida Linnaeus, 1758
References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke
& Wrase, 1988: 10.
Notes. I follow the view of Gebert (1995) and refer the literature records for C. hybrida to C.
monticola albanica.
6.2. Cicindela monticola Menetries, 1832
6.2.1. Cicindela monticola albanica Apfelbeck, 1909
References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke
& Wrase, 1988: 10. Cicindela hybrida albanica: Apfelbeck, 1909: 297 (Vlorë, loc. typ.; Berat, loc.
typ.); Apfelbeck, 1917: 219 (Kiri River near Mesi, loc. typ.; Merdita: Munella, loc. typ.; Vlorë,
loc. typ.); Apfelbeck, 1918a: 75 (Shkodër, loc. typ.; Orish, loc. typ.; Vlorë, loc. typ.); Apfelbeck,
1930: 161; Murraj, 1968: 33 (Golem; Suhagora; Tomor Mt.; Shëngjin); Cassola, 1973: 26. Cicindela
sahlbergi albanica: Gebert, 1995: 16 (Buenë River near Shkorër; Mesi; Vlorë; Merdita: Munella;
Berat). Cicindela monticola albanica: Puchkov & Matalin, 2003: 106.
Material studied. “Skutari Bojana” / “C. v. albanica Apfb. Coll. Apfelbeck”, 2 s. (HNHM);
“Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania coll. Lichtn.”,
1 s. (HNHM); Shkumbi - Kavajes, 7.V.1958, 1 f. (MNST); Golem, 24.III.1961, 2 m., 3 f.
(MNST); Divjake, 18.VI.1964, 1 f. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G.
Blagoev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 2 s., B. Guéorguiev leg. (NMNHS);
Leskovac - Barmachi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Apfelbeck published four consecutive descriptions of the ssp. albanica (Apfelbeck, 1909;
Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930: 161), the oldest one is correctly treated as valid.
6.3. Cicindela sylvatica Linnaeus, 1758
6.3.1. Cicindela sylvatica sylvatica Linnaeus, 1758
References. Cicindela sylvatica sylvatica: Puchkov & Matalin, 2003: 107.
6.4. Cicindela transversalis Dejean, 1822
6.4.1. Cicindela transversalis transversalis Dejean, 1822 [= riparia Dejean, 1822]
References. Cicindela hybrida riparia: Apfelbeck, 1907a: 496 (Kiri River near Mesi).
Notes. Puchkov & Matalin (2003: 108) have omitted the species for Albania, therefore its
occurrence in Albania needs further confirmation.

29. 28 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
7. Genus Cylindera Westood, 1831
Subgenus Cylindera s.str.
7.1. Cylindera germanica (Linnaeus, 1758)
7.1.1. Cylindera germanica germanica (Linnaeus, 1758)
References. Cicindela germanica: Hieke & Wrase, 1988: 10. Cylindera germanica germanica: Puchkov
& Matalin, 2003: 109.
7.1.2. Cylindera germanica muelleri (Magistretti, 1966)
References. Cylindera germanica muelleri: Puchkov & Matalin, 2003: 109.
Subgenus Eugrapha Rivalier, 1950
7.2. Cylindera arenaria (Füessly, 1775)
7.2.1. Cylindera arenaria viennensis (Schranck, 1781)
References. Cicindela arenaria: Hieke & Wrase, 1988: 11. Cylindera arenaria viennensis: Puchkov
& Matalin, 2003: 110.
7.3 Cylindera trisignata (Dejean, 1822)
7.3.1. Cylindera trisignata trisignata (Dejean, 1822)
References. Cicindela literata sic!: Murraj, 1968: 33 (Shëngjin). Cylindera trisignata: Cassola, 1973:
33, Fig. 2. Cylindera trisignata trisignata: Puchkov & Matalin, 2003: 111.
Material studied. Shëngjin, 16.V.1963, 4 m., 4 f. (MNST); Radona, Shkiela, 21.VI.1994, 8 s.,
B. Guéorguiev leg. (NMNHS).
Notes. Cicindela litterata Sulzer, 1776 is synonym of Cylindera arenaria. Revision of material
published by Murraj (1968) revealed that it is identical with C. trisignata.
8. Genus Myriochila Motschulsky, 1858
Notes. New genus for the fauna of Albania.
Subgenus Myriochila s.str.
Notes. New subgenus for the fauna of Albania.
8.1. Myriochila melancholica (Fabricius, 1798)
8.1. Myriochila melancholica melancholica (Fabricius, 1798)
Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 8-
15.VII.2006, 1 s., M. Komnenov leg. (cSH).
Notes. New species for the fauna of Albania.

30. COLEOPTERA: CARABIDAE: CARABINAE 29
Subfamily CARABINAE Latreille, 1802
Tribe CARABINI Latreille, 1802
9. Genus Calosoma Weber, 1801
Subgenus Calosoma s.str.
9.1. Calosoma inquisitor (Linnaeus, 1758)
9.1.1. Calosoma inquisitor inquisitor (Linnaeus, 1758)
References. Calosoma inquisitor inquisitor: Bousquet et al., 2003: 119.
Material studied. “Albania Rausa Dr. Grabowski”, 2 s. (HNHM); North Albanian Alps, Bogë,
1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Mal i Dajtit Mt., 27.V.2000, 1 f.,
A. Paparistos leg. (cAVT).
Notes. First detailed records for Albania.
9.2. Calosoma sycophanta (Linnaeus, 1758)
References. Calosoma sycophanta: Murraj, 1968: 33 (Sauk; Voskopojë; Shkodër); Hieke et al.,
1986: 183 (Borshi near Vlorë); Bousquet et al., 2003: 120.
Notes. Due to misidentifications found in the collection of Xh. Murraj, his literature record
for C. sycophanta (Murraj, 1968) is included here with minimal reservation because of the easily
recognition of this species.
Subgenus Microcallisthenes Apfelbeck, 1918
9.3. Calosoma pentheri Apfelbeck, 1918
References. Calosoma pentheri: Apfelbeck, 1918c: 160-163 (eastern part of the North Albanian
Alps, 1700 m, loc. typ.); Bousquet et al., 2003: 121.
Material included (studied D.W. Wrase). “Prokletia-Geb. Alban., leg Popp”, 1 m. (cWR).
9.4. Calosoma relictum Apfelbeck, 1918 [= pentheroides Roubal, 1932]
References. Calosoma relictum var. pentheroides: Roubal, 1932: 19. Calosoma relictum: Csiki, 1940:
209 (Korab Mt., Tejas); Bousquet et al., 2003: 121.
Material examined. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 53 s. (HNHM);
“Korab-Gebirge Albanien coll. Winkler”, 2 s. (HNHM).
10. Genus Carabus Linnaeus, 1758
Notes. I accept Březina’s (1999) “well geographical and morphologically defined” subspecies
when dealing with the subgenera Megodontus, Procerus and Procrustes.

31. 30 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subgenus Archicarabus Seidlitz, 1887
10.1. Carabus montivagus Palliardi, 1825
10.1.1. Carabus montivagus vellepiticus Hampe, 1850 [= sutomorensis Reitter, 1885]
References. Carabus montivagus m. sutomorensis: Breuning, 1933: 677 (Vlorë; Korab Mt.); Breuning,
1978: 194. Carabus montivagus: Hieke & Wrase, 1988: 18. Carabus montivagus vellepiticus: Březina,
1999: 25; Bousquet et al., 2003: 129; Turin et al., 2003: 198.
Subgenus Carabus s.str.
10.2. Carabus granulatus Linnaeus, 1758
10.2.1. Carabus granulatus interstitialis Duftschmid, 1812 [= aetolicus Schaum, 1857; miriditus
Apfelbeck, 1901]
References. Carabus granulatus var. miridita: Apfelbeck, 1901b: 121-122 (Lezhë, loc. typ.). Carabus
granulatus miridita: Apfelbeck, 1904: 33-34 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë
River Outfall); Monzini & Angelini, 1997: 39. Carabus granulatus aetolicus: Apfelbeck, 1904: 34
(Durrës; Vlorë); Vigna Taglianti et al., 2001: 72-73 (Velipoja near Lezhë). Carabus granulatus
granulatus n. interstitialis: Breuning, 1933: 533 (Vlorë; Durrës); Breuning, 1978: 183. Carabus
granulatus granulatus m. merdita sic!: Breuning, 1933: 534 (Velipoja near Lezhë); Breuning, 1978:
184. Carabus granulatus interstitialis v. miridita: Arnold, 1939: 158; Březina, 1999: 8 (Velipoja near
Lezhë). Carabus granulatus m. miridita: Magistretti, 1965: 26. Carabus granulatus: Hieke & Wrase,
1988: 17. Carabus granulatus interstitialis: Bousquet et al., 2003: 136.
Material included (studied A. Vigna Taglianti). Vlorë, 10.IV.1999, 1 f., A. Paparistos leg. (cAVT).
Subgenus Chaetocarabus C.G. Thomson, 1875
10.3. Carabus intricatus Linnaeus, 1761
10.3.1. Carabus intricatus intricatus Linnaeus, 1761 [= montenegrinus Kraatz, 1876; chionobatus Born,
1918; weiratheri Breuning, 1934]
References. Carabus intricatus montenegrinus: Apfelbeck, 1904: 28 (Durrës). Carabus intricatus var.:
Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m). Carabus intricatus: Klaptocz, 1910:
399 (Theth); Horion, 1941: 42. Ruspoli, 1965: 8; Geisthardt, 1975: 7; Hieke & Wrase, 1988: 16;
Turin et al., 2003: 243 (North Albania). Carabus intricatus var. montenegrinus: Apfelbeck, 1918c: 160,
163 (North Albanian Alps, 1300-2000 m). Carabus intricatus chinobatus: Buresch & Kantardjieva,
1928: 84. Carabus intricatus intricatus m. montenegrinus: Breuning, 1934: 1043, 1045 (Merdita; Tiranë;
Durrës; Vlorë). Carabus intricatus intricatus n. weiratheri: Breuning, 1934: 1046 (Vlorë); Magistretti,
1965: 45 (Vlorë); Breuning, 1978: 234 (Vlorë). Carabus intricatus intricatus var. weiratheri: Březina,
1999: 53. Carabus intricatus intricatus: Bousquet et al., 2003: 138.
Material studied. Bizë, 27.VII.1966, 1 f. (MNST); North Albanian Alps, Bogë, 1400-1800 m,
1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100
m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë,
1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Razma, 1560 m,
12.VI.1995, 1 m., V. Sakalian leg. (NMNHS).

32. COLEOPTERA: CARABIDAE: CARABINAE 31
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
1900 m, 10.VII.1993, 1 m., L. Latella leg. (cAVT).
# subgenus Eucarabus Géhin, 1876
# Carabus catenulatus Scopoli, 1763
# Carabus catenulatus catenulatus Scopoli, 1763 [= catenatus Panzer, 1803; albanensis Géhin, 1885]
References. Carabus catenatus var. albanensis: Apfelbeck, 1904: 35.
Notes. Probably, the original record for this species from Albania is that in the description of
albanensis Géhin, 1885. Apfelbeck (1904: 35) has based indirectly on this record, indicating no
exact locality for the county. I cannot find the original description of this variation, and believe
the species does not inhabit Albania (see also Turin et al., 2003).
[subgenus Hygrocarabus C.G. Thomson, 1875]
[Carabus variolosus Fabricius, 1787]
[Carabus variolosus nodulosus Creutzer, 1799]
References. Carabus variolosus nodulosus: Breuning, 1926: 19; Breuning, 1928a: 113; Buresch &
Kantardjieva, 1928: 88. Carabus nodulosus: Turin et al., 2003: 240.
Notes. Actually, the record “Albanien: Ljuboten” (Breuning, 1926: 22) concerns the territory
of Republic of Macedonia (see also Breuning, 1935: 1169). The following authors had followed
this data mentioning the species for Albania. In spite of this, C. variolosus may inhabit the
marginal northern or northeastern regions of the country.
[subgenus Limnocarabus Géhin, 1876]
[Carabus clathratus Linnaeus, 1761]
[Carabus clathratus auraniensis J. Müller, 1903]
Notes. I cannot find any exact record for this species from Albania even though Turin et al.
(2003: 155) showed in the map it occurs there. This rare Carabus-species may inhabit the
country as Purkyně (1928) described the variation epirensis Purkyně, 1928 (synonym of C.
clathratus auraniensis) from the Greek Epirus.
Subgenus Megodontus Solier, 1848
10.4. Carabus caelatus Fabricius, 1801
10.4.1. Carabus caelatus caelatus Fabricius, 1801 [= sarajevoensis Apfelbeck, 1890; malissorum
Apfelbeck, 1919; metalkanus Apfelbeck, 1919]
References. Carabus caelatus sarajevensis sic!: Apfelbeck, 1907a: 498, 502 (Mal i Shenjt Mt.: top
region, 1500 m). Carabus caelatus: Apfelbeck, 1907a: 500; Apfelbeck, 1917: 220 (Merdita);
Apfelbeck, 1930: 162 (Merdita); Horion, 1941: 41; Březina, 1999: 73. Carabus caelatus malissorum:
Apfelbeck, 1919a: 226-267 (North Albanian Alps, Maja e Jezerzë, loc. typ.); Apfelbeck, 1919b:
45-46 (North Albanian Alps, Maja e Jezerzë, loc. typ.). Carabus caelatus caelatus n. metalkanus:
Breuning, 1935: 1297 (Merdita; Gjalicë e Lumes; Sarisaltik near Kruja; “Male Dojtitz”; Mal e
Krujes); Breuning, 1978: 258. Carabus caelatus caelatus m. malissorum: Breuning, 1935: 1298 (North
Albanian Alps; Maja e Jezercë); Breuning, 1978: 258. Carabus caelatus metalkanus: Arnold, 1939:
189. Carabus caelatus metalkanus v. malissorum: Arnold, 1939: 189. Carabus caelatus caelatus: Březina,
1999: 73; Bousquet et al., 2003: 163. Carabus caelatus sarajevoensis: Turin et al., 2003: 253.

33. 32 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 2 m., P. Beron & B.
Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 3 m., P. Beron
& B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 1 m., 1 f., P.
Dioli leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg.
(cAVT); Valbona, Kollatës Mt., 1800 m, 9.VII.1993, 1 f., F. Sirtori leg. (cAVT).
Notes. Apfelbeck published two description of the form malissorum (Apfelbeck, 1919;
Apfelbeck, 1920), the older one is correctly treated as valid.
10.4.2. Carabus caelatus dalmatinus Duftschmid, 1812 [= procerus Reitter, 1885]
References. Carabus caelatus dalmatinus: Apfelbeck, 1904: 25 (Velipoja near Lezhë); Wohlberedt,
1909: 127 (near Liqen i Shkodrës); Březina, 1999: 74; Bousquet et al., 2003: 163; Turin et al.,
2003: 253 (Velipoja; Lezhë). Carabus dalmatinus procerus: Apfelbeck, 1918c: 160 (North Albanian
Alps). Carabus caelatus var. procerus: Apfelbeck, 1918c: 163 (North Albanian Alps, 1300-2000 m).
Carabus caelatus dalmatinus n. procerus: Breuning, 1935: 1300 (Lezhë District); Breuning, 1978: 258.
Material included (studied A. Vigna Taglianti). Durrës, 13,VI.1998, 1 f., A. Paparistos leg. (cAVT).
Notes. The mentioning of the locality “Vilajet” (Turin et al., 2003: 253) is misunderstanding.
Actually this Turkish word is an equivalent to the English one “district”.
10.5. Carabus croaticus Dejean, 1826
10.5.1. Carabus croaticus durmitorensis Apfelbeck, 1904 [= ljubetensis Apfelbeck, 1917; babinjensis
Apfelbeck, 1919]
References. Carabus croaticus babinjensis: Apfelbeck, 1919a: 226 (North Albanian Alps, loc. typ.);
Apfelbeck, 1919b: 45 (North Albanian Alps, loc. typ.); Krätschmer & Drovenik, 1977: 215 (Abb.1);
Turin et al., 2003: 254 (Babinje Planina Mt.). Carabus croaticus ljubetensis n. ljubetensis: Breuning,
1935: 1291 (North Albanian Alps; Babinje Planina Mt.); Breuning, 1978: 257. Carabus croaticus
ljubetensis m. durmitorensis: Breuning, 1935: 1291 (Korab Mt.); Breuning, 1978: 257. Carabus croaticus:
Horion, 1941: 41; Tabaković-Tošić, 1988: 135; Březina, 1999: 74. Carabus croaticus durmitorensis:
Březina, 1999: 74 (North Albanian Alps); Bousquet et al., 2003: 163. Carabus croaticus durmitorensis
var. babinjensis: Březina, 1999: 74 (North Albanian Alps). Carabus croaticus durmitorensis var. ljubetensis:
Březina, 1999: 74. Carabus croaticus ljubetensis: Turin et al., 2003: 254.
Notes. Apfelbeck published three consecutive descriptions of Carabus croaticus ljubetensis
(Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930), the second one has long time been
accredited to be correct. However it is wrong and the original description must be assigned to
theoldestone.Thesameauthorgavealsotwodescriptionsof Carabus croaticus babinjensis (Apfelbeck,
1919a; Apfelbeck, 1919b), as for the present it is not clear which of both has priority.
10.6. Carabus violaceus Linnaeus, 1758
10.6.1. Carabus violaceus azurescens Dejean, 1826 [= rilvensis Kolbe, 1887; zabljakensis Eidam,
1927; shljebensis Mandl, 1985]
References. Carabus violaceus azurescens n. zabljakensis: Breuning, 1935: 1261 (Maranajt Mt.; Merdita);
Breuning, 1978: 254. Carabus violaceus azurescens m. zabljakensis: Štěrba, 1945: 152 (Maranajt Mt.;
Merdita). Carabus violaceus: Hieke & Wrase, 1988: 15. Carabus violaceus azurescens var. shljebensis:
Březina, 1999: 77 (North Albanian Alps). Carabus violaceus azurescens: Turin et al., 2003: 260.

34. COLEOPTERA: CARABIDAE: CARABINAE 33
10.6.2. Carabus violaceus dryas Gistl, 1857 [= merditanus Apfelbeck, 1917; shardagensis Apfelbeck,
1917; korabensis Csiki, 1944]
References. Carabus violaceus merditanus: Apfelbeck, 1917: 220 (Merdita: Zepë Mt. near Fandi,
loc. typ.); Apfelbeck, 1918a: 75-76 (Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1930: 162 (Merdita:
Zepë Mt., loc. typ.); Cleu, 1968: 28 (Merdita); Cleu, 1969: 167. Carabus violaceus azurescens m.
merditana: Breuning, 1935: 1262 (Vermosh; Zepë Mt. near Oroshi; Peshtrik Mt.; Gjalicë e
Lumes); Breuning, 1978: 254 (Zepë Mt. near Oroshi). Carabus violaceus zabljakensis v. merditanus:
Arnold, 1939: 186. Carabus violaceus var. shardagensis: Csiki, 1940: 210 (Gjalicë Mt.; Korab Mt.).
Carabus violaceus var. korabensis: Csiki, 1944: 50-51 (Korab Mt., loc. typ.). Carabus violaceus azurescens
m. merditanus: Štěrba, 1945: 152 (Vermosh; Zebë Mt.). Carabus violaceus shardagensis: Cleu, 1968:
28 (Korab Mt.); Cleu, 1969: 167 (Korab Mt.). Carabus picenus peristericus: Tarrier, 1974: 242
(Korab Mt.). Carabus violaceus dryas: Březina, 1999: 77; Bousquet et al., 2003: 165; Turin et al.,
2003: 260. Carabus violaceus dryas var. merditanus: Březina, 1999: 77 (Merdita: Zepë Mt.; Gjalicë
e Lumes). Carabus violaceus dryas var. korabensis: Březina, 1999: 77 (Korab Mt.).
Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m,
11.VII.1993, 1 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000 m, 14.VII.1993,
rests of 1 s., A. Gobetti leg. (cAVT).
Notes. Apfelbeck published three descriptions of both C. violaceus merditanus and C. violaceus
shardagensis, respectively in Serbo-Croatian, in Latin and in German (Apfelbeck, 1917; Apfelbeck,
1918a; Apfelbeck, 1930), the second ones have long time been accredited to be correct. However
it is incorrect and the original descriptions must be assigned to the oldest ones. A revision of
the type of the var. shardagensis Apfelbeck, 1917 is needed to elucidate to which subspecies,
azurescens or dryas, it belongs. Csiki (1944: 50, Fig. 1a) represented a figure of this variation
which let me assume that it belongs rather to the former subspecies.
# Carabus violaceus exasperatus Duftschmid, 1812 [= vlasuljensis Apfelbeck, 1894]
References. Carabus violaceus vlasuljensis: Apfelbeck, 1907a: 502 (Merdita); Apfelbeck, 1918c:
160, 163 (North Albanian Alps, Vermosh).
Notes. The easternmost known records of ssp. exasperatus come from Bosnia Herzegovina, so that
the literature data for it should be referred to some of both subspecies that really live in Albania.
Subgenus Oreocarabus Géhin, 1876
Notes. In my opinion, C. hortensis, C. neumeyeri and C. preslii represent separate species by
reason of its clear and constant differences in both the overall aspect of the pronotum and
the penis (Casale et al., 1982: 185, 190 – fig. 75, 195). Externally, the first species can be easily
distinguished from the other two in the more quadrate aspect of the pronotum (vs. subovate
or trapezium-shaped), the lateral border of pronotum posteriorly sinuate (vs. straight to slightly
sinuate) and in the lateral lobes of pronotum clearly horizontal or slightly oblique (vs. oblique).
Additionally C. neumeyeri has lateral borders of pronotum posteriorly more convergent than
those in C. hortensis. The following nomenclatorial acts are proposed: Carabus (Oreocarabus)
neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri Schaum, 1856;
Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii
Dejean, 1830 (see Casale et al., 1982: 194).

35. 34 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
10.7. Carabus hortensis Linnaeus, 1758 [= rhodopensis Apfelbeck, 1904]
References. Carabus hortensis: Apfelbeck, 1918c: 163 (North Albanian Alps: Vermosh). Carabus
hortensis rhodopensis: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near Vermosh, 1100-
1750 m). Carabus hortensis hortensis: Breuning, 1933: 709 (Peshtrik Mt.; Vermosh); Bousquet et
al., 2003: 179.
Material included (studied A. Vigna Taglianti). Tropoia, Valle di Valbona, 900 m, 8-
16.VII.1993, 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 1900 m, 10.VII.1993, 1 f., L.
Latella leg. (cAVT); Valbona, Kollatës Mt., 1000 m, 14.VII.1993, 1 f., L. Latella leg. (cAVT).
10.8. Carabus neumeyeri Schaum, 1856 [= herzegowinensis Apfelbeck, 1904; pindicus Apfel-
beck, 1904; albanicola Breuning, 1965; hilleri Mandl, 1964]
References. Carabus hortensis neumeyeri: Apfelbeck, 1904: 42 (Velipoja); Apfelbeck, 1907a: 496
(Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m); Apfelbeck, 1909:
297 (Berat); Schatzmayr, 1943b: 94 (Shendelliut); Březina, 1999: 46; Bousquet et al., 2003: 179.
Carabus hortensis neumeyeri n. neumeyeri: Breuning, 1933: 710 (Sarisaltik near Kruja; Vlorë;);
Breuning, 1978: 198. Carabus hortensis neumeyeri n. herzegowinensis: Breuning, 1933: 712 (Velipoja
near Lezhë). Carabus hortensis: Hieke & Wrase, 1988: 19. Carabus hortensis neumeyeri var. pindicus:
Březina, 1999: 46. Carabus hortensis neumeyeri var. albanicola: Březina, 1999: 46 (Llogora). Carabus
hortensis neumeyeri var. hilleri: Březina, 1999: 46 (Reza e Kanalit near Llogora). Carabus preslii
neumeyeri: Turin et al., 2003: 228.
Material studied. Kurbneshi, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); North
Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS);
North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg.
(NMNHS).
10.9. Carabus preslii Dejean, 1830 [= jonicus Apfelbeck, 1904]
References. Carabus hortensis preslii m. jonica: Breuning, 1933: 713 (Gjirokastër); Breuning,
1978: 198.
Material studied. Pogyara, 1027 m, 4.VI.1994, 2 f., G. Blagoev leg. (NMNHS); Leskovac,
650 m, 2.VI.1995, 1 f., V. Sakalian leg. (NMNHS).
Notes. Bousquet et al. (2003: 179) has omitted the taxon for Albania, so that it is confirmed
here for the country.
Subgenus Pachystus Motschulsky, 1866
10.10. Carabus cavernosus I. Frivaldszky von Frivald, 1838
10.10.1. Carabus cavernosus cavernosus I. Frivaldszky von Frivald, 1838 [= sterbai Breuning, 1936]
References. Carabus cavernosus cavernosus: Breuning, 1933: 798 (Korab Mt.; Gjalicë e Lumes);
Guéorguiev et al., 1997: 8; Březina, 1999: 35; Bousquet et al., 2003: 181; Turin et al., 2003: 234.
Carabus cavernosus var. sterbai: Csiki, 1940: 210 (Korab Mt.). Carabus cavernosus n. cavernosus:
Breuning, 1978: 207. Carabus cavernosus: Hieke & Wrase, 1988: 15.

36. COLEOPTERA: CARABIDAE: CARABINAE 35
10.11. Carabus graecus Dejean, 1826
10.11.1. Carabus graecus morio Mannerheim, 1830 [= winkleri Breuning, 1933; neowinkleri Deuve, 1991]
References. Carabus graecus winkleri: Breuning, 1933: 802 (Maja e Tartarit Mt., 800 m, loc. typ.).
Carabus graecus n. winkleri: Breuning, 1978: 207 (Maja e Tartarit Mt.). Carabus graecus: Hieke & Wrase,
1988: 14; Březina, 1999: 36. Carabus graecus morio: Březina, 1999: 36; Bousquet et al., 2003: 182.
Carabus graecus morio var. winkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus morio var.
neowinkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus graecus: Turin et al., 2003: 236.
? subgenus Platycarabus A. Morawitz, 1886
? Carabus irregularis Fabricius, 1792
? Carabus irregularis ramanus Sokolář, 1909 [= bucephalus Kraatz, 1879]]
References. Carabus irregularis bucephalus: Turin et al., 2003: 249.
Notes. Turin et al. (2003) showed the species range reaching North Albania. According to
Březina (1999: 53) ssp. ramanus is known from the Bjelasièa Mt. in Northeastern Montenegro
not very far from the studied territory. The species may live in Albania.
Subgenus Procerus Dejean, 1821
10.12. Carabus gigas Creutzer, 1799
10.12.1. Carabus gigas gigas Creutzer, 1799 [= parnassicus Kraatz, 1884]
References. Carabus gigas parnassicus: Březina, 1999: 72. Carabus gigas gigas: Bousquet et al., 2003: 185.
Material studied. Malesia e Polisit Mt., 1850 m, near Sopoti, VI.1966, 1 s. (cXM).
Notes. First detailed record for Albania.
Subgenus Procrustes Bonelli, 1810
10.13. Carabus coriaceus Linnaeus, 1758
10.13.1. Carabus coriaceus cerisyi Dejean, 1826 [= hopffgarteni Kraatz, 1877; subrugosus Kraatz,
1877; weiratherianus Breuning, 1936]
References. Carabus coriaceus cerisyi m. weiratheriana: Breuning, 1937: 1406 (Korab Mt.); Breuning,
1978: 270. Carabus coriaceus subrugosus v. weiratherianus: Arnold, 1939: 192. Carabus coriaceus var.
hopffgarteni: Csiki, 1940: 209 (Kula Ljums; Gjalicë Mt.; Korab Mt.). Carabus coriaceus subrugosus:
Hieke et al., 1986: 183 (Borshi near Vlorë). Carabus coriaceus: Hieke & Wrase, 1988: 13.
10.13.2. Carabus coriaceus excavatus Charpentier, 1825 [= rugosus Dejean, 1826; dalmaticus Géhin,
1885; joniensis Breuning, 1932; albanicus Breuning, 1943]
References. Carabus coriaceus rugosus: Apfelbeck, 1904: 20 (Velipoja; Berat; Vlorë); Apfelbeck,
1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 297 (Ardenica; Vlorë;
Pojani). Carabus coriaceus rugosus n. joniensis m. albanicus: Breuning, 1932: 20 (Maja e Tartarit Mt.,
loc. typ.); Mařan, 1949: 3 (Maja e Tartarit Mt.). Carabus coriaceus excavatus m. dalmatica: Breuning,
1937: 1403 (Velipoja; Kruja; Durrës; Merdita; Tiranë; Berat; Vlorë; south of Vlorë); Breuning,
1978: 269. Carabus coriaceus excavatus n. joniensis: Breuning, 1937: 1404 (Vlorë; Gjirokastër);
Magistretti, 1965: 55; Breuning, 1978: 269. Carabus coriaceus excavatus: Arnold, 1939: 191; Hieke

37. 36 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
et al., 1986: 183 (Mal i Dajtit Mt.: Shkall Priskë); Březina, 1999: 62; Bousquet et al., 2003: 187;
Turin et al., 2003: 278. Carabus coriaceus excavatus dalmaticus: Schatzmayr, 1943b: 94 (Lushnje;
Karbunara e Madhe; Dukati, south of Vlorë). Carabus coriaceus joniensis: Březina, 1999: 62;
Bousquet et al., 2003: 187. Carabus coriaceus joniensis: var. albanicus: Březina, 1999: 62 (Maja e
Tartarit Mt.). Carabus coriaceus mediterraneus: Turin et al., 2003: 278.
Material studied. Sauk, 18.II.1960, 1 m. (MNST); Ibë, 3.VII.1962, 1 f. (MNST); Bizë,
27.VII.1966, 1 f. (MNST); Tiranë, 27.III.1968, 1 f. (MNST); Tiranë Park, 22-31.X.1997, 2 s.,
R. Ward leg. (CMNH); Tiranë Park, 1-4.I.1998, 2 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
2250 m, 11.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000
m, 14.VII.1993, 1 f., A. Gobetti leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley
of Vijosë River, 300-360 m, 18.IV.2003, 2 m., P. Crucitti & C. Cavalieri leg. (cAVT).
Note: According to Turin et al. (2003) C. coriaceus mediterraneus Born, 1906 has transionian
distribution and lives in Albania.
Subgenus Tachypus Weber, 1801
10.14. Carabus cancellatus Illiger, 1798
10.14.1. Carabus cancellatus alessiensis Apfelbeck, 1901 [= hypsobius Apfelbeck, 1918]
References.Carabuscancellatusvar.alessiensis:Apfelbeck,1901b:121(Lezhë,loc.typ.).Carabuscancellatus
alessiensis:Apfelbeck,1904:30;33(VelipojanearLezhë);Apfelbeck,1907a:496(BuenëRiverOutfall);
Breuning, 1933: 594 (Velipoja near Lezhë; Shkodër; Pulaj); Arnold, 1939: 161 (North Albania);
Breuning, 1978: 187 (Velipoja); Březina, 1999: 41 (Velipoja near Lezhë); Bousquet et al., 2003: 195;
Turin et al., 2003: 194. Carabus cancellatus subgraniger var. alessiensis: Bernau, 1913: 198 (Lezhë). Carabus
cancellatus emarginatus: Bernau, 1913: 200. Carabus cancellatus: Hieke & Wrase, 1988: 17.
Material studied. Shkodër, 14.V.1963, 2 m. (MNST).
Notes. The type locality of the form hypsobius Apfelbeck, 1918c: 163 is Vunšaj (in Albanian)
or Vusanje (in Serbo-Croatian) in the North Albanian Alps. It is situated in the Montenegrin
territory very closely to Albania. C. cancellatus emarginatus Duftschmid, 1812 is a valid subspecies
reaching to Bosnia Herzegovina in the east.
Subgenus Tomocarabus Reitter, 1896
10.15. Carabus convexus Fabricius, 1775
10.15.1. Carabus convexus dilatatus Dejean, 1826 [= chionophilus Apfelbeck, 1904; miliusi Breuning, 1927]
References. Carabus convexus var.: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500
m). Carabus convexus dilatatus: Breuning, 1933: 874 (Vlorë; Gjirokastër); Bousquet et al., 2003:
198; Turin et al., 2003: 230. Carabus convexus dilatatus n. miliusi: Breuning, 1933: 876 (North
Albanian Alps; Merdita); Breuning, 1978: 214. Carabus convexus miliusi: Arnold, 1939: 178. Carabus
convexus var. chionophilus: Csiki, 1940: 210 (Korab Mt.). Carabus convexus: Hieke & Wrase, 1988:
19. Carabus convexus dilatatus var. miliusi: Březina, 1999: 33.
Material studied. Mal i Dajtit Mt., 11.V.1965, 1 m. (MNST); North Albanian Alps, Maj’e
Radohimes, 1900-2200, 28.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS);

38. COLEOPTERA: CARABIDAE: ELAPHIRINAE 37
North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS);
North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 3f., P. Beron & B. Petrov leg. (NMNHS).
? subgenus Trachycarabus Géhin, 1876
? Carabus scabriusculus Olivier, 1795
? Carabus scabriusculus bulgarus Lapouge, 1908
References. Carabus scabriusculus bulgarus: Turin et al., 1993: 42.
Notes. This species is questionable for Albania due to lack of exact records. On the other
side, it was quoted for the neighboring countries Montenegro and Serbia.
Tribe CYCHRINI Laporte, 1834
11. Genus Cychrus Fabricius, 1794
11.1. Cychrus semigranosus Palliardi, 1825
# Cychrus semigranosus semigranosus Palliardi, 1825
References. Cychrus semigranosus semigranosus: Häckel, 2003: 205.
Notes. This record is referred to the ssp. montenegrinus because the nominotypical race does
not populate Albania.
# Cychrus semigranosus balcanicus Hopffgarten, 1881
References. Cychrus semigranosus n. balcanicus: Mařan, 1940c: 133-134 (North Albanian Alps).
Notes. This East Balkan race does not inhabit Albania, therefore it is removed from the species list.
# Cychrus semigranosus montanus Winkler, 1939
References. Cychrus semigranosus montanus: Häckel, 2003: 205.
Notes. Winkler (1939: 199) described this subspecies on material from two different regions - the
EasternCarpathiansandWesternBalkans.Mandl(1989:69)designatedlectotypeandparalectotypes
for the specimens from the former region, and syntypes of montanus from the latter region were
assigned to C. semigranosus dinaricus Mandl, 1989 (“Cvrstnica”, loc. typ.). Presumably, the subspecies
mentioning for Albania (Häckel, 2003: 205) is based on incorrect interpretation of the locality
“Zljeb planina nördlich der Nord-albanischen Alpen” (Winkler, 1939: 199; Mařan, 1940c: 133;
Mandl, 1989: 68). In fact the Zljeb Mt. is situated in the Kosovo Region of Serbia.
11.1.1.Cychrus semigranosus montenegrinus Apfelbeck, 1904[=albanicusHopp,1929;dinaricusMandl,1989]
References.Cychrussemigranosussemigranosus:Röschke,1907:255;Häckel,2003:205.Cychrussemigranosus:
Apfelbeck, 1907a: 498, 503 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 210 (Shkëlzen Mt.;
Korab Mt.). Cychrus semigranosus albanicus: Hopp, 1929: 2 (Tomor Mt, 2000 m, loc. typ.); Schatzmayr,
1943b: 94 (Tomor Mt.); Mandl, 1989: 69 (Peshtrik Mt.; Gjalicë Mt.; Çukall Mt.; Merdita); Häckel,
2003:205.Cychrussemigranosusssp.:Schatzmayr,1943b:93(Shendelliut).Cychrussemigranosusmontenegrinus:
Winkler, 1939: 200 (Llogora; Tomor Mt.; Kruja). Cychrus semigranosus n. montenegrinus: Mařan, 1940c:
128, 130-132, 135-136 (Llogora). Cychrus semigranosus montanus: Häckel, 2003: 205.
Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Alban.
Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exp.’18. Pashtrik 4-14.7”, 3 s. (NMW);
“Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Giallica” / “Albania leg. Bischoff 1936”, 1
s. (NMW).

39. 38 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Winkler (1939) synonymized the ssp. albanicus with ssp. montenegrinus. Mařan (1940c)
accepted this point of view, but Mandl (1989) treated both as separate subspecies. Samples of
the ssp. montenegrinus, ssp. albanicus, as well as the holotype and paratypes of the ssp. dinaricus
have been studied in NMW. Differences between specimens from different localities exist
mostly in the sizes of the body and shape of the pronotum, but they also have occurred in the
limits of one and the same population. In reality characters that can support well-defined
geographical races in the Western Balkans, excluding the well-differentiated ssp. travnikanus
Apfelbeck, 1904 (type series in NMW) and the not studied ssp. jakupicensis Mařan, 1943, are
absent. Therefore the following synonymy is proposed: Cychrus semigranosus albanicus Hopp,
1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck, 1904; Cychrus semigranosus dinaricus
Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck, 1904. In MNHUB, I
have furthermore seen 69 s. from different localities, which remained unregistered.
Subfamily ELAPHRINAE Erichson, 1837
Tribe ELAPHRINI Erichson, 1837
12. Genus Elaphrus Fabricius, 1775
Subgenus Neoelaphrus Hatch, 1951
12.1. Elaphrus uliginosus Fabricius, 1792
References. Elaphrus uliginosus: Murraj, 1968: 33 (Bizë).
Material studied. “N 88”, 1 f. (MNST).
Notes. The record of Murraj (1968) is included here with minimal reservation because of his
correct determination of the female above. Goulet (2003: 207) omitted the species for Albania,
but herewith it is confirmed for the country.
Subfamily OMOPHRONINAE Bonelli, 1810
Tribe OMOPHRONINI Bonelli, 1810
13. Genus Omophron Latreille, 1802
Subgenus Omophron s.str.
13.1. Omophron limbatum (Fabricius, 1777)
References. Omophron limbatum: Hieke et al., 1986: 183 (Bizë near Shëngjergji); Hieke & Wrase,
1988: 23; Hůrka, 2003: 208.
Material studied. “Skutari Bojana”, 3 s. (NMW); “Oboti Bojana” / “Apfelb. Alban. spt. IV-V.05”,
1 s. (NMW); “Berat” / “Albania leg. Bischoff 1931”, 17 s. (MNHUB); “Ibe-Erzen” / “Albania leg.
Bischoff 1932”,2s.(MNHUB);“Tepelene”/“Albanialeg.Bischoff 1932”,4s.(MNHUB);“Vojusa

40. COLEOPTERA: CARABIDAE: BRACHININAE 39
Dragot” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff
1937”, 1 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).
Subfamily BRACHININAE Bonelli, 1810
Tribe BRACHININI Bonelli, 1810
14. Genus Aptinus Bonelli, 1810
Subgenus Aptinus s.str.
14.1. Aptinus acutangulus Chaudoir, 1876
Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.6.1993, 1 f., P. Beron & B. Petrov
leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.6.1993, 1 m., 1 f., P. Beron & B.
Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 15 m., 14 f., P.
Dioli leg. (cAVT).
Notes. New species for the fauna of Albania.
14.2. Aptinus bombarda (Illiger, 1800)
References. Aptinus bombarda: Apfelbeck, 1918c: 168 (North Albanian Alps, Vermosh, 1200
m); Hůrka, 1988: 290-291 (Shkëlzen Mt.); Hrdlička, 2003: 212.
Material studied. “Vermosa’14 Penther”, 1 m. (NMW); “Prokletija 1300-2000 m” / “Alban.
montenegr. Grenze, Penther’14”, 1 f. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban.
montenegr. Grenze, Penther’14”, 1 m. (NMW); “Maja e Jezerce Albania sept.”, 1 m. (NMW);
“Shkëlzën” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); North Albanian Alps, Bogë, 1400-
1800 m, 1.6.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
2000 m, 11.VII.1993, 2 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 1000 m,
14.VII.1993, 1 m., L. Latella leg. (cAVT).
Notes. A. bombarda occurs sympatrically with A. acutangulus in the North Albanian Alps (see
also Casale & Vigna Taglianti, 1983: 36; Hůrka, 1988: 289); the former lives at the same altitude
or higher than the latter. The study of the penis reveals that the male from Bogë has somewhat
differing apex compared with those from Vermosa and Maja e Jezercoë. The penis apex in the
latter is identical with the illustration of Casale & Vigna Taglianti (1983: 32, Fig. 8). Very likely
the population from the massif of Bogë forms a separate race.
14.3. Aptinus merditanus Apfelbeck, 1917
14.3.1. Aptinus merditanus merditanus Apfelbeck, 1917 [= granulipennis Apfelbeck, 1917]
References. Aptinus lugubris rufipes: Apfelbeck, 1907a: 499, 501 (Mal i Shenjt Mt.: top region,
1500 m). Aptinus merditanus: Apfelbeck, 1917: 227 (Merdita: Zepë Mt. near Fandi, loc. typ.);
Apfelbeck, 1918b: 89 (Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1930: 171-172

41. 40 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
(Merdita: Zepë Mt. near Fandi, loc. typ.); Casale & Vigna Taglianti, 1983: 29, 39, 41 (Merdita:
Zepë Mt.; Shen Thanas Mt. near Vlorë); Hůrka, 1988: 293-294 (Mal i Shenjt Mt.; Zepë Mt.;
Mal i Dajtit Mt.; Mal i Krujës Mt.; Shen Ili; Maja e Tartarit Mt.; Cafa Kjorës; Tomor Mt.; Shen
Thanas Mt.; Llogora); Wrase, 1991: 18. Aptinus merditanus var. granulipennis: Apfelbeck, 1917:
227 (South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt.
near Joannina, loc. typ.); Apfelbeck, 1918b: 89 (South Albania: Shen Thanas Mt. near Vlorë,
loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.); Apfelbeck, 1930: 172
(South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt.
near Joannina, loc. typ.). Aptinus merditanus merditanus: Hrdlička, 2003: 212.
Materialstudied. “MerditaM.Schèit” /“Coll. Apfelbeck”/“merditanus Typ. Apf.”,1m. (HNHM);
1 s. from “Çafa Kjores”, 53 s. from “Llogora” and “Logara, 8 s. from “Maja e Tartarit”, 40 s. from
“Mal i Dajtit”, 1 s. from “Mal i Krujes”, 10 s. from “Shen Ili”, 1 s. from “Tomor” (MNHUB).
Notes. Apfelbeck published three consecutive descriptions for both A. merditanus and A.
merditanus var. granulipennis (Apfelbeck, 1917; Apfelbeck, 1918b; Apfelbeck, 1930) and the second
ones have long time been accredited to be valid. Actually this is incorrect and the original
descriptions must be assigned to the oldest ones. Aptinus lugubris rufipes (Apfelbeck, 1907a:
499) is nomen nudum (see also Hůrka, 1988: 292). In that case, Apfelbeck had most probably the
intention to introduce a name for A. merditanus earlier than the year of 1917.
15. Genus Brachinus Weber, 1801
Subgenus Brachinus s.str.
# Brachinus alexandri F. Battoni, 1984
References. Brachinus alexandri: Guéorguiev & Guéorguiev, 1995: 239.
Notes. Dealing with B. plagiatus data in the paper of Hùrka (1988), I noted that B. alexandri
was recorded by mistake for Albania (Guéorguiev & Guéorguiev, 1995). Most likely
Hrdlička(2003: 213) followed this mistake marking the species for the country. Here B. alexandri
is removed from the list of Albanian ground beetles.
15.1. Brachinus crepitans (Linnaeus, 1758) [= fallax Apfelbeck, 1904]
References. Brachinus crepitans: Apfelbeck, 1904: 348 (Vlorë; Berat); Apfelbeck, 1909: 298 (Vlorë;
Berat); Csiki, 1940: 222 (Kula Ljums; Gjalicë Mt.); Hieke & Wrase, 1988: 163; Hrdlička, 2003:
213. Brachinus crepitans ab. fallax: Apfelbeck, 1904: 348 (Vlorë); Csiki, 1940: 222 (Kula Ljums).
Material studied. “Avlona”, 1 s. (HNHM); “Han Holfit 5.-10. Mai” / “Alban. montenegr. Grenze,
Penther’14”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Alban-Exp.’18. Pashtrik, 4-14.7.”, 1 s. (NMW); “Dures” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Lume i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Iballe” / “Albania leg. Bischoff
1939”, 1 s. (MNHUB); “Albania Fush-Dukati” / “7.V.1958 leg. Xh. Murraj”, 1 s. (HNHM);
Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);
Ljukove, 100 m, 5.VI.1995, 1 f., G. Blagoev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2
m., 1 f., D. Antonelli leg. (cAVT).

42. COLEOPTERA: CARABIDAE: BRACHININAE 41
15.2. Brachinus ejaculans Fischer von Waldheim, 1828
References. Brachinus ejaculans: Apfelbeck, 1904: 351 (Velipoja; Vlorë); Hieke & Wrase, 1988:
165; Hrdlička, 2003: 213.
Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Miloti, Alb. Matzenauer”, 3
s. (NMW); “Shullaz” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 11 s. (MNHUB);
“Rushkull” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).
15.3. Brachinus elegans Chaudoir, 1842 [= ganglbaueri Apfelbeck, 1904]
References. Brachynus psophiae or ganglbaueri: Klaptocz, 1910: 399 (Shkodër); Schatzmayr, 1943b:
113 (Lushnje). Brachinus ganglbaueri: Hieke & Wrase, 1988: 164.
Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania
leg. Bischoff 1936”, 5 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. Hrdlička (2003: 213) has omitted the species for Albania. Here it is confirmed for the
country.
15.4. Brachinus plagiatus Reiche, 1868
References. Brachinus plagiatus: Apfelbeck, 1904: 350 (Velipoja; Vlorë); Horion, 1941: 353;
Schatzmayr, 1943b: 112 (Skrofotina); Hieke & Wrase, 1988: 165; Hùrka, 1988: 205 (Tiranë).
Material studied. “Avlona”, 6 s. (HNHM); “Durazzo”, 2 s. (HNHM); “Alban. Wgth. Valona”, 9
s. (NMW); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Rushkull” / “Albania leg.
Bischoff 1936”, 3 s. (MNHUB).
Notes. Hrdlička (2003: 213) has omitted the species for Albania. Here it is confirmed for the
country.
15.5. Brachinus psophia Audinet-Serville, 1821
References. Brachinus psophia: Apfelbeck, 1904: 349 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);
Hieke & Wrase, 1988: 164; Hrdlička, 2003: 213. “Brachynus psophiae aut Ganglbaueri”: Schatzmayr,
1943b: 113 (Lushnje).
Material studied. “Avlona”, 1 s. (HNHM); “Alban. Wgth. Valona”, 2 s. (NMW); Radona,
Shkiela, 21.VI.1994, 1 m., B. Guéorguiev leg. (NMNHS).
Subgenus Brachynidius Reitter, 1919
15.6. Brachinus bodemeyeri Apfelbeck, 1904
References. Brachinus bodemeyeri: Apfelbeck, 1904: 352 (Vlorë); Hieke & Wrase, 1988: 166;
Hrdlička, 2003: 213.
Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Alban. Wgth. Valona”, 4 s.
(NMW); “Dukati” / “ex Coll. Scheibel”, 1 s. (MNHUB).

43. 42 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
15.7. Brachinus brevicollis Motschulsky, 1844 [= peregrinus Apfelbeck, 1904]
References. Brachinus peregrinus: Apfelbeck, 1904: 351 (Vlorë, loc. typ.); Reitter, 1919: 140;
Schatzmayr, 1935: 245; Schatzmayr, 1943b: 112 (Tërpan; Dukati; Lushnje; Vlorë); Hieke &
Wrase, 1988: 165. Brachinus brevicollis: Hrdlička, 2003: 213.
Material included (studied A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District),
valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
15.8. Brachinus explodens Duftschmid, 1812 [= glabratus Dejean, 1824; strepens Fischer von
Waldheim, 1828]
References. Brachinus explodens: Apfelbeck, 1904: 353 (Vlorë; Berat); Apfelbeck, 1909: 298
(Vlorë; Berat); Csiki, 1940: 222 (Ura i Lopez); Schatzmayr, 1943b: 113 (Dukati); Hieke et al.,
1986: 190 (North Albanian Alps near Theth); Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214.
Brachinus explodens ab. strepens: Csiki, 1940: 222 (Ura i Lopez; “Stičen” near Gjalicë Mt.). Brachinus
explodens ab. glabratus: Csiki, 1940: 222 (Kula Ljums; Ura i Lopez).
Material studied. “Avlona”, 2 s. (HNHM); “Skutari-Holcekai” / “Alban. montenegr. Grenze,
Penther’14”, 12 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM);
“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura i
Lopez” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 8
s. (NMW); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Cerrik”
/ “16.IV.1958 leg. Xh. Murraj”, 7 s. (HNHM); Tiranë, 7.IV.1966, 1 s. (MNST); Vlorë, 1.V.1994,
2 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg.
(NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev
leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-
4.I.1998 / 20-25.I.1998 / 1.III.1998 / 8.III.1998, 13 s., shaded lake edge under leaves and wet,
rotten wood, R. Ward leg. (CMNH).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near
Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti &
C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M.
Komnenov leg. (cSH).
Notes. I have seen furthermore 160 s. from different localities in MNHUB, which remained
unregistered.
15.9. Brachinus sichemita Reiche & Saulcy, 1855
References. Brachinus explodens sichemita: Apfelbeck, 1904: 353 (Berat; Vlorë).
Notes. Hrdlička (2003: 214) has omitted the species for Albania, therefore its occurrence in
the country needs further confirmation.
Subgenus Brachynolomus Reitter, 1919
15.10. Brachinus immaculicornis Dejean, 1826
15.10.1. Brachinus immaculicornis immaculicornis Dejean, 1826
References. Brachinus immaculicornis: Apfelbeck, 1909: 298 (Vlorë).

44. COLEOPTERA: CARABIDAE: SIAGONINAE 43
Notes. Hrdlička (2003: 214) has omitted the species for Albania; therefore its occurrence in
the country needs further confirmation.
Subgenus Cnecostolus Reitter, 1919
15.11. Brachinus bayardi Dejean, 1831
References. Brachinus bayardi: Schatzmayr, 1943b: 112 (Mal i Krujes).
Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);
“Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Notes. Hrdlička (2003: 214) has omitted the species for Albania. Here it is confirmed for the
country.
15.12. Brachinus bipustulatus Quensel, 1806
References. Brachinus bipustulatus: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909:
298 (Vlorë); Reitter, 1919: 135; Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214.
Material studied. “Scutari” / “collectio Paganetti”, 1 s. (NMW); “Mustajbeg Velipoja Albania”,
1 s. (MNHUB); “Durazzo Apfelbeck”, 1 s. (HNHM); “Oboti Bojana”, 4 s. (HNHM); “Alban.
Wgth. Valona”, 1 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB);
“Myzeqe” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
15.13. Brachinus exhalans (P. Rossi, 1792)
References. Brachinus exhalans: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909: 298
(Vlorë); Schatzmayr, 1943b: 112 (Lushnje; Mal i Krujes); Hrdlička, 2003: 214.
Material studied. “Avlona”, 3 s. (HNHM); “Avlona Albania mer.”, 1 s. (HNHM); “Albanien
Avlona v. Oertzen.”, 5 s. (MNHUB); “Durazzo Apfelbeck”, 2 s. (HNHM); “Alban. Wgth.
Valona”, 6 s. (NMW); “Murshi” / “Albania leg. Bischoff 1936”, 111 s. (MNHUB); Radona,
Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).
Species insertae sedis
15.14. Brachinus nigricornis Gebler, 1829 [= incertus Brullé, 1834]
References. Brachinus incertus: Apfelbeck, 1904: 350 (Velipoja near Lezhë); Apfelbeck, 1909:
298 (Vlorë); Hieke & Wrase, 1988: 165. Brachinus nigricornis: Hrdlička, 2003: 216.
Material studied. “Durazzo”, 1 s. (HNHM); “Albania Avlona”, 5 s. (HNHM); “Bojana
Mundung”, 3 s. (MNHUB); “Alban. Wgth. Valona”, 12 s. (NMW).

45. 44 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subfamily SIAGONINAE Bonelli, 1813
Tribe SIAGONINI Bonelli, 1813
16. Genus Siagona Latreille, 1804
16.1. Siagona europaea Dejean, 1826
16.1.1. Siagona europaea europaea Dejean, 1826 [= depressa Bedel, 1887 (nec Fabricius, 1798)]
References. Siagona europaea: Apfelbeck, 1904: 358 (Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes;
Lushnje; Skrofotina); Hieke, 1981: 76; Hieke & Wrase, 1988: 27. Siagona depressa: Apfelbeck,
1909: 297 (Vlorë). Siagona europaea europaea: Baehr, 2003a: 218.
Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Krabe” /
“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 2 s.
(MNHUB); “Liqen i Butrinten Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tirane”
/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB); “Tirane” / “leg. Bischoff Albania 1937”, 2 s. (MNHUB).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near
Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., P. Crucitti
& C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M.
Komnenov leg. (cSH).
? Siagona europaea oberleitneri Dejean, 1830
References. Siagona europaea var. oberleitneri: Ganglbauer, 1904: 648.
Notes. This race is endemic for the Ionic Islands. Taking into account the proximity of these
islands to South Albania further collectings are needed to determine the subspecies status of
the populations in the studied territory.
Subfamily SCARITINAE Bonelli, 1810
Tribe CLIVININI Rafinesque, 1815
Subtribe CLIVININA Rafinesque, 1815
17. Genus Clivina Latreille, 1802
17.1. Clivina collaris (Herbst, 1784)
References. Clivina collaris: Balkenohl, 2003: 219.
Material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 8-
16.VII.1993, 1 m., L. Latella leg. (cAVT).
Notes. First detailed record for Albania.

46. COLEOPTERA: CARABIDAE: SCARITINAE 45
17.2. Clivina fossor (Linnaeus, 1758)
17.2.1. Clivina fossor fossor (Linnaeus, 1758)
References. Clivina fossor: Apfelbeck, 1904: 68 (Velipoja near Lezhë; Vlorë); Apfelbeck, 1909:
297 (Vlorë); Hieke et al., 1986: 184 (Shkodër); Hieke & Wrase, 1988: 27. Clivina fossor fossor:
Balkenohl, 2003: 219.
Material studied. “Scutary Bojana”, 3 s. (NMW); “Scutary i. See Ufer”, 2 s. (NMW); “Scutary
Vraka”, 3 s. (NMW); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mamurash” /
“Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet,
rotten wood, R. Ward leg. (CMNH).
17.3. Clivina laevifrons Chaudoir, 1842
References. Clivina laevifrons: Apfelbeck, 1904: 69 (Velipoja); Rambousek, 1912: 69; Hieke &
Wrase, 1988:28; Balkenohl, 2003: 220.
Material studied. “Scutary Bojana”, 1 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.”, 1 s. (NMW);
“Scutary i. See Ufer”, 2 s. (NMW); “Scutary Vraka”, 1 s. (NMW).
Subtribe REICHEINA Jeannel, 1957
! genus Dalmatoreicheia Magrini & Bulirsch, 2005
! Dalmatoreicheia sp.
Material studied. “Kruja, Alban., Mader”, 1 m. (NMW).
Notes. The specimen belongs to a new species.
18. Genus Reicheadella Reitter, 1913
Notes. The Albanian species of this subtribe are object of taxonomic work (Bulirsch &
Guéorguiev, in preparation).
Subgenus Reicheadella s.str.
18.1. Reicheadella bischoffi (Meschnigg, 1933)
References. Reicheia bischoffi: Meschnigg, 1933:172 (Tomor Mt. near Berat, loc. typ.); Turin,
1981: 72. Reicheadella bischoffi: Jeannel, 1957: 179; Casale et al., 1998: 95; Balkenohl, 2003: 222.
Material studied. “Tomor Juli 1932” / “Albania leg. Bischoff 1932”, 2 syntypes (MNHUB);
“Tomor: Tomor Alban. mer.” / “lg. Winkler Mai 1931” / “bischoffi Meschn. det. A. Winkler”,
2 m. (NMW).
! Reicheadella sp.
Material studied. “Albania mer. Liqueni i Butrintit, 1.V.1958; A. Smetana leg.”, 1 f. (NMPC).
Notes. The specimen belongs to a new species.

47. 46 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subtribe DYSCHIRIINI W. Kolbe, 1880
19. Genus Dyschirius Bonelli, 1810
Subgenus Chiridysus Fedorenko, 1996
19.1. Dyschirius strumosus Erichson, 1837
References. Dyschirius strumosus: Balkenohl, 2003: 224.
Subgenus Dyschiriodes Jeannel, 1941
Species group “integer”
19.2. Dyschirius intermedius Putzeys, 1846
References. Dyschirus intermedius: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 78
(Oboti); Hieke & Wrase, 1988: 31; Fedorenko, 1993: 819; Balkenohl, 2003: 225.
Species group “nitidus”
19.3. Dyschirius agnatus Motschulsky, 1844 [= lucidus Putzeys, 1846]
References. Dyschirius lucidus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës). Dyschirius
agnatus: Balkenohl, 2003: 224.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 9 s. (HNHM);
“Tepelene” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durrës” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Bojana
Shkoder” / “Albania leg. Bischoff 1937”, 5 s. (MNHUB).
19.4. Dyschirius nitidus (Dejean, 1825)
19.4.1. Dyschirius nitidus nitidus (Dejean, 1825)
References. Dyschirius nitidus: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 63-64 (Berat);
Csiki, 1940: 212 (Kula Ljums); Horion, 1941: 98; Ieniștea, 1979: 224; Gryuntal, 1984b: 514;
Hieke & Wrase, 1988: 29.
Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).
Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here it is confirmed for the country.
Species group “minutus”
19.5. Dyschirius aeneus (Dejean, 1825)
19.5.1. Dyschirius aeneus aeneus (Dejean, 1825)
References.Dyschiriusaeneus:Apfelbeck,1904:72(VelipojanearLezhë);Apfelbeck,1907a:495(Buenë
River); Müller, 1922b: 76; Hieke & Wrase, 1988: 31. Dyschirius aeneus aeneus: Balkenohl, 2003: 224.

48. COLEOPTERA: CARABIDAE: SCARITINAE 47
Material studied. “Durrazo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Durres” /
“Albania leg. Bischoff 1932”, 1 s. (MNHUB).
19.6. Dyschirius apicalis Putzeys, 1846
References. Dyschirius apicalis: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 77
(Durrës); Hieke & Wrase, 1988: 31; Balkenohl, 2003: 224.
Material studied. “Alban. Wgth. Valona”, 1 s. (HNHM); “Skutari Bojana” / “Mündg. Apf. 08.”,
3 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Liqen Butrintës” /
“Albania leg. Bischoff 1936”, 5 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).
[Dyschirius auriculatus (Wollaston, 1867)]
Notes. An expected species for Albania due to its occurring in Montenegro (P. Bulirsch,
personal communication)
19.7. Dyschirius bacillus Schaum, 1857
19.7.1. Dyschirius bacillus arbensis J. Müller, 1911
References. Dyschiriodes bacillus arbensis: Fedorenko, 1996: 191 (Vlorë); Balkenohl, 2003: 224.
Dyschirius bacillus bacillus: Balkenohl, 2003: 224.
Material studied. Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).
Notes. According to Fedorenko (1996: 191) the specimen from Vlorë shows transitional
characters between the subspecies arbensis and lucanus (Focarile, 1959). P. Bulirsch (personal
communication) identified the specimen from Shkiela and referred it to spp. arbensis.
19.8. Dyschirius cylindricus (Dejean, 1825)
19.8.1. Dyschirius cylindricus hauseri A. Fleischer, 1898
References. Dyschirius cylindricus: Apfelbeck, 1907a: 496 (Buenë River Outfall).
Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” /
“Albania leg. Bischoff 1932”, 21 s. (MNHUB).
Notes. Balkenohl (2003: 225) has omitted the species for Albania. Here it is confirmed for the country.
19.9. Dyschirius laeviusculus Putzeys, 1846
References. Dyschirius laeviusculus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës).
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).
Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here D. laeviusculus is
confirmed for the country. The locality Kula e Lumës outlines the southern limit of the
species distribution in the Balkan Peninsula.
19.10. Dyschirius luticola Chaudoir, 1850
19.10.1. Dyschirius luticola luticola Chaudoir, 1850
Material studied. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania.

49. 48 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
19.11. Dyschirius macroderus Chaudoir, 1850
19.11.1. Dyschirius macroderus macroderus Chaudoir, 1850
References. Dyschirius macroderus: Apfelbeck, 1907a: 496 (Buenë River Outfall).
Notes. Balkenohl (2003: 226) has omitted the species for Albania. Actually D. macroderus needs
further confirmation for the country.
19.12. Dyschirius minutus (Dejean, 1825)
19.12.1. Dyschirius minutus albanicus J. Müller, 1922
References. Dyschirius minutus albanicus: Müller, 1922b: 48, 82-83 (Berat, loc. typ.); Müller, 1937:
122; Focarile, 1959: 51; Guéorguiev & Guéorguiev, 1995: 71; Guéorguiev et al., 1997: 9. Dyschirius
punctatus albanicus: Müller, 1923: 107 (Osum River near Berat, loc. typ.); Drovenik & Peks, 1994:
25 (Berat); Drovenik & Peks, 1999: 29 (Berat). Dyschirius minutus: Balkenohl, 2003: 226.
Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura-Erzen”
/ “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff
1936”, 2 s. (MNHUB).
Notes. Müller published two descriptions of the subspecies albanicus. First he described D.
minutus albanicus (Müller, 1922), and next D. punctatus albanicus (Müller, 1923). Drovenik and
Peks (Drovenik & Peks, 1994; Drovenik & Peks, 1999) erroneously indicated the locality
Berat for Montenegro. Fedorenko (2001) confirmed the subspecific status of albanicus and his
point of view is followed here.
19.13. Dyschirius morio Putzeys, 1866 [= simulator J. Müller, 1922]
References. Dyschirius simulator: Müller, 1922b: 48, 90 (Berat, loc. typ.; Shkodër, loc. typ.);
Müller, 1923: 105-107 (Osum River near Berat, loc. typ.); Ieniștea, 1979: 220 (Berat); Hieke &
Wrase, 1988: 32. Dyschirius morio: Guéorguiev & Guéorguiev, 1995: 66; Balkenohl, 2003: 226.
Material studied. “Berat” / “Albania leg. Bischoff 1931”, 15 s. (MNHUB).
Notes. Müller published two description of D. simulator (Müller, 1922; Müller, 1923), the older
one is correctly treated as valid.
# Dyschirius punctatus (Dejean, 1825)
References. Dyschirius punctatus: Magistretti, 1965: 107; Balkenohl, 2003: 227.
Notes. A primarily West-Mediterranean species (Fedorenko, 1996: 161-162, Fig. 382) which
presence in Albania is highly doubtful. Likely the confusion about its presence there is due to
the view of Magistretti (1965: 106-107) who treated D. minutus as synonym of D. punctatus.
Species removed from the list of the Albanian Carabidae.
[Dyschirius pusillus (Dejean, 1825)]
Notes. An expected species for Albania as it was recently collected in North Greece (P. Bulirsch,
personal communication)

50. COLEOPTERA: CARABIDAE: SCARITINAE 49
Species group “chalybeus”
19.14. Dyschirius chalybeus Putzeys, 1846
19.14.1. Dyschirius chalybeus gibbifrons Apfelbeck, 1899
References. Dyschirius gibbifrons: Apfelbeck, 1899: 290; Focarile, 1959: 48 (Shkodër; Vlorë).
Dyschirius chalybeus gibbifrons: Balkenohl, 2003: 225.
Material studied. “Alban. Wgth. Valona”, 7 s. (HNHM; MNHUB); “Skutari l. See-Ufer”, 9 s.
(HNHM; MNHUB); “Skutari 29.3.16”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”,
1 s. (MNHUB); “Liqen Butrintës” / “Albania leg. Bischoff 1936”, 10 s. (MNHUB); Tiranë
Park, 22.II.1998, 1 s., from grasses along small sluggish stream, R. Ward leg. (CMNH).
19.15. Dyschirius salinus Schaum, 1843
19.15.1. Dyschirius salinus striatopunctatus Putzeys, 1846
References. Dyschirius salinus: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 72
(Durrës); Hieke & Wrase, 1988: 30. Dyschirius salinus striatopunctatus: Balkenohl, 2003: 227.
Material studied. “Berat” / “Albania leg. Bischoff 1931”, 14 s. (MNHUB); “Durrazo” /
“Albania leg. Bischoff 1931”, 21 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB): “Durrës” /
“Albania leg. Bischoff 1935”, 2 s. (MNHUB).
Subgenus Dyschirius s.str.
Notes. New subgenus for the fauna of Albania.
19.16. Dyschirius latipennis Seidlitz, 1867
Material included (studied P. Bulirsch). Labinoti-Fushe (Elbasan District), Shkumbinit River,
2.VII.2001, 4 s., Fencl & Moravec leg. (cPB).
Notes. New species for the fauna of Albania.
Subgenus Eudyschirius Fedorenko, 1996
Species group “globosus”
19.17. Dyschirius globosus (Herbst, 1784)
References. Dyschirius globosus: Balkenohl, 2003: 229.
Species group “lafertei”
19.18. Dyschirius gracilis (Heer, 1837)
19.18.1. Dyschirius gracilis gracilis (Heer, 1837) [= lafertei Putzeys, 1846]
References. Dyschirius lafertei: Apfelbeck, 1907a: 495 (Buenë River).

51. 50 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB).
Notes. Balkenohl (2003: 229) omitted this species for the Balkans in spite of at least three
localities from the region that were showed by Fedorenko (1996: 129, Fig. 355). P. Bulirsch
(personal communication) informed me that there is a reliable record of D. gracilis from
Northwestern Greece. The species is confirmed here for the fauna of Albania.
19.19. Dyschirius importunus Schaum, 1857
19.19.1. Dyschirius importunus importunus Schaum, 1857
References. Dyschirius importunus: Apfelbeck, 1904: 74 (Velipoja near Lezhë); Gryuntal, 1984a:
292; Hieke & Wrase, 1988: 33. Dyschirius importunus importunus: Balkenohl, 2003: 229.
Material studied. “Alban. Wgth. Valona”, 3 s. (HNHM).
Subgenus Paradyschirius Fedorenko, 1996
Species group “substriatus”
19.20. Dyschirius parallelus Motschulsky, 1844
19.20.1. Dyschirius parallelus ruficornis Putzeys, 1846
References. Dyschirius ruficornis: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 79
(Shkodër; Berat); Messner, 1964: 277; Hieke & Wrase, 1988: 31. Dyschiriodes parallelus ruficornis:
Fedorenko, 1996: 202 (Vijosë River, Tepelenë District). Dyschirius parallelus ruficornis: Balkenohl,
2003: 230.
Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Vojusa Tepelene”
/ “Albania leg. Bischoff 1936”, 5 s. (MNHUB).
19.21. Dyschirius substriatus (Duftschmid, 1812)
19.21.1. Dyschirius substriatus priscus J. Müller, 1922
References. Dyschirius substriatus priscus: Müller, 1922b: 80 (Berat); Horion, 1941: 104; Ieniștea,
1979: 221; Hieke et al., 1986: 184 (Kula e Lumës near Kukës); Balkenohl, 2003: 230. Dyschirius
substriatus: Hieke & Wrase, 1988: 32. Dyschiriodes substriatus: Fedorenko, 1996: 204.
Tribe SCARITINI Bonelli, 1810
Subtribe SCARITINA Bonelli, 1810
20. Genus Scarites Fabricius, 1775
Subgenus Parallelomorphus Motschulsky, 1849
20.1. Scarites laevigatus Fabricius, 1792
References. Scarites laevigatus: Apfelbeck, 1904: 67 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke
& Wrase, 1988: 27; Balkenohl, 2003: 232.

52. COLEOPTERA: CARABIDAE: TRECHINAE 51
Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Himara Juni”
/ “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Liqen i Butrintes Vivari” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); 2 km
N Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Shkodër, 12.VI.1993, 2 s.,
P. Beron & B. Petrov leg. (NMNHS).
20.2. Scarites terricola Bonelli, 1813
20.2.1. Scarites terricola terricola Bonelli, 1813
Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);
“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “16.VI.1958
leg. Xh. Murraj”, 1 s. (HNHM); Saranda District, Butrinti, 16.IV.1994, 1 s., S. Beshkov leg.
(NMNHS); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).
Notes. New species for the fauna of Albania.
Subgenus Scallophorites Motschulsky, 1858
20.3. Scarites buparius (Forster, 1771)
References. Scarites buparius: Hieke et al., 1986: 184 (Vlorë).
Material examined. “Vom Alb. Amerik Institut…” / “Zweifellos Alb. Konvenienc”, 1 s.
(MNHUB); “S Albanien Vlora, Strand p.7.58 leg. Grebenscikov”, 1 s. (DEI).
Additional material included (studied A. Vigna Taglianti). Tiranë, botanical garden, 6.VI.1998,
1 s., A. Paparistos leg. (cAVT).
Notes. Balkenohl (2003: 233) has omitted the species for Albania. Here it is confirmed for the country.
Subfamily APOTOMINAE LeConte, 1853
Tribe APOTOMINI LeConte, 1853
21. Genus Apotomus Illiger, 1807
21.1. Apotomus rufithorax Pecchioli, 1837
References. Apotomus rufithorax: Schatzmayr, 1943b: 99 (Mal i Krujes; Karbunara e Madhe);
Wrase, 1992: 93 (Durrës); Wrase, 2003a: 237.
Material studied. “Durres” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 1
m., 2 f., D. Antonelli leg. (cAVT).
21.2. Apotomus rufus (P. Rossi, 1790)
References. Apotomus rufus: Wrase, 1992: 77 (Lum i Tiranë).
Notes. Wrase (2003a: 237) has omitted the species for Albania, therefore it occurrence in the
country needs further confirmation.

53. 52 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subfamily TRECHINAE Bonelli, 1810
Tribe BEMBIDIINI Stephens, 1827
Subtribe BEMBIDIINA Stephens, 1827
22. Genus Asaphidion Gozis, 1886
Notes. Hartmann has revised the material studied under this genus from HNHM.
? Asaphidion cyanicorne (Pandellé, 1867)
? Asaphidion cyanicorne cyanicorne (Pandellé, 1867)
Material studied. “Korab” / “A. cyanicornis cyanicornis det. H. Schweiger” / “Fundort wahrsch.
falsch!”, 1 m. with extracted genitalia (NMW); “Korab, Alb. Bischoff 1935” / “A. cyanicornis
cyanicornis det. H. Schweiger”, 1 m. with extracted genitalia (NMW).
Notes. Schweiger noted on the label that this locality might be untrustworthy. For the time
being his remark is taken into consideration despite A. cyanicorne has been recorded for Croatia
and Bulgaria (Mařan, 1933: 89; Marggi et al., 2003: 242).
22.1. Asaphidion flavipes (Linnaeus, 1761)
References. Asaphidion flavipes: Apfelbeck, 1904: 79 (Velipoja near Lezhë); Schatzmayr, 1943b:
95 (Tërpan); Hieke & Wrase, 1988: 42.
Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Mamuras Alban. Kruja Dr.
J. Müller”, 5 s. (NMW); “Argirocastro Albania Weirather”, 2 s. (NMW); “Lumi Bençes Tepelene,
Alb. m.”, 1 s. (NMW); “Berat Alb. Dr. Müller’18”, 1 s. (NMW); “Elbasan Alb., Mader”, 16 s.
(HNHM; NMW); “Tomor-Buranj Juli 1932” / “Albania leg. Bischoff 1932”, 12 s. (MNHUB);
“Bojana Shkoder 1934”, 8 s. (NMW); “Gjinokastrë Umg. Albania 1934 Bischoff”, 1 ex. (NMW);
“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Berat” / “Albania leg. Bischoff 1936”, 5 s. (NMW; MNHUB); Saranda
District, Lyukove, 5.V.1994, 1 m., P. Stoev leg. (NMNHS); Përmeti District, Pelombasi,
2.VI.1994, 1 f., G. Blagoev leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995,
1 f., L. Penev leg. (NMNHS); Tiranë Park, 12-19.XI.1997, 1 f., R. Ward leg. (CMNH).
Notes. Marggi et al. (2003: 242) have omitted the species for Albania. Here A. flavipes is
confirmed for the country.
22.2. Asaphidion nebulosum (P. Rossi, 1792)
22.1.1. Asaphidion nebulosum balcanicum Netolitzky, 1918
References. Asaphidion caraboides: Apfelbeck, 1904: 78 (Durrës). Asaphidion caraboides balcanicus:
Netolitzky, 1918: 215 (Buenë River near Shkodër; Berat); Schatzmayr, 1943b: 95 (Ura Zezes);
Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë; Bizë near Shëngjergji); Hieke &
Wrase, 1988: 41; Sakalian & Guéorguiev, 1997: 46; Marggi et al., 2003: 242. Asaphidion caraboides
var. balcanicus: Csiki, 1940: 212 (Kula Ljums; Gjalicë Mt.). Bembidion elegans: Murraj, 1968: 33
(Tomor Mt.).

54. COLEOPTERA: CARABIDAE: TRECHINAE 53
Material studied. “Shkëlzën Alban.”, 3 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp.
Csiki 1917”, 14 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s.
(HNHM); “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 62 s. (NMW); “Skutari, Alb. Klenka”, 3
s. (NMW); “Vojusa – Shkoder 1934”, 1 s. (NMW); “Dajtit Alb. Bischoff 1935”, 8 s. (NMW);
“Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); Tomor Mt., 21.IX.1961, 2 m., 3 f., Xh.
Murraj leg. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).
Notes. Bonavita & Vigna Taglianti (2005) concluded that A. caraboides and A. nebulosum are
two distinct species, the first monotypic and living in Central Europe, the second polytypic
(with four subspecies) and living in South Europe and Anatolia. The record of Murraj (1968)
about Bembidion elegans is based on misidentification. Except the material mentioned above the
present author has seen further more than 40 s. in MNHUB, which remained unregistered.
22.3. Asaphidion rossii (Schaum, 1857)
References. Asaphidion rossii: Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë); Marggi
et al., 2003: 243.
Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tiranë” / “Albania leg. Bischoff
1931”, 2 s. (NMW; MNHUB); “Bushek Alb. Elbasan Bischoff 1932”, more than 45 s. (NMW;
MNHUB); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 2 s. (NMW; MNHUB); “Gjirakastrë
Ung. Albania 1937 Bischoff 1934”, 3 s. (NMW); “Dajtit, Albania Bischoff 1935”, 8 s. (NMW);
“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 11 s. (MNHUB); “Berat” / “Albania leg. Bischoff
1936”, 2 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Vojusa –
Tepelene 1936”, 4 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);
“Lem i Kuri Ura Mes” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania
leg. Bischoff 1939”, 2 s. (MNHUB).
22.4. Asaphidion stierlini (Heyden, 1880)
References. Asaphidion flavipes: Csiki, 1940: 212 (Kula Ljums).
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM).
Notes. New species for the fauna of Albania.
23. Genus Bembidion Latreille, 1802
Subgenus Bembidion s.str.
? Bembidion humerale Sturm, 1825
References. Bembidion humerale: Murraj, 1968: 33 (Tomor Mt.).
Notes. The record of Murraj (1968) might be doubtful because of some misidentifications
found in his collection. Unfortunately, the cited material cannot be found.
? Bembidion quadrimaculatum (Linnaeus, 1761)
? Bembidion quadrimaculatum quadrimaculatum (Linnaeus, 1761) [= quadriguttatum (Fabricius, 1775)
nec Pontoppidan, 1763]
References. Bembidion quadrimaculatum: Csiki, 1940: 213 (Kula Ljums). Bembidion quadriguttatum
(Fabricius): Csiki, 1940: 213 (Kula Ljums).

55. 54 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. In HNHM, I found single specimen identical with that cited by Csiki (1940) to belong
to B. quadripustulatum. The occurrence of B. quadrimaculatum in Albania is very probable due to
the distribution of this common species on the Balkans.
23.1. Bembidion quadripustulatum Audinet-Serville, 1821
23.1.1. Bembidion quadripustulatum quadripustulatum Audinet-Serville, 1821 [= quadriguttatum
(Olivier, 1795) nec Fabricius, 1775]
Material studied. “Skutari Bojana” / “Apfelbeck”, 1 s. (NMW); “Alban., Wgth. Valona” /
“Apfelbeck”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania
Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Skutari, Alb. Klenka”, 3 s.
(NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Bojana - Ufer b. Shkoder, Alb.” / “Alb. Winkler
1934”, 1 s. (NMW); “Durazzo, Alb. Bischoff”, 1 s. (NMW); Tiranë Park, 1.III.1998, 1 s., R.
Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
Subgenus Bembidionetolitzkya E. Strand, 1929
Notes. This is one of the most complex subgenera from the genus in the Balkans. Apparently,
the center of diversity of this subgenus in the Peninsula is situated in the northern half of
Albania and the adjacent territories of Montenegro and Kosovo. The taxonomical problems
with several yet not specified forms inhabiting the studied territory deserve special endeavour.
Species group “tibiale”
23.2. Bembidion rhodopense Apfelbeck, 1902
References. Bembidion rhodopense: Hieke & Wrase, 1988: 47 (Korab Mt.); Guéorguiev &
Guéorguiev, 1995: 106; Guéorguiev et al., 1997: 14; Sakalian & Guéorguiev, 1997: 46.
Material studied. “…Albanien”, 5 s. (NMW); “Shkëlzën Alb. Bischoff 1935”, 1 s. (NMW);
“Korab” / “Albania leg. Bischoff 1935”, 2 s. (NMW; MNHUB); “Korab” / “Albania leg.
Bischoff 1936”, 5 s. (NMW; MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 27 s. (NMW;
MNHUB).
Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for
the country.
23.3. Bembidion tibiale (Duftschmid, 1812)
References. Bembidion tibiale: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke & Wrase,
1988: 47.
Material studied. “Shkëlzën Alban.”, 1 s. (NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9
s. (MNHUB).
Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for
the country.

56. COLEOPTERA: CARABIDAE: TRECHINAE 55
Species group “complanatum-geniculatum”
23.4. Bembidion geniculatum Heer, 1837
23.4.1. Bembidion geniculatum geniculatum Heer, 1837
References. Bembidion geniculatum: Hieke & Wrase, 1988: 48.
Materialstudied.“Albania,Mts.GyalicaLums”/“Akad.Balk.Exp.Csiki1918”,1s.(HNHM);“Korab”
/ “Albania Bischoff 1935”, 1 s. (MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (NMW).
Notes. Marggi et al. (2003: 245) have omitted the species for Albania. Here B. geniculatum is
confirmed for the country with first detailed records.
23.5. Bembidion justinae Meschnigg, 1947
References. Bembidion justinae: Meschnigg, 1947: 94 (North Albanian Alps: Dedaj, 780 m, south of
Maja e Radohines Massif, near a little brook, loc. typ.); Turin, 1981: 81; Marggi et al., 2003: 245.
Notes. Finding and revising the type of B. justinae is needed to clarify its systematic position
and relationships with the related species.
? Bembidion relictum Apfelbeck, 1904
References. Bembidion relictum: Netolitzky, 1943a: 116; Hieke & Wrase, 1988: 47; Nitzu, 1995: 158.
Material studied. “Dr. Penther Prokletija” / “Sammlung Apfelbeck”, 1 f. (HNHM).
Notes. Hieke & Wrase (1988) and Nitzu (1995) noted this species for Albania. Obviously,
these authors have followed the record “Albanian (Drin, Orsk leg. Winkler)” (Netolitzky, 1943a:
116). Actually this locality is situated in Western Kosovo close to the border with Montenegro.
Due to lack of exact records B. relictum is still regarded as questionable for Albania; the material
studied above is without definite state attribution.
! Bembidion sp. (aff. complanatum Heer, 1937)
Material studied. “Jezerce K. Wgth.18”, 1 f. (NMW).
Notes. Netolitzky has identified the female above as B. complanatum Heer, 1837. I examined it
and think also that this specimen is externally most similar to B. complanatum. The female at
hand has pointed apex of elytra and a body-size of 6.6-6.7 mm. The specimen from Maja e
Jezerzë differs from B. justinae and B. relictum in the black antennomeres, excluding the reddish
scapus. The study of more male specimens, including such from the same locality, and their
comparison with specimens of B. complanatum, B. relictum and the type of B. justinae, could
establish the correct status of this form.
Species group “varicolor”
23.6. Bembidion conforme (Dejean, 1831)
References. Bembidion conforme: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Horion,
1941: 132.
Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); Leskovac
- Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 245) have omitted for Albania. Here it is confirmed for the country.

57. 56 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
23.7. Bembidion meschniggi Netolitzky, 1943
References. Bembidion meschniggi: Netolitzky, 1943a: 113 (Dedaj, loc. typ.); Turin, 1981: 82;
Marggi et al., 2003: 246.
Material studied. “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” / “meschniggi n.
sp. det. Netolitzky” / “Type” / “Genital - Nr 458 präpariert G. Müller 1978”, 1 syntype
(NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Paul Meyer” / “Cotypus det. Netolitzky”
/ “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus anno 1996 revidit Miloš
Fassati”, 1 syntype (NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” /
“Cotypus det. Netolitzky” / “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus
anno 1996 revidit Miloš Fassati”, 1 syntype (NMW).
Notes. I consider B. meschniggi being a distinct species and most probably a sister taxon of B.
atrocaeruleum (Stephens, 1828), but further study should differentiate the former from the latter
and to prove its species status. There are in Albania two places with the name Dedaj (? =
Markdedaj), situated not far from each other. The first place is north of the Drin River (Shkodër
District), at the southern slopes of the North Albanian Alps, at 1012 meters altitude, whereas
the second one is south of the Drin River (Pukë District), at 500 meters altitude. For the
present one cannot specify what of both the type locality of B. meschniggi is.
23.8. Bembidion varicolor (Fabricius, 1803)
23.8.1. Bembidion varicolor varicolor (Fabricius, 1803) [= tricolor (Fabricius, 1801)]
References. Bembidion tricolor: Apfelbeck, 1904: 87 (Velipoja); Hieke & Wrase, 1988: 46.
Material studied. “Shkëlzën Alban.”, 2 s. (NMW); “Shkëlzën Alb. Alfons Bischoff ”, 1 s.
(NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9 s. (MNHUB).
Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for
the country.
Species group “fasciolatum”
# Bembidion ascendens K. Daniel, 1902
References. Bembidion ascendens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi
et al., 2003: 245.
Notes. Sure records for this species from the Balkan Peninsula are known from Slovenia,
Croatia, and Western Bosnia Herzegovina. Since no reliable records are known eastwards of
the city of Sarajevo, B. ascendens is removed from the list of the Albanian ground beetles. Most
likely, the literature data for it refer to B. brunoi.
23.9. Bembidion brunoi (Bonavita, 2001)
References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 468 (Kiri River near Mesi); Hieke et
al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi et al., 2003: 245. Bembidion ascendens:
Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al., 2003: 245.
Material studied. “Skutari Mesi”, 1 f. (NMW); “Merdita Oroshi” / “Alban. spt. IV.V.’05”, 2
m., 4 f. (NMW); “Mesi Kiri Ufer” / “Albania leg. Bischoff 1931”, 1 m., 2 f. (MNHUB); 5 km
from Tepelenë, Drina River, 3.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS).

58. COLEOPTERA: CARABIDAE: TRECHINAE 57
Notes. In both the shape and the internal structure of the median lobe this species seems
most related to B. fasciolatum. On the other hand, in the wider margin of the pronotum and
bigger size of the body, it resembles B. ascendens.
23.10. Bembidion concoeruleum Netolitzky, 1943
References. Bembidion coeruleum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Hieke &
Wrase, 1988: 47. Bembidion concoeruleum: Netolitzky, 1943a: 114 (Oroshi; Shkodër; Dukati); Marggi et
al., 2003: 245. Bembidion coeruleum concoeruleum: Hieke et al., 1986: 185 (Borshi near Vlorë; Uji Ftohte,
south of Tepelenë). Ocydromus concoeruleus: Bonavita & Vigna Taglianti, 1993: 84.
Material studied. “Orosi (Ptr.) Alban. spt.”, 5 s. (HNHM; NMW); “Skutari Mesi” / “Apfelb.
Alban. spt. IV.V.’05”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s.
(NMW); “Prokletija”, 1 s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze,
Penther’14”, 1 s. (NMW); “Dukati”, 1 s. (NMW); “Skutari, Alb. Klenka”, 4 s. (NMW); “Skutari,
Alb. Mader”, 1 s. (NMW); “Lumi, Benges Tepelene, Alb. m.”, 2 s. (NMW); “Argirokastron
Albania mai 1929 Weirather”, 6 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona,
Bischoff V.1931”, 1 s. (NMW); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (MNHUB); Tepelenë
District, Vijosë River, 2.VI.1994, 2 m., G. Blagoev leg. (NMNHS).
Notes. All previously published data for B. coeruleum Audinet-Serville, 1821 from the Balkans
refer to this species (see also Bonavita & Vigna Taglianti, 1993).
# Bembidion fasciolatum (Duftschmid, 1812)
References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i
Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al.,
2003: 245.
Notes. I failed to find material of this species from the Balkan Peninsula in the collections,
excluding a single specimen from the locality “Travnik”. There are two places with this
name - one in Slovenia and one in Bosnia Herzegovina. B. fasciolatum is removed from the
list of the Albanian Carabidae. The records cited for it in the past most probably concern
either an unnamed species (the record near Oroshi) or B. brunoi (the records by Mesi River
and in South Albania).
! Bembidion sp. (ex. gruppo “fasciolatum”)
References. Bembidion fasciolatum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi);
Marggi et al., 2003: 245.
Material studied. “Orosi (Ptr.) Alban. spt.”, 6 s. (DEI; HNHM; MNHUB).
Notes. The specimens from DEI and HNHM have been identified as Bembidion merditanum
Apfelbeck as two of them from the first depository were incorrectly designated as syntypes
of this form. In fact, Bembidion merditanum Apfelbeck is nomen nudum. The series from Oroshi
may belong to separate species.
[subgenus Bracteon Bedel, 1879]
[Bembidion litorale (Olivier, 1790) [= elegans Germar, 1824]]
References. Bembidion elegans: Murraj, 1968: 33 (Tomor Mt.).
Notes. Murraj (1968) cited this species from the Tomor Mt. Among the material loaned from
MNST, I found five specimens from this locality bearing label data as the published ones and

59. 58 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
identified by the author as B. elegans. In fact, all the specimens concern Asaphidion nebulosum
balcanicum. For the present B. litorale is treated as an expected species for Albania because it
inhabits most of the Balkan countries.
Subgenus Chlorodium Motschulsky, 1864
23.11. Bembidion splendidum Sturm, 1825
23.11.1. Bembidion splendidum pincum De Monte, 1957
References. Bembidion splendidum: Apfelbeck, 1907a: 495 (Buenë River).
Material studied. “Scutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW); “Vojusa
Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg.
Bischoff 1937”, 3 s. (NMW).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for
the country. According to Bonavita and Vigna Taglianti (2005) the Balkan populations of the
species belong to ssp. pincum.
Subgenus Diplocampa Bedel, 1896
23.12. Bembidion assimile Gyllenhal, 1810
References. Bembidion assimile: Apfelbeck, 1904: 111 (Velipoja); Hieke & Wrase, 1988: 57.
Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for
the country.
23.13. Bembidion fumigatum (Duftschmid, 1812)
References. Bembidion fumigatum: Apfelbeck, 1904: 111 (Velipoja near Lezhë); Hieke & Wrase,
1988: 57.
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Actually, further
confirmation for the country is needed.
Subgenus Emphanes Motschulsky, 1850
23.14. Bembidion axillare (Motschulsky, 1844)
23.14. Bembidion axillare occiduum Margi & Huber, 2001 [= rivulare (Dejean, 1831)]
References. Bembidion rivulare: Hieke & Wrase, 1988: 58.
Material studied. “Durrës Alban.”, 1 s. (NMW); “Albanien Ruskuli 24.IX.18 Dr. H. Priesner”,
1 s. (NMW); “Ruskuli Alb., Mader”, 1 s. (NMW); “Durazzo, Alb. Ravasini, Lona”, 2 s. (HNHM);
“Durrazo, Alb. Bischoff ”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 9 s.
(MNHUB); “Valona, Alb. Bischoff 1936”, 2 s. (NMW); Radona, Shkiela, 21.VI.1994, 3 s., B.
Guéorguiev leg. (NMNHS).
Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here B. axillare is
confirmed for the country with first detailed records.

60. COLEOPTERA: CARABIDAE: TRECHINAE 59
23.15. Bembidion azurescens Dalla Torre, 1877
23.15.1. Bembidion azurescens azurescens Dalla Torre, 1877
References. Bembidion azurescens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Kula e Lumës
near Kukës); Hieke & Wrase, 1988: 58. Bembidion azurescens azurescens: Marggi et al., 2003: 248.
Material studied. “Oboti Schirel” / “Apfelbeck”, 1 s. (NMW); “Berat, Alb. Ravasini, Lona”,
3 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Vojusa Tepelene” /
“Albania leg. Bischoff 1936”, 8 s. (NMW).
23.16. Bembidion latiplaga Chaudoir, 1850
23.16.1. Bembidion latiplaga latiplaga Chaudoir, 1850
References. Bembidion latiplaga: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988: 59.
Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Oboti Schirei”
/ “Apfelbeck”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW); “Vojusa Tepelene” /
“Albania leg. Bischoff 1936”, 1 s. (NMW; MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff
1937”, 3 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb. Winkler 1937”, 1 s. (NMW); “Bojana-
Ufer b. Shkoder, Alb.” / “dch. Winkler XI 1942”, 1 s. (NMW); “Bojana Oboti, Alb.” / “ex coll.
Winkler 1948”, 1 s. (NMW); “Tirana Ungb. Albania” / “ex coll. Winkler 1948”, 2 s. (NMW).
Notes. Marggi et al. (2003: 249) have omitted the species for Albania. Here it is confirmed for
the country.
23.17. Bembidion normannum Dejean, 1831
23.17.1. Bembidion normannum mediterraneum Csiki, 1928 [= meridionale Ganglbauer, 1891]
References. Bembidion normannum meridionale: Apfelbeck, 1904: 103 (Velipoja near Lezhë).
Bembidion normannum apfelbecki: Müller-Motzfeld, 1986a: 262 (Raskuli); Hieke & Wrase, 1988:
59. Bembidion normannum mediterraneum: Hieke & Wrase, 1988: 59; Marggi et al., 2003: 249.
Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM).
Notes. Hieke & Wrase (1988) have noted that both subspecies of B. normannum inhabit Albania.
However, I consider the material studied and reference data as belonging to ssp. mediterraneum
(see also Marggi et al., 2003: 249).
23.18. Bembidion tenellum Erichson, 1837
23.18.1. Bembidion tenellum tenellum Erichson, 1837
References. Bembidion tenellum: Csiki, 1940: 213 (Kula Ljums).
Material studied. “Skutari Vraka” / “Apfelbeck”, 1 s. (NMW); “Oboti Schiren” / “Apfelbeck”,
1 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 7 s. (HNHM); “Albania
Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” / “Akad.
Balk. Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. Dr. J. Müller”, 3 s. (NMW); “Berat, Alb.
Ravasini, Lona”, 1 s. (NMW); “Liqen Butrintës Alb. 1936. Bischoff ”, 1 s. (NMW); “Bojana
Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb.
Winkler 1937”, 1 s. (NMW); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Tepelenë
District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 /
22.II.1998, 2 s., from grasses along small sluggish stream, R. Ward leg. (CMNH).

61. 60 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Marggi et al. (2003: 249) have omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Euperyphus Jeannel, 1941
23.19. Bembidion combustum Ménétries, 1832
23.19.1. Bembidion combustum combustum Ménétries, 1832
References. Bembidion combustum: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);
Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 48. Bembidion
combustum combustum: Marggi et al., 2003: 249.
Materialstudied.“Albania,Berat”,1s.(NMW);“MerditaOroschi” /“Apfelb.Alban.spt.IV.V.’05”,
9 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 11 s. (HNHM; MNHUB); “Berat, Alb. Ravasini, Lona”, 3 s.
(NMW); “Skutari, Alb. Klenka”, 1 s. (NMW); “Tiranë, Alb. Alf. Bischoff”, 2 s. (NMW); “Cucale
Geb.” / “Albania leg. Bischoff 1931”, 1 s. (NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”,
1 s. (MNHUB); “Lum i Tiranës Alb. V.1934 Mjr. Strupi”, 2 s. (NMW); “Lum Erzen Alb. V.1934
Mjr. Strupi”, 2 s. (NMW); “Tiranë” / “lgt. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “lgt.
Bischoff Albania 1935”, 2 s. (MNHUB); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST).
23.20. Bembidion scapulare Dejean, 1831
23.20.1. Bembidion scapulare oblongum Dejean, 1831
References. Bembidion oblongum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi).
Bembidion scapulare oblongum: Netolitzky, 1943b: 17; Hieke & Wrase, 1988: 50.
Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); “Berat Alb.
Dr. J. Müller’18”, 1 s. (NMW); “Elbasan, Alb. 4.IX.18. Dr. H. Priesner”, 1 s. (NMW); “Elbasan
Alb., Mader”, 3 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s.
(NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomorica” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 2 s. (NMW;
MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 250) have omitted the species for Albania. Here it is confirmed for
the country. Presently the subspecific status of the Balkan populations remains unsettled.
23.21. Bembidion testaceum (Duftschmid, 1812)
23.21.1. Bembidion testaceum testaceum (Duftschmid, 1812)
Material studied. “Berat, Alb. Ravasini, Lona”, 1 s. (NMW); “Bojana-Schkodër Alb., Bischoff
1934”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
Subgenus Metallina Motschulsky, 1850
23.22. Bembidion lampros (Herbst, 1784)
References. Bembidion lampros: Apfelbeck, 1904: 83 (Velipoja; Vlorë); Apfelbeck, 1909: 297
(Vlorë; Sop); Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 44; Marggi et al., 2003: 251.

62. COLEOPTERA: CARABIDAE: TRECHINAE 61
Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
North Albanian Alps, Bogë, 1000-1100 m, 30.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);
North Albanian Alps, Bogë, 1600 m, near cave B 52, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS);
Tiranë Park, 22-31.X.1997 / 1-9.IX.1997 / 8.II.1998 / 11.II.1998 / 15.II.1998 / 22.II.1998, 9
s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Except the specimens mentioned above, I have seen additionally nearly 100 s. in NMW
and MNHUB, which remained unregistered.
23.23. Bembidion properans (Stephens, 1828)
References. Bembidion lampros var. properans: Csiki, 1940: 212 (Kula Ljums). Bembidion properans:
Hieke & Wrase, 1988: 44.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan
Alb., Mader”, 1 s. (HNHM); “Lumi i Tiranës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff
1935”, 4 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB);
Tiranë Park, 8.II.1998, 2 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here it is confirmed for
the country. Except the specimens mentioned above, in NMW the present author has seen
other specimens, which remained not registered.
Subgenus Microserrullula Netolitzky, 1921
23.24. Bembidion quadricolle (Motschulsky, 1844) [= inserticeps Chaudoir, 1850; cyaneomi-
cans Mařan, 1933]
References. Bembidion inserticeps: Apfelbeck, 1907a: 495 (Buenë River). Bembidion quadricolle:
Hieke & Wrase, 1988: 42; Nitzu, 2003: 180.
Material studied. “Albania Scutari”, 1 s. (MNHUB); “Skutari Bojana”, 2 s. (HNHM); “Skutari
Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW; MNHUB); “Berat, Alb. Ravasini, Lona”,
1 s. (NMW); “Berat, Alb. Ravasini, Lona” / “cyaneomicans det. Paul Meyer”, 1 s. (NMW); “Berat”
/ “Albania leg. Bischoff 1931”, 1 s. (NMW); “Lumi Berati Alban. mer.” / “Winkler, Lona,
Bischoff, V.1931”, 1 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 5 s. (NMW;
MNHUB); “Ura Erzen” / “Albania leg. Bischoff. 1932”, 2 s. (NMW); “Bojana Shkoder” /
“Albania leg. Bischoff 1934”, 2 s. (NMW).
Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here B. quadricolle is
confirmed for the country.
Subgenus Nepha Motschulsky, 1864
23.25. Bembidion callosum Küster, 1847
23.25.1. Bembidion callosum callosum Küster, 1847 [= laterale Dejean, 1831; pseudolaterale Netolitzky,
1943a: 103]
References. Bembidion laterale: Apfelbeck, 1904: 108 (Velipoja). Bembidion laterale pseudolaterale:
Netolitzky, 1943a: 103.

63. 62 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Materialincluded(studiedP.Bonavita).“MajaeTartarit”/“Albanialeg.Bischoff 1936”,1f.(NMW).
Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for
the country.
23.26. Bembidion caucasicum (Motschulsky, 1844) [= substriatum Chaudoir, 1846]
References. Bembidion substriatum: Apfelbeck, 1907a: 501 (Merdita); Csiki, 1940: 213 (Shkëlzen
Mt.; Gjalicë Mt.). Bembidion caucasicum: Müller, 1937: 124; Hieke & Wrase, 1988: 56.
Material studied. “Merdita Munela”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk.
Exp. Csiki 1917”, 4 s. (HNHM); “Albania, Mts Gyalicalums” / “Akad. Balk. Exp. Csiki 1917”,
1 s. (HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Tomor:
Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Gjallica, Alb. Bischoff”,
1 s. (NMW); “Maja e Jezercë Albania Bischoff leg.”, 2 s. (NMW); “Korab, Guri Kua Alban.
Bischoff 35”, 5 s. (NMW); “Shkëlzen, Alb. Bischoff 1935”, 1 s. (NMW); “Maja e Policanit” /
“Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (NMW); “Korab” / “Albania leg. Bischoff
1936”, 1 s. (NMW); “Shkëlzen, Alb. Bischoff 5.8.1936”, 3 s. (HNHM; NMW); North Albanian
Alps, Maj’e Radohimes, 2200-2400 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 2 f., L. Latella leg. (cAVT).
Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, in NMW and MNHUB I have seen other
numerous specimens from the massifs of the North Albanian Alps, Korab Mt., Koritnik Mt.,
Gjalicë Mt. and Maja e Policanit Mt.
23.27. Bembidion vseteckai Mařan, 1936
23.27.1. Bembidion vseteckai dissimile G. Müller, 1943
References. Bembidion genei illigeri: Müller, 1943: 13; Hieke et al., 1986: 185 (Uji Ftohte, south
of Tepelenë; Kula e Lumës near Kukës); Hieke & Wrase, 1988: 55. Bembidion illigeri: Marggi et
al., 2003: 253.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Elbasan,
Alb. Leop. Mader leg.”, 2 s. (NMW); “Kruja Alb. Leop. Mader”, 2 s. (NMW); “Terbaci Alban.
mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); Mal i Dajtit Mt., 6.V.1962, 6 s. (MNST).
Notes. Except the specimens studied above I have seen several additional ones in NMW,
which remained unregistered. Bonavita (2005), as well P. Bonavita and A. Vigna Taglianti
(personal communication), noticed that all records or literature data about B. genei illigeri
Netolitzky, 1914 from the southern parts of Balkan Peninsula concern B. vseteckai dissimile.
Subgenus Notaphemphanes Netolitzky, 1920
23.28. Bembidion ephippium (Marsham, 1802)
References. Bembidion ephippium: Hieke & Wrase, 1988: 46.
Material studied. “Valona, Albania”, 4 s. (NMW); “Durazzo Alb. Dr. J. Müller”, 5 s. (NMW);
“Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” / “Albania leg. Bischoff
1932”, 9 s. (MNHUB).

64. COLEOPTERA: CARABIDAE: TRECHINAE 63
Notes. Marggi et al. (2003: 253) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
Subgenus Notaphus Dejean, 1821
23.29. Bembidion varium (Olivier, 1795) [= rumelicum Apfelbeck, 1902; dyrrhachii De
Monte, 1947]
References. Bembidion varium: Apfelbeck, 1894: 547; Apfelbeck, 1904: 85 (Velipoja near Lezhë);
Hieke & Wrase, 1988: 46; Marggi et al., 2003: 254. Bembidion varium dyrrhachii: De Monte, 1947:
52 (Durrës, loc. typ.). Bembidion rumelicum: Guéorguiev & Guéorguiev, 1995: 100.
Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Oboti Schirei” / “Apfelbeck”, 1 s.
(NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 12 s. (MNHUB); “Durres” / “Albania leg.
Bischoff 1932”, 7 s. (MNHUB); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW);
Radona, Shkiela, 21.VI.1994, 4 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 22.II.1998, 30
s., from grasses along small sluggish stream, R. Ward leg. (CMNH).
Notes. Except the material studied above the present author has seen more specimens in
NMW, which remained unregistered.
Subgenus Ocydromus Clairville, 1806
23.30. Bembidion aetolicum Apfelbeck, 1901
23.30.1. Bembidion aetolicum aetolicum Apfelbeck, 1901
References. Bembidion aetolicum: Netolitzky, 1943b: 12 (Tomor Mt.). Bembidion aetolicum
aetolicum: Müller-Motzfeld, 1996b: 164 (Tomor Mt.: Buranj; Tomor Mt.: Kulmak); Marggi
et al., 2003: 254.
Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor Buranj Alb.
10.4.1932 Alf. Bischoff ”, 4 s. (NMW); “Barshë-i-Vjetër Wildbach-Quellen Bischoff
21.VII.1940”, 2 s. (NMW).
23.31. Bembidion decorum (Zenker in Panzer, 1799)
23.31.1. Bembidion decorum decorum (Zenker in Panzer, 1799)
References. Bembidion decorum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i Madh
River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near
Lura Mt.); Hieke & Wrase, 1988: 49. Bembidion decorum decorum: Müller-Motzfeld, 1996b: 153
(Shkodër; Mal i Dajtit Mt.; Shkëlzen Mt.; Bushek near Elbasan; Tiranë; Zeza-Brüche near
Kruja; Lumi i Bençes near Tepelenë; Tomor Mt.: Kulmak; Miloti; Gjirokastër; Prespa Lake;
Cafa Stams; Kiri River near Mesi); Marggi et al., 2003: 255.
Material studied. Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);
Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
Notes. Except the specimens studied above, I have seen several more ones in NMW, which
remained unregistered.

65. 64 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
23.32. Bembidion siculum Dejean, 1831
23.32.1. Bembidion siculum smyrnense Apfelbeck, 1904
References. Bembidion siculum: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576
m; Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Lurë e Vjetër near Lura Mt.); Hieke
& Wrase, 1988: 49. Bembidion siculum smyrnense: Müller-Motzfeld, 1996b: 170 (Shkodër; Maje
Shbogut; Himara; Llogora; Radavac; Sarapta; Lura Mt.: Lurë e Vjetër); Marggi et al., 2003: 256.
Material studied. “Merdita Oroschi”, 3 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 5 s. (HNHM);
“Winkler, Lona, Bischoff, V.1931” / “Terbaci Alban. mer.”, 1 s. (NMW); “Llogora” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Himara Juni” / “Albania leg. Bischoff 1932”, 2 s.
(MNHUB); Përmeti District, Pelombashi, 2.VI.1994, 3 s., G. Blagoev leg. (NMNHS).
Notes. Besides the material studied above, I have seen other more in NMW, which remained
unregistered.
Subgenus Ocyturanes Müller-Motzfeld, 1986
23.33. Bembidion balcanicum Apfelbeck, 1899
23.33.1. Bembidion balcanicum balcanicum Apfelbeck, 1899
References. Bembidion balcanicum: Müller, 1918b: 85 (Munella Mt.); Müller, 1926: 61 (Munella
Mt.); Csiki, 1940: 213 (Gjalicë Mt.; Korab Mt.); Horion, 1941: 136; Netolitzky, 1943a: 42
(Korab Mt.); Hieke & Wrase, 1988: 52. Bembidion balcanicum balcanicum: Marggi et al., 2003: 256.
Material studied. “Latif Koritnik”, 11 s. (HNHM; NMW); “Merdita Munela”, 1 s. (HNHM);
“Merdita Munela” / “Wgth 1906 Merdita”, 10 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” /
“Alban. montenegr. Grenze, Penther,’14”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk.
Exp. Csiki 1917”, 1 s. (HNHM); “Albania, Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”,
21 s. (HNHM); “Alban Exped. Gialica Ljums 17.-26.VI’18”, 2 s. (NMW); “Albania Mts. Korab”
/ “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Jezerce Albania Bischoff leg.”, 4 s.
(NMW); “Maja e Jezerce Alban. sept.” / “Alb. Winkler 1934”, 1 s. (NMW); “Albania Korab
1935 Bischoff leg.”, 1 s. (NMW); “Giallica, Alb. Bischoff 1935”, 3 s. (NMW); “Tomor - Tojk
Bischoff Alban.”, 3 s. (NMW); “Tomor - Tojk Bischoff 1935”, 3 s. (NMW); “Maja Policanit”
/ “Albania 1936 leg. Ramadan coll. Bischoff ”, 3 s. (NMW); “Shkelzën, Alb. Bischoff, 5.8.1936”
/ “Albania leg. Bischoff 1936”, 8 s. (HNHM; NMW).
Notes. Besides the material studied above, in NMW and MNHUB I have seen numerous
other specimens from the massifs of Shkëlzën Mt., Gjalicë Mt., Korab Mt. and Tomor Mt.,
which remained unregistered.
23.34. Bembidion gudenzii (Neri, 1981)
References. Bembidion gudenzii: Müller-Motzfeld, 1985: 225 (“Hochgebirge bei Valona (Epyrus)”).
Notes. Marggi et al. (2003: 256) have omitted B. gudenzii for Albania. Actually, the species
needs further confirmation for the country.
23.35. Bembidion pindicum Apfelbeck, 1901
References. Bembidion pindicum: Netolitzky, 1943a: 49 (Korab Mt.); Guéorguiev, 1999: 71.

66. COLEOPTERA: CARABIDAE: TRECHINAE 65
Material studied. “Korab, Guri-Kua Alban. Bischoff 35”, 1 s. (NMW); “Tomor-Tojk Alban
1935 Bischoff leg.” / “pindicum var.? dtt. Netolitzky”, 1 s. (NMW); “Tomor - Tojk Alban. 1935
Bischoff leg.”, 2 s. (NMW).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. The specimens studied
have been found to be conspecific with three syntypes of B. pindicum labelled “Apfelbeck
Peristeri” / “(Peryphus) pindicum typ. Apf.” / “Typus” (NMW), therefore the species is here
confirmed for Albania.
23.36. Bembidion praeustum Dejean, 1831
23.36.1. Bembidion praeustum praeustum Dejean, 1831
References. Bembidion praeustum: Hieke et al. 1986: 186 (Uji Ftohte, soth of Tepelenë); Hieke
& Wrase, 1988: 52. Bembidion praeustum praeustum: Marggi et al., 2003: 257.
Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Tomor: Kulmak Albania mer.” /
“Ravasini & Lona”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.” / “Alb. Winkler”, 1 s.
(NMW); “Bushek b. Elbasan, Alb. Alf. Bischoff”, 1 s. (NMW); “Maj’e Shbogut Alnania mer.” /
“Winkler, Lona, Bischoff, V.1931”, 2 s. (NMW); “Humara juni” / “Albania leg. Bischoff 1932”,
2 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB); “Maranaj, Alb. Bischoff,
1934”, 1 s. (NMW); “Lumi i Tiranës Alb. V.1934 Mjr. Strupi”, 1 s. (NMW); “Köleyrë” / “Albania
leg. Bischoff 1936”, 1 s. (NMW); “Boge ober d. Quelle” / “Albania leg. Mueller 1939”, 1 s.
(NMW); “Borshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 8 s. (HNHM; NMW);
“Borshë, Alb. Bischoff 1940”, 1 s. (NMW); “Tirane, Alb. dch. Winkler 42”, 1 s. (NMW); Tomor
Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G.
Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);
Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS).
Notes. A part of the material in NMW has been determined as B. praeustum viridifluum J. Müller,
1929. However, I refer provisionally all the specimens studied to the nominotypical race.
23.37. Bembidion signatipenne Jacquelin du Val, 1852
References. Bembidion signatipenne: Hieke & Wrase, 1988: 53.
Notes. Marggi et al. (2003: 257) have omitted B. signatipenne for Albania. Actually, the species
needs further confirmation for the country.
23.38. Bembidion stolfai J. Müller, 1943
References. Bembidion stolfai: Müller, 1943: 12 (North Albanian Alps: Okol near Bogë, loc.
typ.; Maja e Jezercë); Netolitzky, 1943a: 41 (North Albanian Alps: Okol near Bogë; Maja e
Jezercë); Marggi et al., 2003: 257.
Material studied. “Maja e Jezerce Albania sept.” / “Alb. Winkler 1934” / “coll. Netolitzky”
/ “stolfai m. n. sp. det. J. Müller” / “Co-typus” / “Coll. Netolitzky”, 1 f. syntype (NMW).
Notes. The syntype of B. stolfai was studied and compared with specimens of B. reiseri reiseri
Apfelbeck, 1902 from Bosnia Herzegovina (including a syntype labelled “reis. 1900 Cvrstica”
/ “Testediolum bosnicum typ. Apf.” / “Reiseri” / “Typus”), topotypical material of B. reiseri vranense
Apfelbeck, 1902 and extensive material of B. balcanicum balcanicum. B. stolfai is distinct from the
other two species in the pronotum wider than long being narrower in the anterior part and in

67. 66 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
the elytra having distinct transverse microsculpture. Besides, it differs from B. reiseri s.l. in the
wrinkles on outer basal fovea of pronotum clear and long (vs. reduced), the shoulders of
elytra distinct (vs. round) and the sides of elytra after middle parallel (vs. slightly round). B.
stolfai is also living sympatrically with B. balcanicum balcanicum in the massif of Maja e Jezercë.
Subgenus Odontium LeConte, 1848
23.39. Bembidion foraminosum Sturm, 1825
References. Bembidion foraminosum: Hieke & Wrase, 1988: 43.
Material studied. “Tiranë” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for
the country with first detailed record.
23.40. Bembidion striatum (Fabricius, 1792)
References. Bembidion striatum: Netolitzky, 1918: Beilage zu Heft 4-6 (Shkodër; Oboti);
Geisthardt, 1975: 8. Bembidion striatul sic!: Panin, 1942: 567 (Shkodër).
Material studied. “Oboti Schirei”, 1 s. (NMW); “Skutari Bojana”, 5 s. (NMW); “Skutari
Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW).
Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Omoperyphus Netolitzky, 1931
23.41. Bembidion hypocrita Dejean, 1831
23.41.1. Bembidion hypocrita illyricum Netolitzky, 1918
References. Bembidion hypocrita illyricum: Müller, 1918b: 89; Müller, 1926: 66; Netolitzky, 1943a: 122.
Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Mader”,
1 s. (NMW); “Argyrókastron, Alb. leg. Weirather”, 1 s. (NMW); “Mal i Dajtit” / “Albania leg.
Bischoff 1935”, 3 s. (NMW; MNHUB); “Shkelzën, Alb. Bischoff, 5.8.1936”, 1 s. (NMW);
“Bischoff 21.VII.1940 Börshë-i-Vjetër Wildbach-Quellen”, 7 s. (NMW); “Börshë-i-Vjetër Alb.
21.7.1940 Bischoff”, 1 s. (MNHUB).
Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
Subgenus Peryphanes Jeannel, 1941
23.42. Bembidion brunnicorne Dejean, 1831
23.42.1. Bembidion brunnicorne brunnicorne Dejean, 1831
References. Bembidion brunnicorne: Netolitzky, 1943a: 50; Csiki, 1940: 213 (Kula Ljums).
Material studied. “Albania Kula Ljums”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk.
Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Kruja,
Alban., Mader”, 1 s. (NMW); “Tomor: Kulmak Albania mer.”, 3 s. (NMW); “Mal i Dajtit” /

68. COLEOPTERA: CARABIDAE: TRECHINAE 67
“Albania leg. Bischoff 1935”, 2 s. (HNHM; MNHUB); “Maranaj 1937” / “Albania bei Skutari”,
1 s. (NMW); “Börshë i Vjetër Wildbach-Quellen Bischoff, 21.VII.1940”, 4 s. (NMW).
Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material listed above, I have seen additionally more than 70 s. from
different localities in MNHUB, which remained unregistered. All the material in the last museum
was identified as B. milleri.
23.43. Bembidion dalmatinum Dejean, 1831
23.43.1. Bembidion dalmatinum dalmatinum Dejean, 1831
References. Bembidion dalmaticum sic!: Anonymous, 1907: 286. Bembidion dalmatinum: Apfelbeck,
1907a: 498 (Fani i Madh River near Oroshi); Csiki, 1940: 213 (Gjalicë Mt.); Schatzmayr, 1943b:
95 (Dukati; Lushnje); Hieke & Wrase, 1988: 53. Chlaenius nigricornis: Murraj, 1968: 33 (Tiranë;
Tomor Mt.). Bembidion dalmatinum dalmatinum: Marggi et al., 2003: 258.
Material studied. Tomor Mt., 21.IX.1961, 9 s., Xh. Murraj leg. (MNST); Theth, 800-1200 m,
28.V.1993, 7 s., P. Beron & B. Petrov leg. (cPN; NMNHS); North Albanian Alps, Bogë, 1400-
1800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë,
1000-1100 m, 5-9.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District, Merkurth,
11.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); Përmeti District, Pelombashi, 2.VI.1994,
1 s., G. Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 27 s., G. Blagoev
leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS);
Leskoec - Barmashi, 880 m, 2.VI.1995, 2 s., G. Blagoev leg. (NMNHS); 5 km Tepelenë, Drina
River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Ljukove near Vlorë, 6.VI.1995, 4 s., G.
Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH).
Notes. About the record of Murraj (1968) above see the notes under Chlaenius nigricornis.
Besides the material studied, the present author has seen numerous other specimens in NMW
and MNHUB, which remained unregistered.
23.44. Bembidion deletum Audinet-Serville, 1821
23.44.1. Bembidion deletum deletum Audinet-Serville, 1821
Material studied. “Borshë, Alb. Bischoff 1940”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
23.45. Bembidion grandipenne Schaum, 1862
23.45.1. Bembidion grandipenne grandipenne Schaum, 1862
References. Bembidion grandipenne: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);
Hieke & Wrase, 1988: 54.
Material studied. “Oroschi (Ptr.) Alban. spt.”, 4 s. (HNHM; MNHUB); “Merdita Oroschi” /
“Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Albania Malčija” / “Akad. Balk. Exp. Csiki 1917”,
1 s. (HNHM); “Skutari Alb., Mader”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.”, 2 s.
(NMW); “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff
1932”, 2 s. (NMW; MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Shkëlzën, Alb. Bischoff 1935”, 1 s. (NMW); “Borshë-i-Vjetër Wildbach-Quellen
Bischoff 21.VII.1940”, 4 s. (NMW; MNHUB); “Borshë, Alb. Bischoff 1940”, 1 s. (NMW).

69. 68 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Marggi et al. (2003: 259) have omitted the species for Albania. Here it is confirmed for
the country.
# Bembidion milleri Jacquelin du Val, 1852
# Bembidion milleri milleri Jacquelin du Val, 1852
References. Bembidion milleri: Hieke & Wrase, 1988: 54.
Notes. According to Müller-Motzfeld (personal communication in 1994) B. milleri does not
live in the Balkans. P. Bonavita and A. Vigna Taglianti (personal communication) stated that it
occurs only in Slovenia and probably in Serbia. Here the upper opinions are followed, though
Marggi et al. (2003: 258-259) noted the species from the Balkan Peninsula.
Subgenus Peryphus Dejean, 1821
23.46. Bembidion bualei Jacquelin du Val, 1852
23.46.1. Bembidion bualei bualei Jacquelin du Val, 1852 [= albanicum J. Müller, 1935]
References. Bembidion andreae: Apfelbeck, 1904: 92 (Durrës). Bembidion andreae albanicum: Müller,
1935: 152 (Berat, loc. typ.; Tiranë; Shkodër); Netolitzky, 1943a: 27 (Berat); Hieke et al. 1986:
186 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 51; Guéorguiev & Guéorguiev,
1995: 111; Guéorguiev et al., 1997: 15. Bembidion andreae bualei: Panin, 1942: 582 (Berat);
Schatzmayr, 1943b: 95 (Ura Zezes). Bembidion cruciatum albanicum: Marggi et al., 2003: 260.
Material studied. “Alban. Exped. Gjalica Ljuma 17.-26.VI.’18”, 6 s. (NMW); “Bojana Shkodër
Alb., Bischoff 1937” / “Albania leg. Bischoff 1937”, 1 s. (HNHM).
Notes. According to a recent study (Coulon, 2006) the Balkan populations of the species
related to B. cruciatum Dejean, 1831 belong to B. bualei bualei, and the form albanicum is a synonym
of the latter. Except the material studied above, I have seen more specimens from different
localities in NMW and MNHUB, which remained unregistered.
23.47. Bembidion femoratum Sturm, 1825
23.47.1. Bembidion femoratum femoratum Sturm, 1825
Material studied. “Scutari, Alb. Klenka”, 1 s. (NMW); “Bojana, Alb. Shkodër Bischoff 1934”,
31 s. (NMW); “Albania leg. Bischoff 1937”, 1 s. (NMW); “Bojana Shkodër Albania leg. Bischoff
1937”, 4 s. (NMW); 5 km from Tepelenë, Drina River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS).
Notes. New species for the fauna of Albania.
23.48. Bembidion subcostatum (Motschuslky, 1850)
23.48.1. Bembidion subcostatum vau Netolitzky, 1913 [= javurkovae Fassati, 1944]
References. Bembidion subcostatum: Hieke & Wrase, 1988: 52. Bembidion subcostatum javurkovae: Hieke
et al. 1986: 186 (Borshi near Vlorë; Lurë e Vjetër near Lura Mt.); Marggi et al., 2003: 263.
Material studied. “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 4 s. (NMW); “Scutari Alb. Dr.
J. Müller”, 1 s. (NMW); “Berat, Alb. sch. Winkler”, 1 s. (NMW); “Tirana Alb. dch. Winkler 42”,
2 s. (NMW); “Bushek Juni” / “Berat Alban. mer.” / “Albania leg. Bischoff 1932”, 4 s. (NMW);
Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994,
1 s., B. Guéorguiev leg. (NMNHS).

70. COLEOPTERA: CARABIDAE: TRECHINAE 69
Notes. Netolitzky identified the specimens in NMW as ssp. vau, ssp. javurkovae Fassati, 1944 or
ssp. spartanum Fassati, 1944. However, in a forthcoming paper these three taxa are synonymized
(Guéorguiev & Lobo, in press). In HNHM I have seen numerous additional specimens mostly
from the localities “Korab” and “Kula Lums”, and in MNHUB - other more than 200 s., as all
those data remained unregistered.
23.49. Bembidion tetracolum Say, 1823
23.49.1. Bembidion tetracolum tetracolum Say, 1823
References. Bembidion ustulatum (Linnaeus) (sic!): Csiki, 1940: 213 (Kula Ljums; Korab Mt.);
Murraj, 1968: 33 (Tomor Mt.).
Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Merdita M. Scheit” / “Mali Scheit
Apfelbeck”, 1 s. (NMW); “Alban. Vermosa Penther”, 1 s. (NMW); “Kruja, Alban., Mader”, 4
s. (NMW); “Berat Alban. mer.”, 2 s. (NMW); “Albania leg. Bischoff 1936”, 2 s. (NMW);
“Kèlcyrë” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Schkoder” / “Albania leg.
Bischoff 1937”, 2 s. (NMW); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Ljukova,
100 m, 5.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 263) have omitted the species for Albania. Here it is confirmed
for the country. The record of Murraj (1968) is included here since the revision of one male
from the Tomor Mt., having the same label data as the published material, has been found
to be correct.
Subgenus Philochthus Stephens, 1828
23.50. Bembidion decolor Apfelbeck, 1911
References. Bembidion decolor: Apfelbeck, 1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916:
340-341 (Buenë River; Vlorë); Netolitzky, 1943a: 63 (Elbasan; Kruja); Hieke & Wrase, 1988:
62; Marggi et al., 2003: 263.
Material studied. “Alban. Wgth Valona” / “Coll. Apfelbeck”, 1 m. (HNHM); “Albania Kula
Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).
Notes. Apfelbeck published two descriptions of this species, the first in Latin and Serbo-
Croatian and second one in Latin and German (Apfelbeck, 1911; Apfelbeck, 1916). The older
one is correctly treated as valid.
23.51. Bembidion guttula (Fabricius, 1792)
23.51.1. Bembidion guttula guttula (Fabricius, 1792)
References. Bembidion guttula: Hieke & Wrase, 1988: 61.
Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Alb.
Winkler leg.”, 1 s. (NMW); “Maje Tartarit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records.

71. 70 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
23.52. Bembidion inoptatum Schaum, 1857
References. Bembidion inoptatum: Apfelbeck, 1904: 112 (Velipoja near Lezhë; Vlorë); Apfelbeck,
1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916: 341 (Buenë River; Vlorë); Csiki, 1940: 213
(Kula Ljums); Hieke & Wrase, 1988: 62.
Material studied. “Bojana Oboti, Alb.” / “ex coll. Winkler 1948”, 1 s. (NMW).
Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for
the country.
23.53. Bembidion iricolor Bedel, 1879
References. Bembidion iricolor: Apfelbeck, 1909: 297 (Vlorë).
Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Actually, it needs further
confirmation for the country.
23.54. Bembidion lunulatum (Geoffroy, 1785)
References. Bembidion lunulatum: Schatzmayr, 1943b: 96 (Stan-Karbunarë).
Material studied. “Elbasan Alb., Mader”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff
1931”, 2 s. (NMW); “Tirana, Alb., dch. Winkler 42”, 1 s. (NMW); Tiranë Park, 22.II.1998, 1 s.,
from grasses along small sluggish stream, R. Ward leg. (CMNH).
Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for
the country.
23.55. Bembidion mannerheimii C.R. Sahlberg, 1827
Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor: Kulmak Albania
mer.” / “lg. Winkler mai 1931”, 5 s. (NMW).
Notes. New species for the fauna of Albania. Netolitzky has identified the material from the
Balkans either as “Bembidion neresheimeri” or as “cfr. neresheimeri”. I determined the material
from Albania as B. mannerheimii. According to D.W. Wrase (personal communication) B.
neresheimeri J. Müller, 1929 is a distinct species that also may inhabit the Balkan Peninsula;
Marggi et al. (2003: 264) have cited the latter only for Slovenia.
Subgenus Phyla Motschulsky, 1844
23.56. Bembidion tethys Netolitzky, 1926
References. Bembidion tethys: Focarile, 1959: 58; Hieke & Wrase, 1988: 61.
Material studied. “Elbasan Alb. 3.IX.18 Dr. H. Priesner”, 4 s. (NMW; MNHUB); “Elbasan
Alb., Mader”, 2 s. (NMW); “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW); “Lum Erzen Alb.
V.1934 Mjr. Strupi”, 1 s. (NMW); Ljukove, 100 m, 5.VI.1995, 2 s., G. Blagoev leg. (NMNHS);
Tiranë Park, 1-9.XI.1997 / 23-28.II.1998, 3 s., R. Ward leg. (CMNH).
Notes. Marggi et al. (2003: 265) have omitted the species for Albania. Here it is confirmed for
the country with first detailed records for the country.

72. COLEOPTERA: CARABIDAE: TRECHINAE 71
Subgenus Princidium Motschulsky, 1864
23.57. Bembidion punctulatum Drapiez, 1820
23.57.1. Bembidion punctulatum punctulatum Drapiez, 1820
References. Bembidion punctulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke et al.,
1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 44.
Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Tiranës”
/ “Albania leg. Bischoff 1935”, 2 s. (MNHUB); Tepelenë District, Drina River, 3.VI.1995, 1 s.,
G. Blagoev leg. (NMNHS).
Notes. Marggi et al. (2003: 266) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, the present author has seen some more
specimens in NMW, which remained unregistered.
Subgenus Talanes Motschulsky, 1864
Notes. New subgenus for the fauna of Albania.
23.58. Bembidion subfasciatum Chaudoir, 1850
Material studied. “Bojana - Umg. Pulaj” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Ruskuli
Alb., Mader”, 1 s. (NMW); “Liqen Butrintes” / “leg. Bischoff. 1936”, 3 s. (NMW).
Notes. New species for the fauna of Albania.
Subgenus Terminophanes Müller-Motzfeld, 1998
Notes. New subgenus for the faunas of the Balkan Peninsula and Albania.
23.59. Bembidion terminale Heer, 1841
! Bembidion terminale ssp.
Materialstudied.“Albania,AlbanAlps,Okol,800m,15-20.VII.1996,leg.G.Holzinger”,1f.(HNHM).
Notes. New species for the faunas of the Balkan Peninsula and Albania. For the time being
the subspecific status of the local population remains unspecified. The specimen in question
has a somewhat different shape of the pronotum, compared with specimens from the Alps
and North Iran.
Subgenus Testedium Motschulsky, 1864
23.60. Bembidion bipunctatum (Linnaeus, 1761)
23.60.1. Bembidion bipunctatum nivale Heer, 1837
References. Bembidion bipunctatum var. pyritosum: Csiki, 1940: 212 (Shkëlzen Mt.; Gjalicë Mt.;
Korab Mt.). Bembidion bipunctatum: Hieke & Wrase, 1988: 45.
Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, more than 50 s.
(HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania Mts.

73. 72 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Korab” / “Akad. Balk. Exp. Csiki 1918”, 15 s. (HNHM); “Korab Preshk, Alb. Bischoff 1936”,
1 s. (HNHM).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 4 m., 1 f., L. Latella leg. (cAVT).
Notes. Marggi et al. (2003: 269) have omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, I have seen other numerous specimens from
different localities in NMW and MNHUB, which remained unregistered.
Subgenus Trepanes Motschulsky, 1864
23.61. Bembidion articulatum (Panzer, 1796)
References. Bembidion articulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Marggi et al.,
2003: 270.
Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW);
“Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania
leg. Bischoff 1937”, 3 s. (NMW).
23.62. Bembidion maculatum Dejean, 1831
23.62.1. Bembidion maculatum maculatum Dejean, 1831
Material studied. “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
23.63. Bembidion octomaculatum (Goeze, 1777)
References. Bembidion octomaculatum: Apfelbeck, 1904: 110 (Velipoja near Lezhë); Apfelbeck,
1907a: 496 (Kiri River near Mesi); Hieke & Wrase, 1988: 60.
Material studied. “Alban. Wgth. Valona” / “Apfelbeck”, 1 s. (NMW); “Fieri, Albania a Semeni,
Müller”, 4 s. (NMW); “Skutari, Alb. Prof. Dr. Müller”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.”
/ “Alb. Winker 1934”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW).
Notes. Marggi et al. (2003: 270) have omitted the species for Albania. Here it is confirmed for
the country.
24. Genus Ocys Stephens, 1829
24.1. Ocys harpaloides (Audinet-Serville, 1821)
References. Ocys harpaloides: Apfelbeck, 1904: 117 (Vlorë); Apfelbeck, 1909: 297 (Pasha Liman).
Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Scutari Alb., Mader”, 2 s. (NMW);
“Tiranë” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Borshë-i-Vjetër Wildbach-Quellen
Bischoff 21.VII.1940”, 1 s. (NMW).
Notes. Marggi et al. (2003: 272) have omitted the species for Albania. Here it is confirmed for
the country.

74. COLEOPTERA: CARABIDAE: TRECHINAE 73
25. Genus Sinechostictus Motschulsky, 1864
Subgenus Pseudolimnaeum Kraatz, 1888
Notes. New subgenus for the fauna of Albania.
25.1. Sinechostictus doderoi (Ganglbauer, 1891)
Material studied. “Tomor Buranj Alb. 10.7.1932 Alf. Bischoff ”, 1 s. (NMW); “Sckëlzën” /
“Albania Bischoff 1935”, 2 s. (NMW).
Notes. New species for the fauna of Albania.
Subgenus Sinechostictus s.str.
25.2. Sinechostictus tarsicus (Peyron, 1858)
References. Bembidion elongatum var. effluviorum: Csiki, 1940: 213 (Kula Ljums). Bembidion elongatum
Dejean: Hieke & Wrase, 1988: 56.
Material studied. “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania
Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 6 s. (HNHM); “Miloti Alban. Matzenauer”, 1 s.
(NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Lumi Berati Alban. mer.”, 1 s. (NMW); “Lumi
Beratit Alban. mer. Lona. Winkler.”, 1 s. (NMW); “Llogora” / “Albania leg. Bischoff 1932”, 8
s. (MNHUB); “Llogora, Alb. Alf. Bischoff 2.VI.1932”, 3 s. (NMW); “Llogora” / “Albania leg.
Bischoff 1935”, 1 s. (NMW).
Notes. Marggi et al. (2003: 267 sub Bembidion elongatum tarsicum) have omitted the species for
Albania. Here it is confirmed for the country. Bonavita & Vigna Taglianti (2005) concluded
that in the Balkan Peninsula is present only S. tarsicus, a distinct species, while S. elongatus
(Ganglbauer, 1891) is a West European species. S. effluviorus (Peyron, 1858) is a separate species
confined only to the territory of Asian Turkey.
25.3. Sinechostictus millerianus (Heyden, 1883)
Material studied. “Shkëlzen” / “Albania leg. Bischoff 1936”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
Subtribe TACHYINA Motschulsky, 1862
26. Genus Lymnastis Motschulsky, 1862
Notes. New genus for the fauna of Albania.
26.1 Lymnastis galilaeus Piochard de la Brûlerie, 1876
Material studied. “Alban. Wgth. Valona”, 2 s. (MNHUB); “Mysege” / “Albania leg. Bischoff
1937”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania.

75. 74 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
27. Genus Paratachys Casey, 1918
27.1. Paratachys bistriatus (Duftschmid, 1812)
References. Tachys bistriatus: Apfelbeck, 1904: 117 (Velipoja); Murraj, 1968: 33 (Vau i Dejës);
Hieke et al., 1986: 184 (Borshi near Vlorë); Hieke & Wrase, 1988: 62.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Mustajbeg Velipoja Albania”,
2 s. (HNHM); “Albania Miloti”, 3 s. (HNHM); “Miloti, Alb. Matzenauer”, 1 s. (HNHM);
“Elbasan Alb., Mader”, 7 s. (HNHM); “Lim i Tiranes” / “Albania leg. Bischoff 1932”, 2 s.
(MNHUB); “Burelle” / “Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Mursi-Kara Alibey” /
“Albania leg. Bischoff 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1934”, 1 s.
(MNHUB); “Durres” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Librazhd, Prenjas, 750
m, 7.V.1995, 1 s., L. Penev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 8.II.1998 / 28.II.1998,
54 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Kopecký (2003: 275) has omitted the species for Albania, so that it is here confirmed
for the country. The record of Murraj (1968) is included here with great reservation in view
of possible misidentification (it was found a male of Bembidion praeustum from the Tomor Mt.
determinated by Murraj as Tachys bistriatus).
27.2. Paratachys fulvicollis (Dejean, 1831)
References. Tachys fulvicollis: Apfelbeck, 1904: 118 (Velipoja near Lezhë); Hieke & Wrase,
1988: 63.
Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Tirane” / “Albania leg.
Bischoff 1932”, 2 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW);
Tiranë Park, 1.III.1998, 9 s., R. Ward leg. (CMNH).
Notes. Kopecký (2003: 275) has omitted the species for Albania. Here it is confirmed for the
country.
27.3. Paratachys micros (Fischer von Waldheim, 1828)
References. Tachys micros: Hieke & Wrase, 1988: 63.
Material studied. “Ibe Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. Kopecký (2003: 276) has omitted the species for Albania. Here P. micros is confirmed
for the country with first detailed record.
28. Genus Polyderis Motschulsky, 1862
Notes. New genus for the fauna of Albania.
28.1. Polyderis cardioderus (Chaudoir, 1850)
Material studied. Tiranë Park, 8.II.1998, 1 s., grass and soil around tree trunks, R. Ward leg.
(CMNH).
Notes. New species for the fauna of Albania.

76. COLEOPTERA: CARABIDAE: TRECHINAE 75
29. Genus Sphaerotachys J. Müller, 1926
29.1. Sphaerotachys haemorrhoidalis (Ponza, 1805)
References. Tachys haemorrhoidalis: Apfelbeck, 1904: 120 (Velipoja); Hieke et al., 1986: 184 (Uji
Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 65.
Material studied. “Bojana Shkoder”, 10 s. (NMW); “Medua, Alb. 10.5.914”, 2 s. (NMW);
“Elbasan. Alb. 29.VIII.18 Dr. H. Priesner”, 3 s. (MNHUB); “Berat Alb. Dr. Müller 18”, 2 s.
(NMW); “Skutari, Alb. Klenka”, 2 s. (NMW); “Elbasan Alb., Mader”, 7 s. (NMW); “Vorra,
Alban., Mader”, 5 s. (NMW); “Bushek Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Lumi i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg.
Bischoff 1933”, 3 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
Tiranë Park, 8.II.1998, 89 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
Notes. Kopecký (2003: 278) has omitted the species for Albania. Here it is confirmed for
the country.
30. Genus Tachys Dejean, 1821
30.1. Tachys scutellaris Stephens, 1828
References. Tachys scutellaris: Hieke & Wrase, 1988: 63.
Material studied. “Durazzo, Alb. Dr. J. Müller”, 9 s. (NMW); “Rusculi Alb., Mader”, 1 s.
(NMW); “Durres” / “Albania leg. Bischoff 1932”, 18 s. (MNHUB); “Humara Juni” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania leg. Bischoff
1934”, 5 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 6 s. (MNHUB); “Liqen i
Butrintes” / “Albania leg. Bischoff 1936”, more than 30 s. (NMW).
Notes. Kopecký (2003: 276) has omitted the species for Albania. Here it is confirmed for the
country with first detailed records.
31. Genus Tachyta Kirby, 1837
Subgenus Tachyta s.str.
31.1. Tachyta nana (Gyllenhal, 1810)
References. Tachyta nana: Apfelbeck, 1904: 121 (Velipoja); Hieke et al., 1986: 184 (Bizë near
Shëngjergji); Hieke & Wrase, 1988: 66.
Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Ricavac 1300 m 25.VI.-
2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Mamuras Alban. Kruja Dr.
J. Müller”, 1 s. (NMW); “Kap Rodeni Alb., leg. Mader”, 1 s. (NMW); “Kruja, Alban., Mader”,
5 s. (NMW); “Llogora” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB).
Notes. Kopecký (2003: 277) has omitted the species for Albania. Here it is confirmed for
the country.

77. 76 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
32. Genus Tachyura Motschulsky, 1862
Subgenus Tachyura s.str.
32.1. Tachyura diabrachys (Kolenati, 1845)
References. Tachys sexstriatus var. bisbimaculatus: Csiki, 1940: 214 (Kula Ljums). Tachys inaequalis:
Hieke et al., 1986: 184 (Borshi near Vlorë). Tachys diabrachys: Hieke & Wrase, 1988: 65.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM);
“Elbasan, Alb. 4.IX.18 Dr. H. Priesner”, 1 s. (MNHUB); “Ibe Erzen” / “Albania leg. Bischoff
1932”, 10 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë”
/ “leg. Bischoff Albania 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff
1936”, 5 s. (NMW); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW); Tepelenë
District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Librazhd, Prenjas, 750 m,
7.V.1995, 7 s., L. Penev leg. (NMNHS); Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev
leg. (NMNHS).
Notes. Kopecký (2003: 279) has omitted the species for Albania. Here it is confirmed for the
country. The most part of the literature data and older identifications of Tachys inaequalis
(Reitter, 1884, nec Kolenati, 1845), T. sexstriatus (Duftschmid, 1812) and T. bisbimaculatus
(Chevrolat, 1860) from the Balkans concern this species.
32.2. Tachyura parvula (Dejean, 1831)
References. Tachys parvulus: Hieke & Wrase, 1988: 64.
Material studied. “Elbasan Alb. 29.VII.18 Dr. H. Priesner”, 1 s. (NMW); “Elbasan Alb.
4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Klenka”, 3 s. (NMW); “Skutari Alb.,
Mader”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB).
Notes. Kopecký (2003: 279) has omitted the species for Albania. Here T. parvula is confirmed
for the country with first detailed records.
# Tachyura sexstriata (Duftschmid, 1812)
Notes. I have not seen any specimen from this species from the Balkan Peninsula, excluding
Crete. The older records for it from the region refer to T. diabrachys.
Tribe PATROBINI Kirby, 1837
33. Genus Deltomerus Motschulsky, 1850
Subgenus Deltomerus s.str.
? Deltomerus malissorum Apfelbeck, 1918
References. Deltomerus malissorum: Zamotailov, 2003: 283.
Material studied. Montenegro. “Vunsaj, Alban. leg. Plec, 1914” / “Deltomerus mallisorum Typus!
Apfelb.” / “Typus”, 1 m. syntype (NMW); “Wgth. 1918 Kom VIII.”, 1 m., 2 f. (NMW); “Zljeb
Pl. Kulina” / “Montenegro leg. Bischoff 1931”, 1 m., 3 f. (MNHUB).

78. COLEOPTERA: CARABIDAE: TRECHINAE 77
Notes. After the original description, only Zamotailov (2003: 283) noted this species for
Albania without any detailed records. However, the type locality, Vunšaj (in Albanian) or
Vusanje (in Serbo-Croatian) is situated in the Montenegrin section of the North Albanian
Alps very close to Albanian territory (Apfelbeck, 1918c: 160-161, 164-165). The massifs
of Kom Mt. and Zljeb Mt. also lie outside the investigated territory. Due to lack of any
reliable record from Albania D. malissorum is treated here as questionable. Very probably it
inhabits the country.
33.1. Deltomerus parumpunctatus Zamotailov, 1990
References. Deltomerus parumpunctatus: Zamotailov, 1990: 133, 135 (North Albanian Alps: “Čafa
Borit”, loc. typ.); Zamotailov, 1992: 56; Zamotailov, 2003: 283.
Material studied. “Wgth. 1918 Menina pl. N. alb. alp.” / “Deltomerus parumpunctatus m. paratype
V. N. Kurnakov det. 1970”, 1 m. designated by mistake as paratype (NMW); “Wgth. 1918
Menina pl.” / “Deltomerus parumpunctatus m. alotype V. N. Kurnakov det. 1970”, 1 f. designated
by mistake as paratype (NMW).
Notes. Zamotailov (1990: 135) have noted no paratypes in the description of D. parumpunctatus.
For this reason, the two specimens in NMW designated as paratypes should not be considered
as part of the type series.
Subgenus Paradeltomerus Apfelbeck, 1908
33.2. Deltomerus paradoxus Apfelbeck, 1908
Notes. Apfelbeck published three consecutive descriptions for both the subgenus and the
species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest ones are correctly
treated as valid.
? Deltomerus paradoxus paradoxus Apfelbeck, 1908
References. Deltomerus paradoxus: Zamotailov, 1992: 56. Deltomerus paradoxus paradoxus:
Zamotailov, 2003: 283.
Notes. Hieke (1981: 80) stated by mistake that D. paradoxus is described from Albania.
In reality, the nominotypical form was described from the alpine zone of the Shar
Planina Mts. (Apfelbeck, 1907c; Apfelbeck, 1908). Dinchev & Atanasov (1998) stated
that only 1 % of the territory of the last mountain massif is situated in Albania. Actually
all specimens of the nominotypical race in the collections originate from Macedonia or
the Kosovo Region of Serbia. Due to lack of reliable data the subspecies is treated as
doubtful for Albania.
33.2.1. Deltomerus paradoxus korabensis J. Müller, 1936
References. Deltomerus paradoxus korabensis: Müller, 1936: 108-110 (Korab Mt., loc. typ.; Bjeshka
Matroxh); Zamotailov, 2003: 283. Deltomerus paradoxus: Zamotailov, 1992: 56.
Material studies. Nearly 15 s. from Korab Mt. (MNHUB).

79. 78 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Tribe POGONINI Laporte, 1834
34. Genus Pogonistes Chaudoir, 1871
Notes. New genus for the fauna of Albania.
34.1. Pogonistes convexicollis Chaudoir, 1871
Material examined. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania. For the time being, the locality near Durrës is
the westernmost locality in the species range.
34.2. Pogonistes gracilis (Dejean, 1828)
Material examined. “Durres” / “Albania leg. Bischoff 1932”, 29 s. (MNHUB).
Notes. New species for the fauna of Albania. For the present, the locality near Durrës is one
of the easternmost finds in the species range.
35. Genus Pogonus Dejean, 1821
Subgenus Pogonus s.str.
35.1. Pogonus littoralis (Duftschmid, 1812)
References. Pogonus litoralis sic!: Apfelbeck, 1904: 149 (Velipoja); Hieke & Wrase, 1988: 67.
Material studied. “Alban. Wgth. Valona”, 2 s. (HNHM); “Latif Pulaj”, 1 s. (HNHM); “Latif
Pulaj” / “Latif Alban. sept.”, 5 s. (NMW).
Notes. Bousquet (2003a: 287) has omitted the species for Albania. Here it is confirmed for
the country. Besides the material studied above, in MNHUB I have seen more than 60 s. (most
of them come from the vicinity of Durrës), which remained unregistered.
? Pogonus luridipennis (Germar, 1823)
References. Pogonus luridipennis: Murraj, 1968: 33 (Kamëz).
Notes. The data of Murraj (1968) are doubtful because of possible misidentification (among
the materials loaned from MNST, I found a male of Bembidion combustum from the Tomor Mt.
which had been determinated by Murraj as P. luridipennis).
35.2. Pogonus riparius Dejean, 1828
References. Pogonus riparius: Apfelbeck, 1904: 149 (Velipoja near Lezhë; Shkodër); Hieke &
Wrase, 1988: 67; Bousquet, 2003a: 287.
Material studied. “Latif Pulaj”, 6 s. (HNHM); “Latif Pulaj” / “Latif Alban. sept.”, 7 s.
(NMW); “Merdita, Alb. Matzenauer”, 2 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”,
6 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB); “Liqen i Butrintes”
/ “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 3 s.
(MNHUB); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Valona” /
“Albania leg. Bischoff 1936”, 2 s. (MNHUB).

80. COLEOPTERA: CARABIDAE: TRECHINAE 79
Tribe TRECHINI Bonelli, 1810
Subtribe PERILEPTINA Sloane, 1903
36. Genus Perileptus Schaum, 1860
Subgenus Perileptus s.str.
36.1. Perileptus areolatus (Creutzer, 1799)
36.1.1. Perileptus areolatus areolatus (Creutzer, 1799)
References. Perileptus areolatus: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Csiki, 1940: 214
(Kula Ljums); Hieke & Wrase, 1988: 34. Perileptus areolatus areolatus: Moravec et al., 2003: 288.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM);
“Skutari Alb., Mader”, 1 s. (HNHM); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Leskovac - Barmashi,
880 m, 2.VI.1995, 4 s., G. Blagoev leg. (NMNHS).
Subtribe TRECHINA Bonelli, 1810
! genus Adriaphaenops Noesske, 1928
! Adriaphaenops sp.
References. Aphaenopsis sp.: Latella, 1990: 148 (North Albanian Alps: Mali i Thatë Mt., cave
“Shpella e Haxhisë”, 1720 m).
Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave
“Shpella e Haxise”, 8.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. According to Vigna Taglianti (personal communication) the female mentioned above
belongs to a hitherto undescribed species.
37. Genus Albanotrechus Casale & V. B. Guéorguiev, 1994
37.1. Albanotrechus beroni Casale & V. B. Guéorguiev, 1994
References. Albanotrechus beroni: Casale & Guéorguiev, 1994: 418-420 (cave “Spela Merkurth”
near to village of Merkurth, Rreshen District); Moravec et al., 2003: 290.
38. Genus Duvalius Delarouzée, 1859
[subgenus Bicharotrechus Bokor, 1922]
[species group “pilifer”]
[Duvalius droveniki Magrini, 1998]
Notes. This species was described from the locality “Hajla Prokletije” in the Montenegrin
part of the North Albanian Alps (Magrini, 1998), very close to the Albanian border. D. droveniki
might live in Albania too.

81. 80 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
# Duvalius maglicensis Winkler, 1933
References. Duvalius maglicensis: Turin, 1981: 112.
Notes. This species lives only in Bosnia Herzegovina and Montenegro (see also Moravec et
al., 2003: 299). Species deleted from the list of the Albanian ground beetles.
Subgenus Duvalius s.str.
Notes. The arrangement of the separate groups is in accordance with the works of Casale
(1983), Casale & Laneyrie (1983), Casale & Vigna Taglianti (1990) and Casale et al. (1996). The
specificity of the group of “fodori” is discussed in a coming paper (Guéorguiev, in preparation).
I. South Aegean groups
Species group “krueperi”
38.1. Duvalius didonnai Magrini, Vanni & Ingusio, 1996
References. Duvalius didonnai: Magrini et al., 1995a: 27-29 (cave “Shpella Lekpetes”, 1150 m, near to
Kurvelesh, loc. typ.; cave “Shpella e Kasarit”, 1100 m, near to Kurvelesh); Moravec et al., 2003: 302.
Species group “georgii”
38.2. Duvalius bischoffi Meschnigg, 1936
References. Duvalius bischoffi: Meschnigg, 1936: 48-49 (cave “Homberg-Höhle” at Maja e Tanasit
Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b:
99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 110; Casale, 1983: 249; Casale & Laneyrie,
1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of
Vlorë); Moravec et al., 2003: 301.
38.3. Duvalius georgii (J. Müller, 1922)
References. Duvalius georgi: J. Müller, 1922a: 87-88 (hot pole in Tomor Mt., loc. typ.); Jeannel,
1928: 567-570 (Tomor Mt.: Toik, Kulmak); Scheibel, 1937a: 18, 20 (Tomor Mt.); Schatzmayr,
1943b: 98 (Tomor Mt.: Toik, Kulmak); Casale, 1983: 249; Casale & Laneyrie, 1982:199; Genest
& Juberthie, 1994: 592 (hot pole and cave at Tomor Mt., east of Berat); Moravec et al., 2003: 303.
Material studied. “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona” / “ex Coll. Bokor”,
1 f. (HNHM); “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona”, 3 s. (NMW); “Tomor:
Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor-Buranj juli
1932” / “Albania leg. Bischoff 1932”, 3 s. (NMW; MNHUB).
38.4. Duvalius muelleri Winkler, 1933
References. Duvalius muelleri: Winkler: 1933a: 75-76 (Shen Elias Mt., cave “Stere gropa e Vasos”,
800 m, south of Vlorë, loc. typ.); Scheibel, 1937a: 18, 20 (Shen Elias Mt., south of Vlorë);
Schatzmayr, 1943b: 98 (Shen Elias Mt., cave “Stere gropa e Vasos”); Turin, 1981: 112; Casale,

82. COLEOPTERA: CARABIDAE: TRECHINAE 81
1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave at Shen Elias
Mt., south of Vlorë); Magrini et al., 1995b: 25 (Llogora Mt., cave “Stera Kullëxhive”, south of
Vlorë); Moravec et al., 2003: 304.
Material studied. “Stere gropa de Vasos” / “Reza e Kanalit, Alban. m.” / “Winkler, Lona,
Bischoff, V.1931”, 8 syntypes (NMW); “Shen-E’Lias Juni 1932” / “Albania leg. Bischoff 1932”,
2 s. (MNHUB); “Shen-Elias” / “leg. Bischoff Albania 1934”, 1 f. (HNHM); “Shen-E’Lias Juni
1932” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB).
Species group “fuchsi”
38.5. Duvalius fuchsi Scheibel, 1937
References. Duvalius fuchsi: Scheibel, 1937a: 15-18, 20 (nameless cave near Llogora, south of
Vlorë, loc. typ.); Turin, 1981: 111; Casale, 1983: 249; Casale & Laneyrie, 1982: 199; Genest &
Juberthie, 1994: 592 (cave at Llogora, south of Vlorë); Moravec et al., 2003: 303.
Material studied. “Höhle in der Llogora” / “Albania leg. Bischoff 1935”, 1 syntype (MNHUB);
“Höhle i.-d. Llgora” / “Bischoff mai 1935”, 4 syntypes (MNHUB).
37.6. Duvalius pominii Schatzmayr, 1943
References. Duvalius pominii: Schatzmayr, 1943b: 96-97, 99 (cave “Stere e Vogel” and cave “Stere
e Madhe” near Dukati, south of Vlorë, loc. typ.); Turin, 1981: 112; Genest & Juberthie, 1994:
592 (cave “Stere e Vogel” and cave “Stere e Madhe” near Dukati); Moravec et al., 2003: 305.
38.7. Duvalius regiszogui Meschnigg, 1936
References. Duvalius regiszogui: Meschnigg, 1936: 46-47 (cave “Homberg-Höhle” Maja e Tanasit
Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b:
99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 113; Casale, 1983: 249; Casale & Laneyrie,
1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of
Vlorë); Moravec et al., 2003: 305.
II. North Aegean groups
Species group “leonhardi”
38.8. Duvalius sturanyi (Apfelbeck, 1904)
38.8.1. Duvalius sturanyi stenocephalus (Apfelbeck, 1918)
References. Trechus stenocephalus: Apfelbeck, 1918c: 164 (North Albanian Alps, Vermosh, 1100
m, loc. typ.). Duvalius leonhardi sturanyi: Jeannel, 1928: 548 (North Albanian Alps: Vermosh,
1100 m). Duvalius sturanyi stenocephalus: Scheibel, 1937a: 18-20 (North Albanian Alps: Vermosh,
Menina Planina Mt.); Pretner, 1978: 130, 135, 148, 167-168 (Vermosh; Menina Planina Mt.);
Moravec et al., 2003: 305. Duvalius leonhardi stenocephalus: Mařan, 1939c: 60 (Vermosh). Duvalius
sturanyi: Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (North Albanian Alps:
Menina Planina Mt. near Vermosh).

83. 82 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14” / “Typus” /
“Trechus stenocephalus Typus! Apfelb.”, 1 f. syntype of Trechus stenocephalus (NMW).
Notes. Jeannel (1928: 582-584) synonymized D. stenocephalus with D. leonhardi sturanyi, but later
Scheibel (1937a: 19-20) proved that the former is a subspecies of D. sturanyi. Further studies are
neededtotestthestatusof D.sturanyi sturanyiandD.sturanyi stenocephalus,havinginmindthedifferent
forms of their aedeagus and copulatory pieces (Scheibel, 1937a). The same concerns the remote
populations of the former near Foča and from the Komovi Mt. as well as the status of D. sturanyi
matejkai Mařan, 1939 described from the Montenegrin section of the North Albanian Alps.
Species group “fodori”
Notes.Thetreatmentof thisspeciesgroupisinaccordancewithaseparateauthor’spaperinprepara-
tion. It treats Duvalius fodori Scheibel, 1937, D. strupii Scheibel, 1937 and D. vignai Casalei, 1983.
38.9. Duvalius strupii Scheibel, 1937
References. Duvalius strupii: Scheibel, 1937b: 282-283 (Korab Mt., 2000 m, loc. typ.); Schatzmayr,
1943b: 98 (Korab Mt.); Turin, 1981: 113; Casale & Laneyrie, 1982:191; Casale, 1983: 250
(Korab Mt.); Genest & Juberthie, 1994: 592 (Korab Mt.); Moravec et al., 2003: 305.
Materialstudied.“Korab”/“Albanialeg.Bischoff 1936”,17specimens,incl.12syntypes(MNHUB).
Subgenus Euduvalius Jeannel, 1929
Species group “gogalai”
38.10. Duvalius zhalovi B. V. Guéorguiev, 2005
References. Duvalius zhalovi: Guéorguiev, 2005: 33-40 (cave “Peshtnik”, 1100 m near Ostreni
i Madh, loc. typ.).
39. Genus Neotrechus J. Müller, 1913
Species group “dalmatinus”
39.1. Neotrechus lonae (J. Müller, 1914)
39.1.1. Neotrechus lonae lonae (J. Müller, 1914)
References. Trechus lonae: J. Müller, 1914: 1003-1005 (Çukall Mt.: cave “Scpedha n’Giuka
Kunors”, loc. typ.; Çukall Mt.: cave “Scpedha n’Ciuk tugit Vukai”, loc. typ.). Neotrechus lonae:
Winkler, 1926: 86, 96 (caves in Çukall Mt.); Scheibel, 1936: 229 (Çukall Mt.); Scheibel, 1937a:
20 (Çukall Mt.). Neotrechus lonai sic!: Jeannel, 1928: 296-297 (Çukall Mt.: cave “Scpedha n’
giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Schatzmayr, 1943b: 97 (Çukall Mt.:
cave “Scpedha n’ giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Casale & Laneyrie,
1983: 180; Genest & Juberthie, 1994: 592 (caves in Çukall Mt., Rapshë). Neotrechus lonai sic!
lonai sic!: Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall Mt.); Pretner, 1978: 147, 166
(Çukall Mt.). Neotrechus lonae lonae: Moravec et al., 2003: 315.

84. COLEOPTERA: CARABIDAE: TRECHINAE 83
Material studied. “Cukali Gebirge N-Albanien Lona leg. 1914”, 4 s. (NMW); “Scpedha n’Giuka
Zukali Albanien Lona Giuno 1914”, 1 s. (DEI); “Cukale Albania sept.” / “Bischoff 1930 coll.
A. Winkler”, 5 s. (NMW); “Cucale Geb. Höhle von Vukaj” / “Albania leg. Bischoff 1931”, 1 s.
(MNHUB); “Shpella Kodra Çorit” / “Cukale Geb.” / “leg. Bischoff Albania 1934”, 3 s.
(MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”, 8 s.
(MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 1 s.
(MNHUB); “Shpella Rasa-Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 4 s.
(MNHUB); “Shpella Çuka Vjit” / “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB);
“Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. N. lonae lonae is sympatric and syntopic with N. malissorum (J. Müller, 1914) in most of
the caves where they have been found. The report of this subspecies from the area of Rapshë
(Genest & Juberthie, 1994: 592) is erroneous since this region is populated by N. lonae atavus.
The nominotypical subspecies differs from both ssp. atavus and ssp. shkodrensis in the wider
and more rounded, almost not narrowed to apex, elytra.
39.1.2. Neotrechus lonae atavus J. Müller, 1934 [= bischoffi Meschnigg, 1936]
References. Neotrechus bischoffi: Meschnigg, 1936: 44-46 (North Albanian Alps: cave “Shpella
e lugstit, Hot-Brigje” near Rapshë, loc. typ.); Schatzmayr, 1943b: 98 (cave “Shpella e lugstit,
Hot-Brigje” near Rapshë); Casale & Laneyrie, 1983: 180. Neotrechus lonae atavus: Scheibel, 1936:
227-228 (cave “Shpella e lugstit, Hot Brgje” near Rapshë); Scheibel, 1937a: 20 (cave “Hot
Brigje”); Moravec et al., 2003: 315. Neotrechus lonai sic! atavus: Absolon & Mařan, 1947: 2;
Mařan, 1947: 11 (Rapshë); Pretner, 1978: 147, 166 (cave “Shpella e lugstit, Hot Brgje” near
Rapshë); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (cave “Shpella e Lugstit”,
cave “Hot Brigje” near Rapshë).
Material studied. “Shpella e Lugstit” / “Brigie Hot” / “leg. Bischoff Albania 1934”, 7 syntypes
of Neotrechus bischoffi (MNHUB); “Shpella e Lugstit” / “Brigie Hot” / “Albania leg. Bischoff
1935”, 11 syntypes of Neotrechus bischoffi (MNHUB).
Notes. Scheibel (1936) established the synonymy of N. bischoffi with N. lonae atavus. Regardless
of that, Schatzmayr (1943b) and Casale & Laneyrie (1983) listed the former as valid species.
The type localities of both N. lonae atavus and N. suturalis pretneri Scheibel, 1936 coincide and
are located in Montenegro very close to the Albanian border.
39.1.3. Neotrechus lonae shkodrensis Scheibel, 1936
References. Neotrechus lonae shkodrensis: Scheibel, 1936: 228-229 (cave “Shpella kufinit, Koplikut-
Hotit” near Ivanaj, north of Liqen i Shkodrës, loc. typ.); Scheibel, 1937a: 20 (Ivanaj); Moravec
et al., 2003: 315. Neotrechus lonai sic! shkodrensis: Absolon & Mařan, 1947: 2; Mařan, 1947: 11
(Ivanaj); Pretner, 1978: 147, 166 (cave “Shpella kufinit, Koplikut-Hotit” near Ivanaj); Casale &
Laneyrie, 1983: 180; Drovenik & Peks, 1994: 45; Genest & Juberthie, 1994: 592 (cave “Kufinit
et Kopikut, Hotit peés” near Ivanaj, north of Liqen i Shkodrës); Drovenik & Peks, 1999: 51.
Material studied. “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1935”, 4
syntypes (MNHUB); “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1937”, 4 s.
(MNHUB).
Notes. Drovenik & Peks (1994: 45) and Drovenik & Peks (1999: 51) indicated that the type
locality of this subspecies is situated in Montenegro. Actually it is located in Albania, very
close to the state border.

85. 84 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
39.2. Neotrechus malissorum (J. Müller, 1914)
References. Trechus malissorum: Müller, 1914: 1005 (Çukall Mt.: cave “Scpedha n’ kunor
Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ Ciuk tujit Vukai”, loc. typ.; Çukall Mt.: cave
“Scpedha e Bacit Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ rasst’ kunors”, loc. typ.;
Çukall Mt.: cave “Scpedha n’ giuka kunors”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs
n’ gur t’ barzodhs”, loc. typ.). Neotrechus dalmatinus malissorum: Winkler, 1926: 86, 96 (caves in
Çukall Mt.); Scheibel, 1937a: 20 (Çukall Mt.). Neotrechus malissorum: Jeannel, 1928: 290-291
(Çukall Mt.: cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave
“Scpedha e bacit Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”;
cave “Scpedha Nre Gegs n’ gur t’ barzodhs Zukali”); Schatzmayr, 1943b: 98 (Çukall Mt.:
cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave “Scpedha e bacit
Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”; cave “Scpedha
Nre Gegs n’ gur t’ barzodhs Zukali”); Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall
Mt.); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (Çukall Mt., cave “n’Kunor
Vukai”, cave “n’Ciuk tujit Vukai”, cave “Lacit Vukai”, cave “n’Rast’Kunors”, cave “n’Giuka
Kunors”); Moravec et al., 2003: 315.
Material studied. “Scpedha n’Giuka Zukali Albanien Lona Giuno 1914”, 6 s. (DEI); “Scpedha-
Nre Geg N’gur Bazzodha Zukali Albanien”, 1 s. (HNHM); “Scpedha-Nre Geg N’gur Barzodha
Zukali Albanien Lona Giugno 1914”, 4 s. (MNHUB); “Cukali Gebirge N-Albanien Lona leg.
1914”, 4 s. (NMW); “Albania m. Cucali C. Lona”, 2 s. (HNHM); “Cukali Albania sept.” / “leg.
Bischoff 1931”, 2 s. (NMW); “Shpellë Quku i Konors Vukaj Cucale” / “Albania leg. Bischoff
18.VI.31”, 2 s. (MNHUB); “Shpella Kodra Carit” / “Cukale Geb.” / “leg. Bischoff Albania
1934”, 2 s. (MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”,
24 s. (NMW; MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”,
3 s. (MNHUB); “Shpella Rasa Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 3 s.
(MNHUB); “Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-
Thatë Mt., cave “Shpella e Haxise”, 8.VII.1993, rests of 3 s., A. Gobetti leg. (cAVT).
39.3. Neotrechus suturalis (Schaufuss, 1864)
39.3.1. Neotrechus suturalis pentheri Winkler, 1926
References. Trechus suturalis: Apfelbeck, 1918c: 164 (North Albanian Alps: small cave near
Rapshë). Neotrechus dalmatinus suturalis var. pentheri: Winkler, 1926: 85-86, 96 (near Rapshë, loc.
typ.). Neotrechus dalmatinus pentheri: Scheibel, 1937a: 20 (Rapshë). Neotrechus suturalis pentheri:
Jeannel, 1928: 295 (North Albanian Alps: caves near Rapshë); Absolon & Mařan, 1947: 2;
Mařan, 1947: 11 (North Albanian Alps: Rapshë); Pretner, 1978: 146-147, 167 (Rapshë); Casale
& Laneyrie, 1983: 180; Drovenik & Peks, 1994: 46; Genest & Juberthie, 1994: 592 (North
Albanian Alps: caves at Rapshë); Drovenik & Peks, 1999: 53; Moravec et al., 2003: 316.
Material studied. “Alban. montenegr. Grenze, Penther’14” / “Höhle bei Rapsa, 727 m”, 15
syntypes of Trechus suturalis (DEI; MNHUB); “Albania Rapsa Dr. Grabowski”, 86 s. (HNHM);
“Höhlen bei Rapsa leg. Bischoff 1931”, 1 s. (DEI); “Höhlen bei Rapsa 20.VII.31” / “Albania
leg. Bischoff 1931”, 4 s. (MNHUB); “Rapsh” / “Albania leg. Bischoff 1935”, 40 s. (MNHUB).
Notes. Drovenik & Peks (1994: 46) and Drovenik & Peks (1999: 53) noted by mistake the
type locality in Montenegro. In fact, the village of Rapshë is situated in Albania.

86. COLEOPTERA: CARABIDAE: TRECHINAE 85
40. Genus Trechus Clairville, 1806
Subgenus Trechus s.str.
Species group “quadristiatus”
40.1. Trechus obtusus Erichson, 1837
40.1.1. Trechus obtusus obtusus Erichson, 1837
References. Trechus obtusus: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m);
Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9. Trechus obtusus obtusus: Jeannel, 1927: 309
(“Petrov planina”, 2000 m); Moravec et al., 2003: 337.
Material studies. “Latif Koritnik”, 1 s. (NMW); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
40.2. Trechus quadristriatus (Schrank, 1781)
References.Trechusquadristriatus:Apfelbeck,1907a:498(MaliShenjtMt.:topregion,1500m);Schatzmayr,
1943b: 96 (Lushnje; cave “Stere e Madhe” near Dukati); Hieke et al., 1986: 184 (Uji Ftohte, south of
Tepelenë); Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9; Moravec et al., 2003: 339.
Material studied. “Elbasan Alb., Mader”, 2 s. (HNHM); “Kruja, Alban., Mader”, 4 s. (HNHM);
“Sarisaltik, 1180 m Alb., leg. Mader”, 3 s. (HNHM); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997
/ 23-28.XI.1997, 47 s., R. Ward leg. (CMNH).
Notes. Except the specimens studied above, in MNHUB the present author has seen additionally
about 210 s., which remained unregistered.
Species group “babinjensis”
40.3. Trechus babinjensis Jeannel, 1927
References. Trechus babinjensis: Jeannel, 1927: 267-268 (North Albanian Alps, loc. typ.); Casale
& Laneyrie, 1983: 141; Moravec et al., 2003: 327.
40.4. Trechus valbonensis Jeannel, 1927
References. Trechus valbonensis: Jeannel, 1927: 268 (North Albanian Alps, loc. typ.); Casale &
Laneyrie, 1983: 141; Moravec et al., 2003: 343.
Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m,
11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Species group “subnotatus”
40.5. Trechus subnotatus Dejean, 1831
40.5.1. Trechus subnotatus subnotatus Dejean, 1831
References. Trechus subnotatus subnotatus: Apfelbeck, 1904: 127 (Vlorë); Jeannel, 1927: 446 (Kruja;
Tomor Mt.: Kulmak); Pawlowski, 1973: 232; Moravec et al., 2003: 341. Trechus subnotatus: Jeannel,

87. 86 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
1925: 23 (Tomor Mt.: Kulmak); Horion, 1941: 181; Schatzmayr, 1943b: 96 (Dukati; cave “Stere
e Madhe” near Dukati); Casale & Laneyrie, 1983: 147; Hieke & Wrase, 1988: 36; Nonveiller et
al., 1994: 9, 15; Guéorguiev & Guéorguiev, 1995: 75; Guéorguiev et al., 1997: 11; Sakalian &
Guéorguiev, 1997: 46.
Material studied. “Kruja, Alban., Mader”, 2 s. (HNHM).
Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 1
m., D. Antonelli leg. (cAVT).
Notes. In MNHUB I have seen also additional 39 s., which remained not registered.
40.5.2. Trechus subnotatus ljubetensis Apfelbeck, 1908
References. Trechus subnotatus ljubetensis: Moravec et al., 2003: 341.
Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Albania” / “ljubetensis Apf.
R. Jeannel”, 1 m. (NMW).
Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908;
Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. The identification
of the two specimens from Vlorë is doubtful since the ssp. ljubetnesis is normally treated as
local endemic of the Shar Planina Mts.
? species group “cardioderus”
? Trechus irenis Csiki, 1912 [= cardioderus balcanicus Jeannel, 1927]
References. Trechus cardioderus balcanicus: Tabaković-Tošić, 1988: 137.
Notes. Tabaković-Tošić (1988) noted the range of this taxon reaching Albania. Due to lack of
reliable records from the country the species is treated as questionable for Albania.
Species group “pulchellus”
40.6. Trechus merditanus Apfelbeck, 1906
References. Trechus merditanus: Apfelbeck, 1906: 1163 (Merdita: Mal i Shenjt Mt. and Zepë
Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Jeannel,
1927: 539-540 (Merdita: Mal i Shenjt Mt. and Zepë Mt.); Pawlowski, 1972: 312-314; Casale &
Laneyrie, 1983: 151; Moravec et al., 2003: 336.
Material studied. “Merdita M. Scheit” / “merditanus n. sp. Apfelb.”, 2 s. probably syntypes (NMW).
Notes. According to Jeannel (1927), T. merditanus is a vicarious species of T. priapus.
40.7. Trechus priapus K. Daniel, 1902
40.7.1. Trechus priapus priapus K. Daniel, 1902
References. Trechus priapus: Meixner, 1939: 309; Pawlowski, 1973: 249; Hieke & Wrase, 1988:
38; Nonveiller et al., 1994: 9; Guéorguiev & Guéorguiev, 1995: 77; Guéorguiev et al., 1997: 10.
Trechus priapus priapus: Jeannel, 1927: 537 (North Albanian Alps; Gjalicë Mt.); Moravec et al.,
2003: 338.
Material studied. “Alban. Exped. Gjalica Ljums 17.-26.VI’18”, 2 s. (NMW).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Notes. Except the two specimens above, in MNHUB I have seen additionally nearly 800 s.,
which remained unregistered.

88. COLEOPTERA: CARABIDAE: TRECHINAE 87
Species group “egreginus”
40.8. Trechus egregius Jeannel, 1927
References. Trechus egregius: Jeannel, 1927: 551-552 (North Albanian Alps, loc. typ.); Casale &
Laneyrie, 1983: 152; Moravec et al., 2003: 330.
Species group “tristis”
40.9. Trechus nigrinus Putzeys, 1847 [= tristis (Duftschmid, 1812)]
References. Trechus nigrinus: Apfelbeck, 1904: 129 (Vlorë); Apfelbeck, 1907a: 498-499 (Mal i Shenjt
Mt.: top region, 1500 m); Jeannel, 1927: 555 (Vlorë); Moravec et al., 2003: 336. Trechus tristis: Csiki,
1940: 214 (Gjalicë Mt.); Pawlowski, 1973: 250; Hieke & Wrase, 1988: 38; Nonveiller et al., 1994: 9.
Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Alban. Exp.’18. Pashtrik 4-14.7”, 1 s. (NMW); “Kruja, Alban. Mader”, 3 s. (HNHM); “Tomor”
/ “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff
1937”, 67 s. (MNHUB); “Mal i Gjer” / “Albania leg. Bischoff 1937”, 15 s. (MNHUB).
Notes. Except the material studied above, in MNHUB the present author has seen more 48 s.,
which remained unregistered.
Species group “obtusiusculus”
40.10. Trechus albanicus Apfelbeck, 1905
References. Trechus albanicus: Apfelbeck, 1905: 241 (Northeastern Albania / Kosovo border
region: alpine belt of Koritnik Mt., loc. typ.); Apfelbeck, 1907b: 636-637 (alpine belt of Koritnik
Mt., loc. typ.); Jeannel, 1927: 563-564 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 214 (Gjalicë Mt.;
Korab Mt.); Casale & Laneyrie, 1982: 153; Drovenik & Peks, 1994: 39 (Koritnik Mt.); Drovenik
& Peks, 1999: 44 (Koritnik Mt.); Moravec et al., 2003: 326.
Material studied. “Latif Koritnik” / “Trechus albanicus Apfb. Typus! Coll. Apfelbeck”, 1 f. syntype
(HNHM); “Latif Koritnik”, 2 syntypes (DEI; NMW); “Latif Alban. spt.”, 1 s. (DEI); “Albania
Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Mts. Gyalica Lums” /
“Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki
1918”,2s.(HNHM);“Alban.Exped.GjalicaLjums17.-26.VI.’18”,1s.(NMW);“Gjalica”/“Albania
leg. Bischoff 1936”, 2 s. (NMW); “Gjallica” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).
Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b)
and the latter has long time been accredited to be correct (see also Casale & Laneyrie, 1982;
Drovenik & Peks, 1994: 39; Drovenik & Peks, 1999: 44; Moravec et al., 2003: 326). However,
it is incorrect and the original description must be assigned to the older one.
40.11. Trechus kobingeri Apfelbeck, 1902
40.11.1. Trechus kobingeri kobingeri Apfelbeck, 1902 [= distinctissimus Roubal, 1920]
References. Trechus kobingeri: Jeannel, 1925: 23 (Çukall Mt.); Jeannel, 1927: 566 (Çukall Mt.);
Casale & Laneyrie, 1982: 153; Nonveiller et al., 1994: 8, 18 (near to Shkodër). Trechus distinctissimus:

89. 88 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Casale & Laneyrie, 1982: 153. Trechus kobingeri bulgaricus: Guéorguiev et al., 1997: 10. Trechus
kobingeri kobingeri: Moravec et al., 2003: 333.
Material studied. “Maja e Jezerce Albania sept.”, 1 m. (NMW); “Cucale” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
Notes. Tråchus kobingeri bulgaricus Pawlowski, 1972 is a junior synonym of T. kobingeri pawlowskianus
P. Moravec & Lompe, 2003, and its record for Albania is erroneous. This race is known only
from a limited region in Western Bulgaria.
# Trechus obtusiusculus Ganglbauer, 1889
References. Trechus obtusiusculus: Nonveiller et al., 1994: 9.
Notes. The species is recorded with detailed records only from Bosnia Herzegovina and
Montenegro. The data for Albania is too doubtful, so that it is removed from the species list.
40.12. Trechus ravasinianus Lorenz, 1998 [= ravasinii Jeannel, 1925]
References. Trechus ravasinii: Jeannel, 1925: 24-25 (Tomor Mt.: Kulmak, Toik, Abbas Ali, loc.
typ.); Jeannel, 1927: 570-571 (Tomor Mt.: Abbas Ali, Kulmak, Toik); Casale & Laneyrie, 1982:
153. Trechus ravasinianus: Moravec et al., 2003: 339.
Material studied. “Tomor: Tomor Albania mer.” / “lg. Winkler Mai 1931”, 1 f. (HNHM);
“Tomor: Kulmak Albania mer.” / “lg. Winkler Mai 1931”, 12 s. (NMW).
Notes. Except the localities mentioned above, in MNHUB I have seen more than 100 s.,
which remained unregistered.
! Trechini gen. indet.
! Trechini gen. & sp. indet.
Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave
“Shpella e Haxise”, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT).
Notes. According to Vigna Taglianti (personal communication) the parts found in the cave
“Shpella e Haxise” may belong to an unknown genus close to Pheggomisetes Knirsch, 1923.
Subtribe TRECHODINA Jeannel, 1926
41. Genus Thalassophilus Wollaston, 1854
41.1. Thalassophilus longicornis (Sturm, 1825)
References. Thalassophilus longicornis: Apfelbeck, 1907a: 496 (Kiri River near Mesi).
Notes. Moravec et al. (2003: 346) have omitted the species for Albania. Actually, it needs
further confirmation for the country.

90. COLEOPTERA: CARABIDAE: HARPALINAE 89
Subfamily HARPALINAE Bonelli, 1810
Tribe CHLAENIINI Brullé, 1834
Subtribe CALLISTINA Laporte, 1834
42. Genus Callistus Bonelli, 1810
42.1. Callistus lunatus (Fabricius, 1775)
42.1.1. Callistus lunatus lunatus (Fabricius, 1775)
References. Callistus lunatus: Schatzmayr, 1943b: 99 (Lushnje); Murraj, 1968: 33 (Hibë; Sauk);
Hieke et al., 1986: 190 (Ibë near Krraba). Callistus lunatus lunatus: Kirschenhofer, 2003: 347.
Material studied. “Lomi Beratit Albanien”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr.
Grenze Penther’14”, 1 s. (NMW); “Hani Grabom Mai” / “Alban. montenegr. Grenze
Penther’14”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); Sauk,
9.VII.1961, 1 f. (MNST).
Notes. The records of Murraj (1968) are included here with minimal reservation since the
revision of a female from Sauk having the same label data as the published material has been
found to be correctly identified by him.
Subtribe CHLAENIINA Brullé, 1834
43. Genus Chlaenius Bonelli, 1810
Subgenus Chlaeniellus Reitter, 1908
43.1. Chlaenius flavipes Ménétries, 1832
References. Chlaenius flavipes: Apfelbeck, 1907a: 496, 498, 501 (Kiri River near Mesi; Fani i
Madh River near Oroshi); Schatzmayr, 1943b: 100 (Zezes River near Ura Zezes); Hieke et al.,
1986: 190 (Borshi near Vlorë); Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 349.
Material studied. “Oboti Bojana”, 1 s. (NMW); “Skutari Mesi”, 1 s. (NMW); “Merdita Oroschi”
/ “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.’18”, 1 s.
(NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 8 s. (NMW); “Elbasan Alb., Mader”, 8 s.
(NMW; MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); Tepelenë District, Vijosë
River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).
43.2. Chlaenius nigricornis (Fabricius, 1787)
References. Chlaenius nigricornis: Apfelbeck, 1904: 158 (Durrës); Hieke & Wrase, 1988: 149;
Kirschenhofer, 2003: 350.
Material studied. “Scutari l. See-Ufer”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Albania
Rittm. Matoha”, 1 s. (NMW); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB).

91. 90 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Murraj (1968: 33) cited this species from two localities (Tiranë; Tomor Mt.). Among
the material loaned from MNST, I found out 9 s. from the second place having the same label
data as the material published by Murraj as Chlaenius nigricornis. Actually, all the nine specimens
refer to Bembidion dalmatinum.
43.3. Chlaenius nitidulus (Schrank, 1781)
References. Chlaenius nitidulus: Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 350.
Material studied. “Maja ë Qori 2050 m, Albania”, 1 s. (NMW); Sauk, 5.III.1961, 1 m. (MNST).
Notes. First detailed records for Albania.
43.4. Chlaenius terminatus Dejean, 1826
References. Chlaenius terminatus: Schatzmayr, 1943b: 100 (Stan-Karbunarë; Skrofotina; Vlorë).
Material studied. “Oboti Bojana”, 3 s. (HNHM).
Notes. Kirschenhofer (2003: 350) has omitted the species for Albania. Here it is confirmed
for the country.
43.5. Chlaenius tristis (Schaller, 1783)
43.5.1. Chlaenius tristis tristis (Schaller, 1783)
References. Chlaenius tristis: Hieke & Wrase, 1988: 149. Chlaenius tristis tristis: Kirschenhofer, 2003: 350.
Material studied. “Oboti Bojana”, 5 s. (HNHM); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB).
Notes. First detailed records for Albania.
43.6. Chlaenius vestitus (Paykull, 1790)
References. Chlaenius vestitus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (Stan-
Karbunarë); Kirschenhofer, 2003: 351.
Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Lumi Dukatit Alban. mer.”, 4 s. (NMW);
“Maj’e Tartarit Albania mer.”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”,
1 s. (HNHM); “Alban. Exped. Kula Ljums 14.20.VII.’18”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr.
H. Priesner”, 4 s. (NMW); “Alessio, Alb. Matzenauer”, 2 s. (NMW); “Miloti, Alb. Matzenauer”, 1 s.
(NMW); 40 s. from different localities (MNHUB); Librazhd, Prenjas, 750 m, 7.V.1995, 1 s., S.
Golovach,B.Petrov,P.Stoevleg.(NMNHS);DrinaRiver, 3.VI.1995,1s.,V.Sakalainleg.(NMNHS).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Subgenus Chlaenites Motschulsky, 1860
43.7. Chlaenius spoliatus (P. Rossi, 1792)
43.7.1. Chlaenius spoliatus spoliatus (P. Rossi, 1792)
References. Chlaenius spoliatus: Schatzmayr, 1943b: 99 (Durrës); Hieke & Wrase, 1988: 147.
Chlaenius spoliatus spoliatus: Kirschenhofer, 2003: 351.

92. COLEOPTERA: CARABIDAE: HARPALINAE 91
Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Oboti Bojana”, 2 s.
(HNHM); “Durazzo” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Himara juni” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 5 s.
(MNHUB); “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i
That” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Subgenus Chlaenius s.str.
43.8. Chlaenius festivus (Panzer, 1796)
43.8.1. Chlaenius festivus festivus (Panzer, 1796)
References. Chlaenius festivus: Apfelbeck, 1904: 157 (Vlorë; Velipoja near Lezhë); Apfelbeck,
1909: 298 (Vlorë); Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (Lushnje); Murraj,
1968: 33 (Radenjë); Hieke & Wrase, 1988: 148. Chlaenius festivus festivus: Gridelli, 1925: 120
(Shkodër; Berat); Kirschenhofer, 2003: 351.
Material studied. “Alban., Wgth. Valona”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Skutari
l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk.
Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); nearly
40 s. from different localities (MNHUB); Sauk, 18.II.1961, 1 f. (MNST); Rodenj, 26-28.VII.1962,
1 s. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 4 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. The specimen collected at Liqen i Shkodrës was determined as C. velutinus (Duftschmid,
1812) by Kirschenhofer, but I refer this record to the nominotypical subspecies. The material
published by Murraj (1968) was found to be correctly determinated.
Subgenus Dinodes Bonelli, 1810
43.9. Chlaenius cruralis Fischer von Waldheim, 1829
References. Chlaenius cruralis: Kirschenhofer, 2003: 352.
Material studied. “Skutari” / “Strasser”, 3 s. (NMW); Ibë, 4.VII.1963, 1 m. (MNST).
Notes. First detailed records for Albania.
43.10. Chlaenius decipiens (L. Dufour, 1820) [= azureus Duftschmid, 1812, nec Fabricius,
1775; laticollis Chaudoir, 1843; ambiguus Csiki, 1931]
References. Chlaenius decipiens: Apfelbeck, 1904: 154 (Vlorë); Hieke et al., 1986: 190 (Ibë near
Krraba); Hieke & Wrase, 1988: 148; Kirschenhofer, 2003: 352. Chlaenius azureus: Apfelbeck,
1909: 298 (Vlorë). Chlaenius decipiens var. ambiguus: Csiki, 1940: 219 (Gjalicë Mt.). Chlaenius
decipiens laticollis: Schatzmayr, 1943b: 100 (Lushnje).
Material studied. “Elbasan Albanien”, 1 s. (NMW); “Albania Mts. Gyalica Lums” / “Akad.
Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tepelene Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”,
1 s. (NMW); “Tirane” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg.

93. 92 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Bischoff 1936”, 1 s. (MNHUB); “Ivan Geb Korce” / “Albania 1936 leg. Ramadan coll. Bischoff”,
1 s. (MNHUB); Tiranë, 27.IX.1958, 1 m. (MNST); Tiranë, 10.V.1962, 1 m. (MNST).
Subgenus Epomis Bonelli, 1810
43.11. Chlaenius circumscriptus (Duftschmid, 1812)
References. Chlaenius circumscriptus: Kirschenhofer, 2003: 353.
Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Liqen i Butrintes”
/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
43.12. Chlaenius dejeani (Dejean, 1831)
References. Chlaenius dejeani: Apfelbeck, 1904: 154 (Vlorë); Schatzmayr, 1943b: 99 (Lushnje);
Hieke & Wrase, 1988: 147. Epomis dejeani: Apfelbeck, 1909: 298 (Vlorë).
Material studied.“Gjormi”/“Albanialeg.Bischoff 1935”, 1s.(MNHUB);“Murthi”/“Albanialeg.
Bischoff 1936”, 2 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB).
Notes. Kirschenhofer (2003: 353) has omitted the species for Albania. Here it is confirmed
for the country.
Subgenus Trichochlaenius Seidlitz, 1887
43.13. Chlaenius aeneocephalus Dejean, 1826
43.13.1. Chlaenius aeneocephalus aeneocephalus Dejean, 1826
References. Chlaenius aeneocephalus: Apfelbeck, 1904: 156 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);
Hieke & Wrase, 1988: 148.
Notes. Kirschenhofer (2003: 355) has omitted the species for Albania, so that it needs of
further confirmation for the country.
Tribe CYCLOSOMINI Laporte, 1834
Subtribe MASOREINA Chaudoir, 1870
44. Genus Masoreus Dejean, 1821
Subgenus Masoreus s.str.
44.1. Masoreus wetterhallii (Gyllenhal, 1813)
44.1.1. Masoreus wetterhallii wetterhallii (Gyllenhal, 1813)
References. Masoreus wetterhallii: Hieke & Wrase, 1988: 152.
Material studied. “Tomor Tojk” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. Bousquet (2003b: 357) has omitted the species for Albania. Here it is confirmed for
the country with a first detailed record.

94. COLEOPTERA: CARABIDAE: HARPALINAE 93
Tribe DRYPTINI Bonelli, 1810
45. Genus Drypta Latreille, 1796
Subgenus Drypta s.str.
45.1. Drypta dentata (P. Rossi, 1790)
References. Drypta dentata: Apfelbeck, 1904: 346 (Velipoja); Hieke & Wrase, 1988: 153; Baehr,
2003b: 358.
Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB).
Tribe HARPALINI Bonelli, 1810
Subtribe AMBLYSTOMINA Fauvel, 1889
46. Genus Amblystomus Erichson, 1837
46.1. Amblystomus levantinus Reitter, 1883
46.1.1. Amblystomus levantinus levantinus Reitter, 1883
References. Amblystomus levantinus: Apfelbeck, 1904 (Velipoja); Schatzmayr, 1943b: 101 (Stan-
Karbunarë; Lushnje); Hieke & Wrase, 1988: 165 (Velipoja). Amblystomus levantinus levantinus:
Jaeger & Wrase, 2003: 360.
Notes. According to D. Wrase (personal communication) the “metallescens-levantinus” group
needs revision. This notable specialist on the Palaearctic Harpalini wrote me: “I don’t know
really what levantinus is, as I haven’t checked yet the type. I have several species, clearly different
in genitalia belonging to that group. For the time being the best solution of that problem it to
continue the classification we have now.”.
46.2. Amblystomus metallescens (Dejean, 1829)
References. Amblystomus metallescens: Csiki, 1940: 219 (Kula Ljums; Gjalicë Mt.); Schatzmayr,
1943b: 101 (Lushnje; Mal i Krujes); Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Oboti Bojana”, 1 s. (HNHM);
“Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” /
“Akad.Balk.Exp.Csiki1918”,1s.(HNHM);“Durres”/“Albanialeg.Bischoff 1932”,1s.(MNHUB).
46.3. Amblystomus niger (Heer, 1841)
References. Amblystomus niger: Hieke et al., 1986: 188 (Ibë near Krraba; Bizë near Shëngjergji);
Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Durres” / “Albania leg.
Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB);
“Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 8.II.1998 / 15.II.1998,
4 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).

95. 94 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subtribe ANISODACTYLINA Lacordaire, 1854
47. Genus Anisodactylus Dejean, 1829
Subgenus Anisodactylus s.str.
47.1. Anisodactylus binotatus (Fabricius, 1787) [= spurcaticornis Dejean, 1829]
References. Anisodactylus binotatus: Apfelbeck, 1909: 298 (Pashaliman); Hieke et al., 1986: 186
(North Albanian Alps near Theth); Hieke & Wrase, 1988: 112; Ito, 2003a: 360. Anisodactylus
binotatus ab. spurcaticornis: Apfelbeck, 1909: 298 (Pasha Liman).
Material studied. “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Theth, 1-8.VIII.
1961, 1 m. (MNST); Theth, 800-1200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).
47.2. Anisodactylus nemorivagus (Duftschmid, 1812)
References. Anisodactylus nemorivagus: Apfelbeck, 1904: 212 (Velipoja); Horion, 1941: 247; Hieke
& Wrase, 1988: 112.
Notes. Ito (2003a: 361) has omitted the species for Albania. Actually, it needs further
confirmation for the country.
47.3. Anisodactylus pueli Schauberger, 1933
47.3.1. Anisodactylus pueli pueli Schauberger, 1933
References. Anisodactylus pueli: Schauberger, 1933: 124 (Vlorë, loc. typ.); Sciaky, 1979: 15 (Vlorë); Turin,
1981: 150; Wrase, 1991: 12; Noonan, 1996: 34, 109 (Vlorë). Anisodactylus pueli pueli: Ito, 2003a: 361.
Material studied. “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Subgenus Pseudodichirius Lutshnik, 1921
47.4. Anisodactylus intermedius Dejean, 1829
References. Anisodactylus intermedius: Hieke & Wrase, 1988: 113; Ito, 2003a: 361.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Maja e Konispol Seraqin” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB);Rodeni,2-22.VII.1962,1s.(MNST);Gjirokastra,6.V.1994,1m.,P.Stoevleg.(NMNHS).
Notes. First detailed records for Albania.
48. Genus Diachromus Erichson, 1837
48.1. Diachromus germanus (Linnaeus, 1758)
References. Diachromus germanus: Ito, 2003a: 362.
Material studied. “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 2 s. (HNHM).
Notes. First detailed record for Albania.

96. COLEOPTERA: CARABIDAE: HARPALINAE 95
49. Genus Gynandromorphus Dejean, 1829
49.1. Gynandromorphus etruscus (Quensel, 1806)
49.1.1. Gynandromorphus etruscus etruscus (Quensel, 1806)
References. Gynandromorphus etruscus: Schatzmayr, 1943b: 105 (Dukati); Hieke & Wrase, 1988:
113. Gynandromorphus etruscus etruscus: Ito, 2003a: 362.
Material studied. “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Radona, Shkiela,
21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).
Subtribe DITOMINA Bonelli, 1810
50. Genus Carterus Dejean, 1830
Subgenus Carterus s.str.
50.1. Carterus dama (P. Rossi, 1792)
References. Ditomus dama: Apfelbeck, 1909: 298 (Vlorë). Carterus dama: Hieke & Wrase, 1988:
145; Wrase, 1989: 39 (Mursi - Kara Ali Bey); Wrase, 2003b: 363.
Material studied. “Murrsi - Kara Ali Bey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);
Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).
Additional material included (studied D.W. Wrase). Oricum, south of Vlorë, X.1994, 2 f.,
K. Resl leg. (cWR).
50.2. Carterus gilvipes (Piochard de la Brûlerie, 1873)
References. Carterus gilvipes: Schatzmayr, 1943b: 101 (Dukati); Wrase, 2003b: 364.
50.3. Carterus rotundicollis Rambur, 1837
Material studied. Borsh, 15-28.V.1961, 1 f. (MNST).
Notes. New species for the fauna of Albania.
51. Genus Ditomus Bonelli, 1810
51.1. Ditomus calydonius (P. Rossi, 1790)
51.1.1. Ditomus calydonius calydonius (P. Rossi, 1790)
References. Ditomus calydonius calydonius: Wrase, 2003b: 364.
51.2. Ditomus tricuspidatus (Fabricius, 1792)
References. Ditomus tricuspidatus: Ganglbauer, 1889: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);
Wrase, 2003b: 365. Carterus tricuspidatus: Apfelbeck, 1904: 173 (Vlorë).

97. 96 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB); “Brataj Ramica Albania mer.”
/ “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “M. e Tartarit juni 1932” / “Albania leg.
Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Borshne” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); Borsh, 15-28.V.1961, 1 f. (MNST);
Ljukove, 100 m, 5.VI.1995, 1 m., G. Blagoev leg. (NMNHS).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 2 s., M.
Komnenov leg. (cSH).
52. Genus Dixus Billberg, 1820
52.1. Dixus clypeatus (P. Rossi, 1790)
References. Aristus clypeatus: Apfelbeck, 1909: 298 (Berat). Ditomus clypeatus: Hieke et al., 1986:
186 (Borshi near Vlorë; Mali me Gropë near Livadhet e Selitës; Mal i Dajtit Mt.: Shkall Priskë);
Hieke & Wrase, 1988: 146. Dixus clypeatus: Wrase, 2003b: 365.
Material studied. “Kruja, Alban., Mader”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff
1933”, 3 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Albanien
Dajti…6.6.55 Burmeister”, 1 s. (MNHUB); “N 111”, 1 s. (MNST), Sauk, 10-20.VI.1961, 1 s.
(MNST).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M.
Komnenov leg. (cSH).
52.2. Dixus eremita (Dejean, 1825)
References. Ditomus eremita: Schatzmayr, 1943b: 102 (Skrofotina; Lushnje). Dixus eremita: Wrase,
2003b: 365; Wrase, 2005: 820.
52.3. Dixus obscurus (Dejean, 1825)
References. Aristus obscurus: Apfelbeck, 1909: 298 (Vlorë; Berat). Ditomus obscurus: Schatzmayr,
1943b: 101 (Lushnje); Hieke et al., 1986: 186 (Borshi near Vlorë; Poliçan near Tomor Mt.).
Dixus obscurus: Wrase, 2003b: 365.
Material studied. “Serandë” Piqeros” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Tirane”
/ “Albania leg. Bischoff 1933”, 1 s. (MNHUB).
53. Genus Graniger Motschulsky, 1864
53.1. Graniger femoralis (Coquerel, 1859)
References. Carterophonus femoralis: Sciaky & Franzini, 1976: 26 (Berat).

98. COLEOPTERA: CARABIDAE: HARPALINAE 97
54. Genus Odontocarus Solier, 1835
Notes. New genus for the fauna of Albania.
54.1. Odontocarus robustus (Dejean, 1831)
Material studied. “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);
“Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania
leg. Bischoff 1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
Butrinti, 4.VI.1995, 1 s., V. Sakalian leg. (NMNHS).
Notes. New species for the fauna of Albania.
[genus Pachycarus Solier, 1835]
[subgenus Mystropterus Chaudoir, 1842]
[Pachycarus macedonicus V.B. Guéorguiev & B.V. Guéorguiev, 1997]
Notes. This species has been described from the Macedonian part of the Galičica Mt. very
close to the Albanian border, so that it may inhabit Albania too.
Subtribe HARPALINA Bonelli, 1810
55. Genus Acinopus Dejean, 1821
Subgenus Acinopus s.str.
55.1. Acinopus baudii A. Fiori, 1913
55.1.1. Acinopus baudii hellenicus Heinz, 1982
References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995:
205; Kataev & Wrase, 2003: 368. Acinopus baudii hellenicus: Heinz, 1982: 16 (Tepelenë; Maja e
Konispol, Seraqin); Kataev & Wrase, 2003: 368.
Material studied. “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Murthe bis Kara Alibey” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB).
55.2. Acinopus picipes (Olivier, 1795) [= emarginatus Chaudoir, 1842]
References. Acinopus emarginatus: Apfelbeck, 1904: 180 (Vlorë). Acinopus picipes: Apfelbeck,
1909: 297 (Vlorë); Schatzmayr, 1943b: 102 (Karbunara e Madhe; Tërpan; Lushnje); Hieke &
Wrase, 1988: 143; Kataev & Wrase, 2003: 368.
Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Mal i That”
/ “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Mal i Dajtit Mt.: Shkall Priskë, 26-30. VI.1961,
1 f. (MNST); Ibë, 4.VII.1962, 1 m. (MNST); Ibë, 8.VII.1966, 1 f. (MNST); “AL (Berat). Paß ca.
5 km S. Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 s. (MNHUB); Ljukove, 100 m, 5.VI.1995,
1 s., G. Blagoev leg. (NMNHS).
Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i
That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., Heinz & Peks leg. (cWR); Hotel “Edva”
at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH); Dhermi, 200 m,
12.VII.2006, 2 s., M. Komnenov leg. (cSH).

99. 98 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
# Acinopus subquadratus Brullé, 1832
References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995:
205; Kataev & Wrase, 2003: 368.
Notes. Guéorguiev & Guéorguiev (1995) mentioned this taxon for Albania, following an
older indication for South Albania (Magistretti, 1965: 272). Kataev & Wrase (2003: 368) have
most probably followed also the same sources. However, according to Heinz (1982) A.
subquadratus inhabits only the southern and central regions of Greece and the data on its
presence in Albania concerns A. baudii hellenicus; both species were often mixed in the past.
The species is removed from the list of the Albanian Carabidae.
Subgenus Oedematicus Bedel, 1897
55.3. Acinopus megacephalus (P. Rossi, 1794)
References. Acinopus megacephalus: Apfelbeck, 1904: 180 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);
Puel, 1934: 79; Schatzmayr, 1943b: 102 (Dukati; Skrofolina); Jeanne, 1984: 24; Hieke & Wrase,
1988: 143; Kataev & Wrase, 2003: 369.
Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 3 s. (MNHUB); “Durrës” / “Albania
leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB);
“Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 3 s. (MNHUB).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Lushnja-Divjaka,
21.VII.1993, 1 m., L. Latella leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of
Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva”
at Radhime (Vlorë District), 8-15.VII.2006, 3 s., leg. M. Komnenov (cSH).
Subgenus Osimus Fischer von Waldheim, 1829
55.4. Acinopus ammophilus Dejean, 1829
References. Acinopus ammophilus: Apfelbeck, 1907a: 498, 501 (Oroshi).
Notes. Kataev & Wrase (2003: 369) have omitted the species for Albania. Actually, it needs
further confirmation for the country.
56. Genus Harpalus Latreille, 1802
Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982
56.1. Harpalus tenebrosus Dejean, 1829 [= centralis Schauberger, 1929]
References. Harpalus tenebrosus var. centralis: Csiki, 1940: 220 (Kula Ljums). Harpalus tenebrosus:
Hieke & Wrase, 1988: 138; Kataev & Wrase, 2003: 370.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM);
“Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).

100. COLEOPTERA: CARABIDAE: HARPALINAE 99
Subgenus Harpalus s.str.
Species group “honestus”
56.2. Harpalus attenuatus Stephens, 1828
References. Harpalus attenuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Csiki, 1940: 220 (Shkëlzen Mt.; Kula Ljums); Schatzmayr, 1943b: 104 (“Poggio Boschetto”);
Hieke et al., 1986: 187 (Borshi near Vlorë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 139;
Kataev & Wrase, 2003: 373.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania Skölsen” / “Akad.
Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”,
1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Burelle” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);
“Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja ae Konispol Seraqin” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Vlesh” / “30.VII.1958 leg. Xh. Murraj”,
1 s. (HNHM); “Albanien-Exp. DEI Borshi südl. Vlora 14.-27.V.1961”, 1 s. (MNHUB); Mal i
Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 2 f. (MNST); 2 km N from Durrës, 24.V.1993, 1 s., P.
Stoev & D. Zaprjanova leg. (NMNHS).
56.3. Harpalus honestus (Duftschmid, 1812)
56.3.1. Harpalus honestus honestus (Duftschmid, 1812)
References. Harpalus honestus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Csiki, 1940: 220 (Korab Mt.); Hieke & Wrase, 1988: 138. Harpalus honestus honestus: Kataev &
Wrase, 2003: 377.
Material studied. “Vermosa Penther”, 20 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr.
Grenze, Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze,
Penther’14”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Alban. Exp. Kula Ljums 18-28.V.’18”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”,
1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg.
Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 3 s.
(MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Gjallica” / “Albania
leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);
Ibë, 18.V.1960, 1 m. (MNST).
56.4. Harpalus rubripes (Duftschmid, 1812)
References. Harpalus rubripes: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Csiki, 1940: 220 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke & Wrase, 1988: 133; Kataev &
Wrase, 2003: 381.
Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania
Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Korab” / “Akad. Balk. Exp.
Csiki 1918”, 1 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1
s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Alban. Exp. 1918 Korab, 23-
31.VII.”, 1 s. (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal

101. 100 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
i Shenjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Okol Shkalla” / “Albania leg.
Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB);
“AL (Tirana) 8 km N Psene. E Tirana 1450 m. 30.IV.1993 leg. H. + S. Peks”, 2 m. (MNHUB);
North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);
Cafasan, 1230 m, 27.VII.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS).
56.5. Harpalus rufipalpis Sturm, 1818
56.5.1. Harpalus rufipalpis rufipalpis Sturm, 1818 [= rufitarsis (Duftschmid, 1812)]
References. Harpalus rufitarsis: Apfelbeck, 1904: 196 (Berat); Apfelbeck, 1907a: 499 (Mal i
Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Berat); Hieke et al., 1986: 188 (Bizë
near Shëngjergji); Hieke & Wrase, 1988: 138. Harpalus rufipalpis rufipalpis: Kataev & Wrase,
2003: 381.
Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mal i Shenjit”
/ “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 1 s.
(MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Maranaj” / “Albania leg.
Bischoff 1937”, 2 s. (MNHUB); “Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 3 s. (HNHM).
Notes. In addition to the material studied above, in NMW I have seen more specimens, which
remained unregistered.
56.6. Harpalus sulphuripes Germar, 1824
56.6.1. Harpalus sulphuripes sulphuripes Germar, 1824
References. Harpalus sulphuripes: Apfelbeck, 1904: 196 (Vlorë); Anonymous, 1907: 284;
Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë);
Schauberger, 1926: 32 (Vlorë; Pasha Liman); Hieke et al., 1986: 188 (Tomor Mt.: Abbas
Ali, 2401 m); Hieke & Wrase, 1988: 139. Harpalus sulphuripes sulphuripes: Kataev & Wrase,
2003: 383.
Material studied. More than 50 s. from different localities (MNHUB); North Albanian Alps,
Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS); Saranda District,
Ljukove, 5.V.1995, 1 s., P. Stoev leg. (NMNHS); Gjirokastra, 6.V.1994, 1 s., P. Stoev leg.
(NMNHS); 12 km NE Erseka, 900 m, 12.V.1995, 1 s., L. Penev leg. (NMNHS).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M.
Komnenov leg. (cSH).
Species group “atratus”
56.7. Harpalus atratus Latreille, 1804 [= subsinuatus (Duftschmid, 1812)]
References. Harpalus atratus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Hieke et al., 1986: 187 (Mal i Dajtit Mt.: Shkall Priskë); Hieke & Wrase, 1988: 137; Kataev &
Wrase, 2003: 373. Harpalus atratus var. subsinuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top
region, 1500 m).
Material studied. “Albania Zalle-Dardha” / “23.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); more
than 60 s. from different localities (MNHUB).

102. COLEOPTERA: CARABIDAE: HARPALINAE 101
Additional material included (studied A. Vigna Taglianti). 5 km S of Tepelenë (crossroads
Permet), 350 m, 21-22.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. In addition to the material studied above, in NMW I have seen more specimens, which
remained unregistered.
Species group “quadripunctatus”
56.8. Harpalus laevipes Zetterstedt, 1828 [= quadripunctatus Dejean, 1829]
References. Harpalus quadripunctatus: Hieke & Wrase, 1988: 133. Harpalus laevipes: Kataev &
Wrase, 2003: 378.
Material studied. “Gjallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. First detailed record for Albania.
Species group “serripes”
56.9. Harpalus flavicornis Dejean, 1829
56.9.1. Harpalus flavicornis flavicornis Dejean, 1829
References. Harpalus flavicornis: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);
Hieke & Wrase, 1988: 136. Harpalus flavicornis flavicornis: Kataev & Wrase, 2003: 376.
Materialstudied.“MaliDajtit”/“Albanialeg.Bischoff 1932”,1s.(MNHUB);“Tirane”/“Albania
leg. Bischoff 1933”,1s. (MNHUB); “LumiiErzen” /“leg.Bischoff Albania1934”, 1s. (MNHUB);
“Mal i Dajtit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tirane” / “leg. Bischoff Albania
1934”, 3 s. (MNHUB); “Mal i That” / “Albania 1936 leg. Ramadan coll. Bischoff”, 1 s. (MNHUB);
“Tiranë” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff
1937”, 1 s. (MNHUB); “Albanien - Tirana Dajti-Vorberge 3.V.39…Müller J.”, 2 s. (MNHUB); Mal
i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 m. (MNST); Zooteknika, 14.V.1963, 1 s. (MNST); “AL
(Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 1 s. (MNHUB); “AL (Berat).
Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB); Cafasan, 1230 m,
27.VII.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 4 s., G.
Blagoev leg. (NMNHS); 5 km Leskovac, 650 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNH); Tiranë
Park, 11.I.1998 / 8.II.1998, 4 s., R. Ward leg. (CMNH).
56.10. Harpalus serripes (Quensel, 1806)
56.10.1. Harpalus serripes serripes (Quensel, 1806) [= subchalybaeus Reitter, 1900]
References. Harpalus serripes: Apfelbeck, 1904: 197 (Vlorë); Anonymous, 1907: 283; Apfelbeck,
1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Csiki, 1940:
220 (Korab Mt.); Hieke et al., 1986: 187 (Ibë near Krraba); Hieke & Wrase, 1988: 135. Harpalus
serripes ab. subchalybaeus: Schauberger, 1926: 25 (Dukati i Ri). Harpalus serripes serripes: Kataev &
Wrase, 2003: 382.
Material studied. L. Selitës, 3-9.VII.1961, 1 m. (MNST); Ibë, 4.VII.1963, 1 f. (MNST); “AL
(Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. + S. Peks”, 10 s. (MNHUB); 2 km N from
Durrës, 24.V.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Cafasan, 1230 m, 27.VII.1993,
3 s., P. Stoev, D. Zaprjanova leg. (NMNHS); Lushnja, Divjaka National Park, 10.V.1995, 1 s., S.

103. 102 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m, 6.VI.1995, 3 s., G. Blagoev
leg. (NMNHS).
Additional material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë
District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).
Notes. In addition to the material studied above, in NMW and ZNHU I have seen more than
130 s. from different localities, which remained unregistered.
Species group “pumilus”
[Harpalus picipennis (Duftschmid, 1812)]
References. Harpalus picipennis: Apfelbeck, 1904: 200; Hieke & Wrase, 1988: 141; Kataev &
Wrase, 2003: 380.
Notes. Hieke & Wrase (1988) noted the species for Albania following Apfelbeck’s record
“Prisren” (Apfelbeck, 1904: 200). In fact this place is situated in the Kosovo Region of Serbia.
Most likely, Kataev & Wrase (2003) have used the same sources. For the time being the species
is treated here as probable but not yet proven for Albania (having in mind recent data from
the north parts of Republic of Macedonia, unpublished information).
56.11. Harpalus pumilus Sturm, 1818 [= vernalis Fabricius, 1801]
References. Harpalus vernalis: Müller, 1931: 69; Hieke & Wrase, 1988: 140; Kataev & Wrase,
2003: 380.
Material studied. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).
Notes. First detailed record for Albania.
Species group “anxius”
56.12. Harpalus anxius (Duftschmid, 1812)
References. Harpalus anxius: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Kataev & Wrase, 2003: 373.
Material studied. “Latif Cucali”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt.
IV.V.’05”, 2 s. (NMW); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B. Petrov, P.
Stoev leg. (NMNHS).
Notes. Hieke & Wrase (1988: 139) noted this species for Albania apparently following the
record “Jannina” (Apfelbeck, 1904: 199). In fact this locality is situated in Greece.
56.13. Harpalus subcylindricus Dejean, 1829
References. Harpalus subcylindricus: Hieke & Wrase, 1988: 140; Kataev & Wrase, 2003: 382.
Material studied. “Skutari” / “Strasser”, 1 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1
s. (NMW); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rapsh” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
Mal i Dajtit Mt., 10.V.1965, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë ~ 600 m 27.IV.1993
leg. Heinz u. Sonja Peks”, 3 s. (MNHUB); “AL (Berat). Paß ca. 5 km S Gllava 600 m. 27.IV.1993
leg. H. + S. Peks”, 1 m., 1 f. (MNHUB); North Albanian Alps, Maj’e Radohimes, 2400-2550

104. COLEOPTERA: CARABIDAE: HARPALINAE 103
m, 29.V.1993, 3 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Radohimes,
1000-1100 m, 5.IX.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS).
Notes. First detailed records for Albania.
Species group “flavescens”
56.14. Harpalus flavescens (Piller et Mitterpacher, 1783)
Material studied. “Elbasan, Alb. 11.IX.18 Dr. H. Priesner”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
Species group “tardus”
56.15. Harpalus albanicus Reitter, 1900
References. Harpalus albanicus: Reitter, 1900: 114 (? Shkodër, loc. typ.); Apfelbeck, 1904: 200
(Shkodër); Rambousek, 1912: 86; Horion, 1941: 230; Roudier, 1949: 176 (Shkodër); Magistretti,
1965: 308; Hieke & Wrase, 1988: 134; Kataev & Wrase, 2003: 372.
Material studied. “Skutari” / “coll. Reitter”, 1 f. syntype (HNHM).
Notes. Tschitschérine (1901: 154) expressed doubt about the correct interpretation of the
species type locality “Scutari”. It may as well represent the present city of Üsküdar located on
the Asian shore of the Bosporus right opposite the peninsula of Istanbul, or the present
Albanian City of Shkodër.
56.16. Harpalus tardus (Panzer, 1796)
References. Harpalus tardus: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top
region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988:134; Kataev & Wrase, 2003: 383.
Material studied. “Latif Oroshi”, 1 s. (NMW); “Maranaj” / “Apfelb. Alban. spt. IV.V.’05”, 2
s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.1918”, 1 s. (NMW).
Species group “latus”
56.17. Harpalus latus (Linnaeus, 1758)
References. Harpalus latus: Kataev & Wrase, 2003: 378.
Materialstudied. “Rapsa10-18.V.” /“Alban. montenegr.Grenze, Penther’14”, 1s. (NMW); North
Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & P. Stoev leg. (NMNHS).
Notes. First detailed records for Albania.
56.18. Harpalus solitaris Dejean, 1829
References. Harpalus solitaris: Kataev & Wrase, 2003: 382.
Material studied. North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 f., P. Beron, B.
Petrov leg. (NMNHS).
Notes. First detailed record for Albania.

105. 104 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Species group “smaragdinus”
56.19. Harpalus smaragdinus (Duftschmid, 1812)
References. Harpalus smaragdinus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500
m); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 382.
Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Mal i
Shenjit” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); Theth, 1-8.VIII.1961, 1 m. (MNST);
“AL (Shkodar). Paß SE Puke 850 m 2.V.1993 leg. H. & S. Peks”, 1 m. (MNHUB); North
Albanian Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & B. Petrov leg.
(NMNHS); Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Rapsh, 15.V.2002,
3 m., 1 f., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,
1 s., leg. M. Komnenov (cSH).
Species group “autumnalis”
# Harpalus anatolicus Tschitschérine, 1898
# Harpalus anatolicus anatolicus Tschitschérine, 1898 [= skutariensis Reitter, 1900]
References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”).
Notes. Reitter (1900: 107) indicated as type locality of H. skutariensis the site “Scutari” (see
also Kataev & Wrase, 1997). “Scutari” was the Greek name for the district of Istanbul now
known as Üsküdar, located on the Asian shore of the Bosporus right opposite the peninsula
of Istanbul. The same name was also used in the past for the present city of Shkodër.
Tschitschérine (1901: 146) has stated that H. skutariensis Reitter is identical with H. tristis. Apfelbeck
(1904: 199)acceptedthisopiniondespiteDaniel&Daniel(1902:250-251)hadalreadysynonymized
H. tristis to H. danieli. Later on, Mlynář (1979: 83) considered H. skutariensis as synonym of H.
anatolicus. Kataev & Wrase (1997) confirmed this synonymy and correctly gave data only for the
Anatolian Peninsula. The species is excluded from the list of the Albanian Carabidae.
56.20. Harpalus autumnalis (Duftschmid, 1812)
References. Harpalus autumnalis: Hieke & Wrase, 1988: 136; Kataev & Wrase, 2003: 373.
Material studied. “Skutari Bojana”, 3 s. (NMW); “Lum i Tiranes” / “Albania leg. Bischoff
1933”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
# Harpalus danieli Reitter, 1900 [= tristis Tschitschérine, 1898]
References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”); Müller, 1931: 65 (“Skutari”).
Notes. Tschitschérine (1901: 146) stated H. skutariensis Reitter is synonym of H. tristis, but a
bit later Daniel & Daniel (1902: 250-251) synonymized H. tristis to H. danieli. Apfelbeck (1904:
199) accepted the opinion of Tschitschérine. Later on, Mlynář (1979) solved the problem (see
notes under H. anatolicus). Recently, Kataev & Wrase (2003: 375) have indicated H. danieli for
European Turkey and former Yugoslavia (Serbia and Montenegro). I consider, however, due
to lack of reliable records, the species should be excluded from the faunas of Albania and
adjacent territories in the north.

106. COLEOPTERA: CARABIDAE: HARPALINAE 105
[Harpalus triseriatus A. Fleischer, 1897]
Notes. This Balkan endemic species lives in Bosnia Herzegovina, Macedonia, Bulgaria and
Greece (Kataev & Wrase, 2003: 383) and most probably inhabits Albania as well.
Species group “cupreus”
56.21. Harpalus cupreus Dejean, 1829
56.21.1. Harpalus cupreus fastuosus Faldermann, 1836
References. Harpalus cupreus fastuosus: Schatzmayr, 1943b: 104 (Lushnje).
Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Murrsi-
Kara Alibey” / “leg. Bischoff Albania 1934” / “cotype” / “Harpalus bischoffi Schaub. det. Dr.
E. Schaub. 35” / “(H. bischoffi Schb. i. l.) = H. cupreus Z. Mlynar det. 1976”, 1 m. with extracted
genitalia (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).
Notes. Kataev & Wrase (2003: 374) have omitted the species for Albania. Here it is confirmed
for the country. Harpalus bischoffi Schauberger is nomen nudum.
56.22. Harpalus euchlorus Ménétriés, 1839
Material studied. “Skutari” / “Strasser” / “collectio Paganetti”, 2 s. (NMW).
Notes. New species for the fauna of Albania. B. Kataev (personal communication) expressed
the opinion that H. euchlorus may be a synonym of the preceding species.
Species group “dimidiatus”
56.23. Harpalus dimidiatus (P. Rossi, 1790)
References. Harpalus dimidiatus: Apfelbeck, 1904: 193 (Vlorë; Berat); Apfelbeck, 1907a: 495
(northeast of Shkodër); Apfelbeck, 1909: 298 (Vlorë; Berat); Csiki, 1940: 220 (Kula Ljums);
Schatzmayr, 1943b: 104 (“Poggio Boschetto”; Lushnje; Dukati); Hieke et al., 1986: 187 (North
Albanian Alps near Theth); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 375.
Material studied. “Merdita M. Scheit”, 2 s. (NMW); “Skutari Mesi”, 2 s. (NMW); “Albania
Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Alban. Exp. Kula Ljums 18-
28.V.’18”, 5 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 2 s. (NMW); more than 60 s. from
different localities (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 2 s. (HNHM);
“Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 12 s. (HNHM); “Albania Shkoder” /
“21.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); Ibë, 24.IV.1962, 2 f. (MNST); Ibë, 4.VII.1963, 2 f.
(MNST); Mal i Dajtit Mt., 10.V.1965, 3 m., 1 f. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m.
(MNST); Tiranë, 7.IV.1966, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993
leg. Heinz u. S. Peks”, 10 s. (MNHUB); “AL (Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. +
S. Peks”, 5 s. (MNHUB); “AL (Shkodar) Paß SE Puke 850 m 2.V.1993”, 1 s. (MNHUB);
Hotolisht - Librazhd, 300 m, 7.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);
Librazhd - Prenjas, 750 m, 7.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);
Tiranë, botanical garden, 8.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Dukati
Mt., 30 km S Vlorë, 450 m, 11.V.1995, 2 s., L. Penev leg. (NMNHS); Kiari Pass, 1160 m,
1.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Tiranë Park, 22.II.1998, 1 s., R. Ward leg. (CMNH).

107. 106 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Additional material included (studied S. Hristovski, A. Vigna Taglianti and D.W. Wrase).
Tiranë, XI.58, 1 s. (collector illegible) (cWR); Tiranë District, Kffaba-Paß, Südseite, 500 m,
29.IV.1993, 5 s., H. & S. Peks leg. (cWR); Korcë District, Mai i That Mt., Zwezde-Sattel, 1000-
1100 m, 4.V.1993, 6 s., Heinz & Peks leg. (cWR); Fier, 4.VII.1993, 1 f., L. Latella leg. (cAVT);
Tiranë, botanical garden, 4.VI.2002, 1 f., A. Paparistos leg. (cAVT); Tiranë, Mal i Dajtit Mt.,
7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley of
Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva”
at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).
Species group “dispar”
56.24. Harpalus punctatostriatus Dejean, 1829
References. Harpalus punctatostriatus: Apfelbeck, 1909: 298 (Vlorë; Sop); Schatzmayr, 1943b:
103 (Lushnje); Hieke & Wrase, 1988: 128; Kataev & Wrase, 2003: 380.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania 1918”, 1 s. (HNHM);
“Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
56.25. Harpalus pygmaeus Dejean, 1829
References. Harpalus pygmaeus: Apfelbeck, 1904: 195 (Vlorë; Berat); Apfelbeck, 1909: 298
(Berat); Hieke et al., 1986: 188 (Ibë near Krraba); Hieke & Wrase, 1988: 139; Kataev & Wrase,
2003: 380.
Material studied. “Albania 1918”, 1 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 10 s.
(NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura Erzen” /
“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1933”,
1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania
leg. Bischoff 1935”, 1 s. (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s.
(HNHM); “Albania Dajti” / “6.V.1959 leg. Xh. Murraj”, 1 s. (HNHM); Sauk, 9.VII.1961, 3 s.
(MNST); Tiranë, 10.V.1962, 1 s. (MNST); Tiranë Park, 22-31.X.1997, 2 s., R. Ward leg. (CMNH);
Tiranë Park, 16-21.II.1998, 2 s., R. Ward leg. (CMNH).
Notes. The specimens from the locality “Tiranë Park” are somewhat bigger (7.1-7.4 mm),
compared with specimens that inhabit Central Europe (cfr. Hůrka, 1996).
Species group “hospes”
56.26. Harpalus hospes Sturm, 1818
56.26.1. Harpalus hospes hospes Sturm, 1818
Material studied. “AL (Korcë) Qaf e Qarrit – 1100 m 4.V.1993, leg. Heinz u. S. Peks”, 9 s.
(MNHUB).
Notes. New species for the fauna of Albania.

108. COLEOPTERA: CARABIDAE: HARPALINAE 107
Species group “affinis”
56.27. Harpalus affinis (Schrank, 1781) [= aeneus (Fabricius, 1775); viridulus (Geoffroy,
1785); limbopunctatus Fuss, 1858]
References. Harpalus aeneus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Schatzmayr, 1943b: 103 (Lushnje). Harpalus affinis: Csiki, 1940: 219 (Gjalicë Mt.); Kataev &
Wrase, 2003: 371. Harpalus affinis ab. viridulus: Csiki, 1940: 219-220 (Gjalicë Mt.; Korab Mt.);
Harpalus affinis ab. limbopunctatus: Csiki, 1940: 220 (Gjalicë Mt.; Korab Mt.).
Material studied. Korcë, 10-20. VIII.1962, 1 m., 1 f. (MNST); North Albanian Alps, Theth,
800-1200 m, 28.V.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e
Radohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, 5-9.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 8-
16.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Notes. In addition to the material studied above, in NMW and MNHUB I have seen more
specimens, which remained unregistered.
Species group “distinguendus”
56.28. Harpalus distinguendus (Duftschmid, 1812)
56.28.1. Harpalus distinguendus distinguendus (Duftschmid, 1812)
References. Harpalus distinguendus: Apfelbeck, 1904: 192 (Vlorë; Durrës); Apfelbeck, 1909:
298 (Vlorë; Berat); Schatzmayr, 1943b: 103 (Dukati; Lushnje); Hieke & Wrase, 1988: 131.
Harpalus distinguendus distinguendus: Kataev & Wrase, 2003: 376.
Material studied. Ibë, 18.V.1960, 1 f. (MNST); Sauk, 5.III.1961, 1 m. / 23.IV.1963, 1 f. (MNST);
Rojan, 6.III.1961, 1 f. (MNST); Rodeni, 26-28.VII.1962, 1 f. (MNST); Tiranë, V.1963, 1 m. /
7.IV.1966, 5 m., 6 f. (MNST); Mal i Dajtit Mt., 10.V.1965, 1 f. (MNST); Kurbneshi, VI.1993, 1
f., P. Beron & B. Petrov leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 m., 2 f., P.
Stoev leg. (NMNHS); Saranda District, Qeparo, 4-5.V.1994, 2 s., P. Stoev leg. (NMNHS);
Saranda District, Ljukove, 3-4.VI.1994, 1 m., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994, 1
f., B. Guéorguiev leg. (NMNHS); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B.
Petrov, P. Stoev leg. (NMNHS); Librazhd - Prenjas, 750 m, 7.V.1995, 1 f., S. Golovach, B.
Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 f., S. Golovach,
B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 22-31.X.1997 / 8.II.1998 / 23-28.II.1998, 41
s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. In addition to the material studied above, in NMW and MNHUB I have seen some
more specimens, which remained unregistered.

109. 108 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
56.29. Harpalus saxicola Dejean, 1829 [= fugax Faldermann, 1836]
References. Harpalus fugax: Apfelbeck, 1907a: 498-499, 501 (Oroshi; Mal i Shenjt Mt.: top
region, 1500 m). Harpalus saxicola: Kataev & Wrase, 2003: 381.
Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 7 s. (NMW); “Rapsa 10-
18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Tartarit” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i That” / “Albania Ramadan coll. Bischoff ”, 1 s.
(MNHUB); “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 1 s. (HNHM).
Species group “oblitus”
56.30. Harpalus oblitus Dejean, 1829
56.30.1. Harpalus oblitus oblitus Dejean, 1829
References. Harpalus oblitus: Hieke & Wrase, 1988: 130. Harpalus oblitus oblitus: Kataev, 1993:
89, fig. 106; Kataev & Wrase, 2003: 379.
Notes. In NMW and MNHUB there are several specimens with labels indicating they come
from Albania. Additional labels with detailed localities revealed however that the material
originates from sites located in the valley of Sava River (Croatia and Serbia). Most probably,
Hieke & Wrase (1988: 130) have based their data for the species occurrence in Albania on the
wrongly labeled specimens in question. In spite of that Kataev (1993) displayed on map one
sure locality for the country.
Subgenus Pseudophonus Motschulsky, 1844
56.31. Harpalus griseus (Panzer, 1796)
References. Harpalus griseus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 103 (Dukati).
Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 3 s. (NMW); Rodeni, 2-
22.VII.1962, 1 m. (MNST); Tiranë, Selit e Vogel, 17.IX.1993, 2 s., S. Beshkov leg. (NMNHS);
“Albania, Alban Alps, Okol, 900 m, swept singled, 16-19.VII.1996, leg. A. Orosz”, 1 m. (HNHM).
Notes. Kataev & Wrase (2003: 385) have omitted the species for Albania. Here it is confirmed
for the country.
56.32. Harpalus rufipes (DeGeer, 1774) [= pubescens (O.F. Müller, 1776)]
References. Harpalus pubescens: Apfelbeck, 1904: 187 (Vlorë; Berat); Schatzmayr, 1943b: 103
(Dukati). Ophonus pubescens: Apfelbeck, 1909: 298 (Berat). Harpalus rufipes: Csiki, 1940: 219
(Kula Ljums); Kataev & Wrase, 2003: 385. Pseudophonus rufipes: Hieke et al., 1986: 187 (Vlorë;
North Albanian Alps near Theth; Shkodër); Hieke & Wrase, 1988: 126.
Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 1 s. (NMW); “Alban., Ricavac
Penther”, 1 s. (NMW); “Alban. Exped. Kula Ljums 14.20.VII’18”, 1 s. (NMW); Tiranë,
28.IX.1958, 1 f. (MNST); Ibë, 18.V.1960, 1 m., 1 f. (MNST); Theth, 1-8.VIII.1961, 3 f. (MNST);
Tiranë, 10.V.1961, 1 m., 2 f. (MNST); Rushkull, 6.IV.1962, 1 f. (MNST); Rodeni, 2-28.VII.1962,
2 m., 2 f. (MNST); Shkodër, 14.V.1963, 2 m. (MNST); Tiranë, 11.IX.1963, 2 f. (MNST); Shkodër,
12. VII.1965, 1 m. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m. (MNST); North Albanian Alps,

110. COLEOPTERA: CARABIDAE: HARPALINAE 109
Bogë, 1000-1100 m, 5-9.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District,
Merkurth, 11.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Tropoia, valley
of Valbona River, 900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995,
1 m., 2 f., D. Antonelli leg. (cAVT); Tiranë, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg.
(cAVT); M. Polisit, 8.VII.2004, 1 f., A. Paparistos leg. (cAVT); Malj Cojepit 1300-1400,
17.VII.2004, 1 m., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-
15.VII.2006, 3 s., leg. M. Komnenov (cSH).
Notes. Besides the material studied above, in NMW and MNHUB I have seen more specimens,
which remained unregistered.
57. Genus Ophonus Dejean, 1821
Subgenus Brachyophonus Sciaky, 1987
57.1 Ophonus krueperi Apfelbeck, 1902
References. Ophonus krueperi: Kataev & Wrase, 2003: 388.
Material included (studied D.W. Wrase). “AL (Kolonjë) Paß ca. 26 km n. Leskovik -1000 m
5.V.1993 Heinz leg.”, 1 m. (cWR; probably more specimens in cWH).
Notes. Apfelbeck published two descriptions for this taxon. First he described Ophonus krueperi
(Apfelbeck, 1902b: 543), and after than – Harpalus krueperi (Apfelbeck, 1904: 185). Incorrectly
the second description has long time been accredited to be the valid. Actually, this is incorrect
and the older description must be assigned as valid. Müller (1931: 48 sub Harpalus krueperi) and
Sciaky (1987: 69 sub Ophonus krueperi) followed the record “Chani Driskos” (Apfelbeck, 1904)
when they cited the species for Albania. However, this locality lies in Northwestern Greece.
According to D.W. Wrase (personal communication), Kataev & Wrase (2003: 388) used the
record near Leskovac, marking the species for Albania. First detailed record for the country.
Subgenus Hesperophonus Antoine, 1959
57.2. Ophonus azureus (Fabricius, 1775) [= oberthuri (Pater, 1938)]
References. Harpalus azureus: Apfelbeck, 1904: 184 (Vlorë); Csiki, 1940: 219 (Kula Ljums;
Ura i Lopez). Ophonus azureus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Apfelbeck, 1909: 298 (Vlorë); Hieke et al., 1986: 187 (North Albanian Alps near Theth; Bizë
near Shëngjergji); Hieke & Wrase, 1988: 125; Kataev & Wrase, 2003: 388. Harpalus azureus
oberthuri: Schatzmayr, 1943b: 102 (Lushnje).
Material studied. “Latif Orosi”, 1 s. (NMW); “Merdita Orosi” / “Apfelb. Alban. spt. IV.V.’05”,
2 s. (NMW); “Merdita M. Sheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Albania Kula
Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura e Loper” / “Akad. Balk.
Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Kula Ljums 18.-28.V.’18”, 12 s. (NMW);
“Alban. Exped. 1918 Bicaj 14.15.VI.”, 8 s. (NMW); “Alban. Exped.’18 Pashtrik, 4-14.7.”, 2 s.
(NMW); 53 s. from different localities (MNHUB); Ibë, 17-25.VI.1961, 1 f. (MNST); Rodens,
2-22. VII.1962, 1 s. (MNST); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. +
S. Peks”, 2 m., 1 f. (MNHUB); “AL (Korce) Qaf e Qarrit – 1100 m, 4.v.1993 leg. Heinz u.

111. 110 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Sonja Peks”, 1 m. (MNHUB); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993,
1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District, Merkurth, 11.VI.1993, 1 s., P. Beron
& B. Petrov leg. (NMNHS); Saranda District, Lyukove, 5.V.1994, 2 s., P. Stoev leg. (NMNHS);
Saranda District, Qeparo, 4-5.V.1995, 1 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove,
5.V.1995, 1 s., P. Stoev leg. (NMNHS); Dukati Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L.
Penev leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 5 s., G. Blagoev leg. (NMNHS); Tiranë
Park, 15.II.1998, 1 s., from grass and soil around tree trunks, R. Ward leg. (CMNH); Tiranë
District, Petrela, 28.XI.2000, 1 s., A. Zhalov leg. (NMNHS).
Notes. In the region of Gllavë O. azureus was found to lives sympatrically with O. similis.
57.3. Ophonus cribricollis (Dejean, 1829)
References. Harpalus cribricollis: Apfelbeck, 1904: 184-185 (Vlorë; Berat); Csiki, 1940: 219
(Shkëlzen Mt.; Kula Ljums; Gostilë). Ophonus cribricollis: Apfelbeck, 1907a: 499 (Mal i Shenjt
Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë; Berat); Hieke & Wrase, 1988: 126;
Kataev & Wrase, 2003: 389.
Material studied. “Merdita Oroshi”, 3 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt.
IV.V.’05”, 3 s. (NMW); “Pasha-Liman Albanien V.1908 H. Hopp”, 1 s. (MNHUB); “Alban.
Exped. Kula Ljums 18-28.V.1918”, 3 s. (NMW); “Devoll-Tal juni 1932” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tepelene
juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania
1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Dukati
Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District),
18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. In addition to the material studied above, in HNHM the present author has seen more
specimens from the localities “Kula Lums”, “Delvine”, “Skölsen” and “Mont Peklen”. Sciaky
(1987: 66) cited the locality “Štimlje” for Albania, but actually this place is situated in the
Kosovo Region of Serbia.
57.4. Ophonus similis (Dejean, 1829)
Material studied. “Hani Grabom Mai” / “Alban. montenegr. Grenze “Alban. Exped. 1918
Bicaj 14.15.VI.”, 1 s. (NMW); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. +
S. Peks”, 2 f. (MNHUB).
Notes. New species for the fauna of Albania.
57.5. Ophonus subquadratus (Dejean, 1829) [= meridionalis (Dejean, 1829)]
References. Ophonus meridionalis: Apfelbeck, 1909: 298 (Berat; Gramata).
Material examined. “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);
“Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M.
Komnenov leg. (cSH).
Notes. Kataev & Wrase (2003: 389) have omitted the species for Albania. Here it is confirmed
for the country.

112. COLEOPTERA: CARABIDAE: HARPALINAE 111
Subgenus Incisophonus Sciaky, 1987
57.6. Ophonus incisus (Dejean, 1829)
References. Ophonus incisus: Hieke & Wrase, 1988: 121; Kataev & Wrase, 2003: 389.
Material studied. “Murthi” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Albania Cerric”
/ “16.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS).
Notes. First detailed records for Albania.
Subgenus Metophonus Bedel, 1897
57.7. Ophonus brevicollis (Audinet-Serville, 1821) [= albanensis (Schauberger, 1933)]
References. Harpalus parallelus albanensis: Schauberger, 1933: 65 (Vlorë, loc. typ.). Harpalus
albanensis: Turin, 1981: 156. Ophonus brevicollis: Sciaky, 1987: 92-93 (Vlorë); Kataev & Wrase,
2003: 389.
57.8. Ophonus cordatus (Duftschmid, 1812)
References. Ophonus cordatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Kataev & Wrase, 2003: 389.
Material examined. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).
57.9. Ophonus laticollis Mannerheim, 1825 [= punctatulus (Duftschmid, 1812)]
References. Harpalus punctatulus: Csiki, 1940: 219 (Gjalicë Mt.). Ophonus punctulatus sic!: Hieke
et al., 1986: 187 (Mali me Gropë near Livadhet e Selitës). Ophonus laticollis: Kataev & Wrase,
2003: 390.
Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM);
“Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. Hieke & Wrase (1988: 122 sub O. punctatulus) noted this species for Albania apparently
following the record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus punctatulus). Actually this
locality is situated in the Kosovo Region of Serbia.
57.10. Ophonus melletii (Heer, 1837)
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania.
57.11 Ophonus parallelus (Dejean, 1829) [= albanicus Schauberger, 1933]
References. Ophonus parallelus albanicus: Schauberger, 1933a: 65 (Vlorë, loc. typ.). Ophonus
parallelus: Hieke & Wrase, 1988: 122; Kataev & Wrase, 2003: 390.
Material studied. “Alban. Exp.’18 Kruma 5.VI.”, 1 s. (NMW); “Maranaj” / “Albania leg.
Bischoff 1937”, 1 s. (MNHUB); “AL (Berat) Paß ca. 5 km s. Gllavë 600 m, 27.IV.1993 leg.
Heinz u. S. Peks”, 1 m. (MNHUB).

113. 112 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
57.12. Ophonus puncticeps Stephens, 1828
References.Harpaluspuncticeps:Horion,1941:212;Magistretti,1965:280;Geisthardt,1975:8.Ophonus
puncticeps: Hieke et al., 1986: 187 (North Albanian Alps near Theth); Kataev & Wrase, 2003: 390.
Material studied. “Mali i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “AL (Berat).
Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB).
57.13. Ophonus puncticollis (Paykull, 1798)
References. Ophonus puncticollis: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Apfelbeck, 1909: 298 (Berat); Kataev & Wrase, 2003: 390. Harpalus puncticollis: Müller, 1931:
50 (Tomor Mt.); Csiki, 1940: 219 (Kula Ljums); Hieke & Wrase, 1988: 122.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM);
“Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).
57.14. Ophonus rufibarbis (Fabricius, 1792)
References. Ophonus rufibarbis: Kataev & Wrase, 2003: 391.
Material studied. “Berat…Alban.”, 1 s. (MNHUB); “AL (Vlorë) Llogora-Paß (N-Seite) 800
m, 29.IV.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB).
Notes. Hieke & Wrase (1988: 124) noted the species for Albania apparently following the
record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus rufibarbis). However, this place is situated
in the Kosovo Region of Serbia. First detailed records for Albania.
57.15. Ophonus rupicola (Sturm, 1818)
References. Harpalus rupicola: Apfelbeck, 1904: 183 (Velipoja); Hieke & Wrase, 1988: 124.
Ophonus rupicola: Kataev & Wrase, 2003: 391.
Material studied. Kjari Pass, 1160 m, 1.VI.1995, 1 s., G. Blagoev leg. (NMNHS).
57.16. Ophonus schaubergerianus (Puel, 1937)
References. Ophonus schaubergerianus: Kataev & Wrase, 2003: 391.
Material studied. “Mal i Shenit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).
Notes. First detailed record for Albania.
Subgenus Ophonus s.str.
57.17. Ophonus ardosiacus (Lutshnik, 1922)
References. Ophonus ardosiacus: Kataev & Wrase, 2003: 391.
Material studied. Ibë, 4.VII.1962, 1 f. (MNST).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Durrës, Plepa,
15.VIII.1995, 2 m., D. Antonelli leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-
15.VII.2006, 3 s., leg. M. Komnenov (cSH).
Notes. First detailed records for Albania.

114. COLEOPTERA: CARABIDAE: HARPALINAE 113
57.18. Ophonus diffinis (Dejean, 1829)
References. Harpalus diffinis: Schatzmayr, 1943b: 102 (Mal i Krujes). Ophonus diffinis: Kataev &
Wrase, 2003: 391.
[Ophonus franziniorum Sciaky, 1987]
Notes. According to D.W. Wrase (personal communication), the occurrence of this winged
species in Albania is very probable. Except its proven presence in Bulgaria (Sciaky, 1987: 49;
Wrase, 1991: 14), there are reliable records for its occurrence in the Greek Ionic Islands and
European Turkey (D.W. Wrase, B. Guéorguiev, unpublished data).
57.19. Ophonus sabulicola (Panzer, 1796) [= columbinus (Germar, 1817)]
References. Ophonus sabulicola: Apfelbeck, 1909: 298 (Vlorë; Berat); Kataev & Wrase, 2003:
392. Harpalus sabulicola: Csiki, 1940: 219 (Kula Ljums). Harpalus sabulicola columbinus: Schatzmayr,
1943b: 102 (Tërpan; Dukati). Ophonus sabulicola columbinus: Hieke et al., 1986: 187 (Poliçan near
Tomor Mt.).
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); 11 s.
from different localities (MNHUB); Ibë, 4.VII.1963, 1 f. (MNST); “AL (Berat) Paß ca. 5 km s.
Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 2 s. (MNHUB); “AL (Korcë) Qaf e Qarrit –
1100 m 4.V.1993, leg. Heinz u. S. Peks”, 18 s. (MNHUB).
Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i
That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., W. Heinz & S. Peks leg. (cWR); “AL
(Kolonjë) Paß ca. 2 km ne. Leskovik -1000 m 5.V.1993 H. & S. Peks leg.”, 16 s. (cWR); 2 km N
from Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Hotel “Edva”, Radhime
(Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).
57.20. Ophonus stictus Stephens, 1828
Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 8-
15.VII.2006, 1 f., leg. M. Komnenov (cSH).
Notes. New species for the fauna of Albania.
58. Genus Parophonus Ganglbauer, 1891
Subgenus Ophonomimus Schauberger, 1923
58.1. Parophonus hirsutulus (Dejean, 1829)
References. Ophonus hirsutulus: Apfelbeck, 1902a: 99 (Velipoja near Lezhë). Harpalus hirsutulus:
Apfelbeck, 1904: 186 (Velipoja). Parophonus hirsutulus: Apfelbeck, 1907a: 495 (Buenë River);
Sciaky, 1992: 50. Ophonomimus hirsutulus: Hieke & Wrase, 1988: 141.
Material studied. “Oboti Bojana”, 1 s. (HNHM); “Oboti Schirei”, 2 s. (HNHM).
Notes. Ito (2003b: 392) has omitted the species for Albania. Here it is confirmed for the
country.

115. 114 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Subgenus Parophonus s.str.
58.2. Parophonus dejeani (Csiki, 1932)
References. Parophonus dejeani: Ito, 2003b: 393.
58.3. Parophonus laeviceps (Ménétriés, 1832) [= fallax (Peyron, 1858); planicollis Apfel-
beck, 1904 non (Dejean, 1829)]
References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë).
Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Tachyophonus fallax: Schatzmayr, 1943b: 104
(Stan-Karbunarë). Parophonus laeviceps: Hieke & Wrase, 1988: 142; Ito, 2003b: 393.
58.4. Parophonus maculicornis (Duftschmid, 1812)
References. Harpalus maculicornis: Apfelbeck, 1904: 186 (Velipoja; Vlorë; Berat). Ophonus
maculicornis: Apfelbeck, 1909: 298 (Vlorë; Berat). Trichotichnus maculicornis: Csiki, 1940: 220 (Kula
Ljums). Parophonus maculicornis: Hieke et al., 1986: 186 (Borshi near Vlorë); Hieke & Wrase,
1988: 142; Ito, 2003b: 393.
Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Pasha-Liman Albanien V.1908
H. Hopp”, 2 s. (MNHUB); “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Albania Kula
Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Elbasan, Alb. 2.IX.18 Dr. H. Priesner”,
1 s. (NMW); “Logara juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” /
“Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff Albania 1934”, 1 s.
(MNHUB); “Lum i Erzen Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murrsi-
Kara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Cerrik”
/ “18.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Lushnja, Divjaka National Park, 10.V.1995, 2 s., S.
Golovach, B. Petrov, P. Stoev leg. (cWR; NMNHS); Tiranë Park, 1.II.1998 / 5.II.1998 / 8.II.1998,
12 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).
58.5. Parophonus mendax (P. Rossi, 1790)
References. Harpalus mendax: Apfelbeck, 1904: 187 (Vlorë). Ophonus mendax: Apfelbeck, 1909:
298 (Vlorë). Tachyophonus mendax: Schatzmayr, 1943b: 104 (Stan-Karbunarë). Parophonus mendax:
Sciaky, 1992: 55; Hieke & Wrase, 1988: 143; Ito, 2003b: 393.
Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alessio, Alb. Matzenauer”,
1 s. (NMW); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 1-4.I.1998 / 11.I.1998
/ 1.II.1998, 10 s., R. Ward leg. (CMNH).
58.6. Parophonus planicollis (Dejean, 1829)
References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë).
Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Parophonus planicollis: Ito, 2003b: 393.
Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj” / “suturifer Rttr. det.
Jedlicka”, 1 f. (HNHM).

116. COLEOPTERA: CARABIDAE: HARPALINAE 115
Notes. Hieke & Wrase (1988: 142) noted that the Apfelbeck’s figures for P. planicollis (Apfelbeck,
1904: 187) refer to P. laeviceps.
59. Genus Trichotichnus A. Morawitz, 1863
Subgenus Trichotichnus s.str.
59.1. Trichotichnus laevicollis (Duftschmid, 1812)
59.1.1. Trichotichnus laevicollis laevicollis (Duftschmid, 1812)
References. Trichotichnus laevicollis: Schatzmayr, 1926: 35 (Çukall Mt.); Horion, 1941: 233; Hieke
& Wrase, 1988: 141. Trichotichnus laevicollis laevicollis: Ito, 2003b: 396.
Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Cucale” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 4 s.
(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 3 f., P. Beron & B.
Petrov leg. (NMNHS).
Subtribe STENOLOPHINA Kirby, 1837
60. Genus Acupalpus Latreille, 1829
Subgenus Acupalpus s.str.
60.1. Acupalpus brunnipes (Sturm, 1825)
Material studied. “Valona”, 1 s. (HNHM); “Oboti Bojana”, 1 s. (HNHM); “Albania Vlesh” /
“30.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM).
Notes. New species for the fauna of Albania.
60.2. Acupalpus elegans (Dejean, 1829)
References. Acupalpus elegans: Hieke & Wrase, 1988: 116; Jaeger, 1992: 224; Jaeger, 1999: 957;
Jaeger & Kataev, 2003: 398.
Material studied. “Skutari Bojana”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).
Notes. First detailed records for Albania.
60.3. Acupalpus luteatus (Duftschmid, 1812)
References. Acupalpus luteatus: Hieke & Wrase, 1988: 118; Jaeger, 1992: 225; Jaeger, 1999: 959;
Jaeger & Kataev, 2003: 398.
Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Oboti Bojana”, 1 s.
(HNHM); “Skutari Bojana”, 1 s. (HNHM); “Skutari Drin”, 1 s. (HNHM).
Additional material included (studied B. Jaeger). “Velipoja” / “Coll. Paganetti”, 1 s. (NMW).
Notes. First detailed records for Albania.

117. 116 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
60.4. Acupalpus maculatus (Schaum, 1860) [= immundus Reitter, 1900]
References. Acupalpus immundus: Apfelbeck, 1904: 206 (? Shkodër). Acupalpus maculatus: Csiki,
1940: 221 (Kula Ljums); Horion, 1941: 238; Focarile, 1959: 81; Hieke & Wrase, 1988: 118;
Jaeger, 1992: 225; Jaeger, 1999: 960; Jaeger & Kataev, 2003: 398.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM);
“Busher Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg.
Bischoff 1935”, 1 s. (MNHUB); “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);
Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).
Additional material included (studied B. Jaeger). “Elbasan, VIII.1918, Priesner”, 2 s. (MTKD);
“Elbasan, Mader”, 1 s. (MTKD); Shkodër, VIII. 1941, 12 s., Capra leg. (MHNG; cBJ);
Rrogozhinë, 1 s. (cBJ).
Notes. The record “Scutari” (Apfelbeck, 1904) is questionable because the material of Strasser
could be collected from the city of Üsküdar - in the Asian seacoast of Bosporus.
60.5. Acupalpus meridianus (Linnaeus, 1761)
References. Acupalpus meridianus: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226; Jaeger, 1999:
960; Jaeger & Kataev, 2003: 398.
Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB).
Notes. First detailed record for Albania.
60.6. Acupalpus notatus Mulsant & Rey, 1861
References. Acupalpus notatus: Horion, 1941: 238; Focarile, 1959: 82; Hieke et al., 1986: 188
(Uji Ftohte, south of Tepelenë; Ibë near Krraba); Hieke & Wrase, 1988: 118; Jaeger, 1988: 20
(Ljum, Shkumbin; Durrës: Rogozina; Elbasan; Mati: Burrelë; Tepelenë: Uji Ftohte; Ibë near
Krraba; Tiranë; Vlorë); Jaeger, 1992: 226; Jaeger, 1999: 961; Jaeger & Kataev, 2003: 398.
Material studied. “Elbasan Alb., Mader”, 3 s. (NMW; MNHUB); “Vorra, Alb. leg. Mader”, 2 s.
(NMW); “Ljum i Shkumbin, Juni 1932” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Burelle” /
“Albanialeg.Bischoff 1934”,16s.(MNHUB);“Shullaz”/“Albanialeg.Bischoff 1934”,5s.(MNHUB);
“Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 3
s.(MNHUB);TiranëPark,22.II.1998,30s.,grassesalongsmallsluggishstream,R.Wardleg.(CMNH).
Additional material included (studied B. Jaeger and A. Vigna Taglianti). “Elbasan, Mader”,
4 s. (MTKD, cBJ); “Elbasan, Priesner”, 4 s. (MTKD); Shkodër, VIII. 1941, 2 s., Capra leg.
(MHNG); Durrës, Plepa, 15.VIII.1995, 1 m., D. Antonelli leg. (cAVT).
60.7. Acupalpus paludicola Reitter, 1884 [= puncticollis Auct. nec (Coquerel, 1859)]
References. Acupalpus paludicola: Apfelbeck, 1904: 206 (Vlorë; Velipoja); Apfelbeck, 1909: 298
(Vlorë-Malket); Jaeger, 1989: 148 (Vlorë; Lum i Tiranes; Shkodër); Jaeger, 1992: 226; Jaeger,
1999: 961; Jaeger & Kataev, 2003: 399. Acupalpus puncticollis: Hieke et al., 1986: 188 (Uji Ftohte,
south of Tepelenë); Hieke & Wrase, 1988: 117.
Material studied. “Albania Avlona v. Oertzen”, 3 s. (MNHUB); “Skutari l. See-Ufer”, 1 s.
(NMW); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB; cBJ).

118. COLEOPTERA: CARABIDAE: HARPALINAE 117
Notes. All older references and collection records in the museums about A. puncticollis (Coquerel,
1859) from Albania refer to A. paludicola.
60.8. Acupalpus parvulus (Sturm, 1825) [= dorsalis (Fabricius, 1787)]
References. Acupalpus dorsalis: Apfelbeck, 1904: 206 (Velipoja; Vlorë); Apfelbeck, 1909: 298
(Vlorë); Csiki, 1940: 221 (Kula Ljums); Hieke & Wrase, 1988: 117.
Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
Tiranë Park, 8.II.1998 / 1.III.1998, 3 s., from grass and soil around tree trunks, R. Ward leg.
(CMNH).
Notes. Jaeger & Kataev (2003: 399) have omitted the species for Albania. Here it is confirmed
for the country. According to B. Jaeger and D.W. Wrase (personal communication) the records
from Albania may concern aberrant specimens of A. maculatus without darkened humeral
macula, but the only certainty for a correct identification is the study of male genitalia.
60.9. Acupalpus planicollis (Schaum, 1857)
References. Acupalpus planicollis: Apfelbeck, 1904: 205 (Velipoja); Apfelbeck, 1909: 298 (Vlorë-
Malket); Hieke & Wrase, 1988: 116; Guéorguiev & Guéorguiev, 1995: 177; Guéorguiev et al.,
1997: 20; Jaeger & Kataev, 2003: 399.
Material studied. “Liqen i Butrintes vivari” / “leg. Bischoff Albani 1934”, 1 s. (MNHUB).
Additional material included (studied B. Jaeger). “Velipoja, Mustjabeg”, 1 s. (ZSM).
60.10. Acupalpus suturalis Dejean, 1829
60.10.1. Acupalpus suturalis suturalis Dejean, 1829
References. Acupalpus suturalis: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226. Acupalpus suturalis
suturalis: Jaeger, 1999: 962; Jaeger & Kataev, 2003: 399.
Material studied. “Oboti Bojana”, 1 s. (HNHM); Tiranë Park, 22-31.X.1997, 1 s., R. Ward
leg. (CMNH); Tiranë Park, 22.II.1998, 3 s., R. Ward leg. (CMNH).
Additional material included (studied B. Jaeger). “Scutari, VIII. 1941, Capra”, 1 s. (MHNG).
Notes. First detailed records for Albania.
61. Genus Anthracus Motschulsky, 1850
61.1. Anthracus longicornis (Schaum, 1857)
References. Anthracus longicornis: Apfelbeck, 1904: 207 (Velipoja); Hieke & Wrase, 1988: 118;
Jaeger & Kataev, 2003: 400.
Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Oboti Bojana”, 4 s.
(HNHM); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB).
61.2. Anthracus quarnerensis (Reitter, 1884)
References. Anthracus quarnerensis: Apfelbeck, 1904: 208 (Velipoja); Hieke & Wrase, 1988:
119; Jaeger & Kataev, 2003: 400.

119. 118 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
62. Genus Bradycellus Erichson, 1837
Subgenus Bradycellus s.str.
62.1. Bradycellus caucasicus (Chaudoir, 1846) [= collaris (Paykull, 1798)]
References. Bradycellus collaris: Csiki, 1940: 221 (Shkëlzen Mt.).
Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM).
Notes. Jaeger & Kataev (2003: 400) have omitted the species for Albania. Here it is confirmed
for the country.
62.2. Bradycellus distinctus (Dejean, 1829)
References. Bradycellus distinctus: Jaeger, 1990: 10; Jaeger, 1999: 966; Jaeger & Kataev, 2003: 401.
Material studied. “Skutari Bojana”, 5 s. (NMW).
Additional material included (studied B. Jaeger). “Skutari Bojana”, 12 s. (MNHUB; DEI;
NMPC; cBJ).
Notes. First detailed record for Albania.
62.3. Bradycellus ganglbaueri Apfelbeck, 1904
References. Bradycellus ganglbaueri: Jaeger, 1997: 352 (Mal i Krujes); Jaeger & Kataev, 2003: 401.
62.4. Bradycellus verbasci (Duftschmid, 1812)
References. Bradycellus verbasci: Hieke & Wrase, 1988: 120; Jaeger & Kataev, 2003: 401.
Material studied. Velipoja, 19.IX.1993, 8 s., S. Beshkov leg. (NMNHS; ZISP); Tiranë Park, 1-
9.XI.1997, 1 s., R. Ward leg. (CMNH).
Additional material included (studied B. Jaeger). “Velipoja Avola v. Oertzen”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
63. genus Dicheirotrichus Jaquelin du Val, 1857
Subgenus Dicheirotrichus s.str.
63.1. Dicheirotrichus obsoletus (Dejean, 1829)
References. Dicheirotrichus obsoletus: Apfelbeck, 1904: 211 (Velipoja); Hieke & Wrase, 1988:
120; Jaeger & Kataev, 2003: 402.
Material studied. “Durres” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB).
Notes. According to B. Jaeger (personal communication) E. Schauberger identified the material
from MNHUB as D. obsoletus, but a revision of that is needed.

120. COLEOPTERA: CARABIDAE: HARPALINAE 119
64. Genus Loxoncus Schmidt-Göbel, 1846
64.1. Loxoncus procerus (Schaum, 1858)
References. Anoplogenius procerus: Apfelbeck, 1904: 202 (Velipoja near Lezhë); Apfelbeck, 1907a:
496 (Buenë River Outfall); Schatzmayr, 1943b: 105 (Lushnje). Loxoncus procerus: Wrase, 1991:
12; Kataev, 2002: 364 (Buenë River; Buenë River at Oboti); Jaeger & Kataev, 2003: 404.
Material studied. “Apfelb. Albanien”, 2 s. (MNHUB); “Apfelb. Albanien” / “Bojana
Mündung”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM); “Skutari Bojana”, 3 s. (HNHM);
“Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
65. Genus Stenolophus Dejean, 1821
Subgenus Egadroma Motschulsky, 1855
65.1. Stenolophus marginatus Dejean, 1829
References. Egadroma marginata: Apfelbeck, 1904: 204 (Vlorë; Berat); Apfelbeck, 1907a: 496
(Buenë River); Hieke et al., 1986: 188 (Durrës; Uji Ftohte, south of Tepelenë); Hieke & Wrase,
1988: 116. Stenolophus marginatus: Apfelbeck, 1909: 298 (Vlorë); Jaeger & Kataev, 2003: 405.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Oboti Bojana”, 3 s.
(HNHM); “Skumbi Elbasan Alb. 10.8.1918 Dr. H. Priesner”, 3 s. (NMW); “Ibe Erzen” /
“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 2 s.
(MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “Albania leg.
Bischoff 1937”, 1 s. (MNHUB); “Myzeqe Ebene” / “Albania leg. Bischoff 1938”, 1 s.
(MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 1 s.
(MNHUB).
Subgenus Stenolophus s.str.
65.2. Stenolophus abdominalis Gené, 1836
65.2.1. Stenolophus abdominalis persicus Mannerheim, 1844
Material studied. Tiranë Park, 22.II.1998, 1 s., shaded lake edge under leaves and wet, rotten
wood, R. Ward leg. (CMNH); Tiranë Park, 1.III.1998, 2 s., shaded lake edge under leaves and
wet, rotten wood, R. Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
65.3. Stenolophus discophorus (Fischer von Waldheim, 1823)
References. Stenolophus discophorus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988:
114; Jaeger & Kataev, 2003: 405. Acupalpus discophorus: Csiki, 1940: 221 (Kula Ljums).
Material studied. “Oboti Bojana”, 5 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp.
Csiki 1918”, 2 s. (HNHM); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB);
“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 4 s. (MNHUB); Radona, Shkiela, 21.VI.1994,
1 s., B. Guéorguiev leg. (NMNHS).

121. 120 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
65.4. Stenolophus mixtus (Herbst, 1784)
References. Stenolophus mixtus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988: 114;
Jaeger & Kataev, 2003: 406.
65.5. Stenolophus proximus Dejean, 1829
References. Stenolophus proximus: Apfelbeck, 1904: 202 (Velipoja); Hieke & Wrase, 1988: 114;
Jaeger & Kataev, 2003: 406.
65.6. Stenolophus skrimshiranus Stephens, 1828
References. Stenolophus skrimshiranus: Apfelbeck, 1904: 203 (Buenë River at Velipoja); Apfelbeck,
1909: 298 (Pasha Liman); Hieke & Wrase, 1988: 115; Jaeger & Kataev, 2003: 406.
Material studied. “Ljum i Schkumbin Juni 1932” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB).
65.7. Stenolophus teutonus (Schrank, 1781)
References. Stenolophus teutonus: Apfelbeck, 1904: 203 (Velipoja); Apfelbeck, 1909: 298 (Vlorë);
Murraj, 1968: 33 (Ibë); Hieke et al., 1986: 188 (Vlorë; Bizë near Shëngjergji); Hieke & Wrase,
1988: 115; Jaeger & Kataev, 2003: 406. Acupalpus teutonus: Csiki, 1940: 221 (Kula Ljums);
Schatzmayr, 1943b: 105 (Fieri).
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Alban. Exp.’18 Kula Ljums
26.VI.-3.VII.”, 2 s. (NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 6 s. (NMW); “Berat,
Juli”, “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Busher, Juni” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara
Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Shkumbin, Juni” / “Albania leg.
Bischoff 1932”, 1 s. (MNHUB); “Maia e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Tepelene” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “Albania leg. Bischoff
1934”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1934”, 7 s. (MNHUB); “Shullaz” /
“Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s.
(MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania
leg. Bischoff 1935”, 2 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Tiranë, 24.VI.1961, 1 f. (MNST);
Rodeni, 2-22.VII.1962, 1 s. (MNST); Tiranë Park, 22.II.1998, 1 s., shaded lake edge under
leaves and wet, rotten wood, R. Ward leg. (CMNH).
Notes. The record of Murraj (1968) is included here with reservation because of the
misidentification found in his collection.

122. COLEOPTERA: CARABIDAE: HARPALINAE 121
Tribe LEBIINI Bonelli, 1810
Subtribe CYMINDIDINA Laporte, 1834
66. Genus Cymindis Latreille, 1806
Subgenus Cymindis s.str.
66.1. Cymindis angularis Gyllenhal, 1810
Material studied. “Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania
leg. Bischoff 1935”, 15 s. (MNHUB); “Taja” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB);
“Gjallica” / “Albania leg. Bischoff 1936”, 20 s. (MNHUB); “Taja” / “Albania leg. Bischoff
1936”, 2 s. (MNHUB).
Notes. New species for the faunas of the Balkan Peninsula and Albania. Most likely, the
populations from the high mountains in North Albania are glacial relics.
66.2. Cymindis axillaris (Fabricius, 1794)
66.2.1. Cymindis axillaris axillaris (Fabricius, 1794)
References. Cymindis axillaris: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);
Klaptocz, 1910: 399 (Oroshi); Hieke et al., 1986: 190 (North Albanian Alps near Theth).
Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Maja e
Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Mali i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Koritnik”
/ “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan
coll. Bischoff”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Boge” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania
Mueller leg. 1939”, 9 s. (MNHUB); “Iballe” / “Albania Mueller leg. 1939”, 1 s. (MNHUB).
Notes. Kabak (2003: 411) has omitted the species for Albania. Here it is confirmed for the
country.
66.2.2. Cymindis axillaris adriatica J. Müller, 1921
References. Cymindis axillaris adriatica: Müller, 1921a: 139 (Shkodër).
66.3. Cymindis coadunata Dejean, 1825
66.3.1. Cymindis coadunata coadunata Dejean, 1825
Material studied. “Çafa Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë”
/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 6
s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB).
Notes. New species for the fauna of Albania.
66.4. Cymindis humeralis (Geoffroy, 1785)
References. Cymindis humeralis: Csiki, 1940: 222 (Shkëlzen Mt.); Hieke & Wrase, 1988: 161.

123. 122 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s.
(NMW); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban. Exp.’18
Pashtrik, 4-14.7.”, 1 s. (NMW); 68 s. from different localities (MNHUB).
Notes. Kabak (2003: 413) has omitted the species for Albania. Here it is confirmed for the
country.
66.5. Cymindis lineata (Quensel, 1806)
References. Cymindis lineata: Apfelbeck, 1904: 343 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt
Mt.: top region, 1500 m); Hieke & Wrase, 1988: 162; Kabak, 2003: 413.
References. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Rapsa 10-18.V.”
/ “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Alban. Exped. Kula Ljums 18.-
28.V.1918”, 2 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 3 s. (NMW); “Mal i Dajtit Tirane”
/ “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 1 s.
(MNHUB); “Mal i Scheinjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tomor” / “leg.
Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll.
Bischoff”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Borsne”
/ “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “AL (Korcë) Vlacisht n. Korcë, 800 m 4.V.1993,
leg. Heinz u. S. Peks (ex. coll H. Peks)”, 1 s. (MNHUB).
66.6. Cymindis lineola L. Dufour, 1820
References. Cymindis axillaris lineola: Apfelbeck, 1907a: 495 (Buenë River).
Material studied. “Latif Skutari”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 2 s. (HNHM).
Notes. Kabak (2003: 414) has omitted the species for Albania. Here it is confirmed for the
country.
? Cymindis pindicola Apfelbeck, 1901
Material studied. “Neres” / “Apfelb. 1906 Albania or.”, 1 s. (HNHM).
Notes. Apfelbeck described this species from the mountains of Peristeri, Veluchi and Oxya
situated in Northwestern and Central Greece. The specimen from “Neres” has been found to
be conspecific with a single syntype of C. pindicola labeled “Veluchi Gb. Stralcia” / “Coll.
Apfelbeck” (HNHM). However, I failed to find a place named “Neres” in Albania or in the
adjacent regions. Because of uncertain localization, this record remains questionable for Albania.
66.7. Cymindis scapularis Schaum, 1857
66.7.1. Cymindis scapularis scapularis Schaum, 1857
References. Cymindis scapularis var.: Apfelbeck, 1909: 298 (Vlorë-Fieri).
Notes. Kabak (2003: 411) has omitted the species for Albania. Actually, it needs further
confirmation for the country.
Subgenus Menas Motschulsky, 1864
Notes. New subgenus for the fauna of Albania.

124. COLEOPTERA: CARABIDAE: HARPALINAE 123
66.8. Cymindis miliaris (Fabricius, 1801)
Material studied. “Albania Rapsa Dr. Grabowski”, 3 s. (HNHM).
Notes. New species for the fauna of Albania.
Subgenus Tarulus Bedel, 1906
66.9. Cymindis vaporariorum (Linnaeus, 1758)
References. Cymindis vaporariorum: Hieke & Wrase, 1988: 162.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s.
(NMW); “Korab” / “Albania leg. Bischoff 1935”, 20 s. (MNHUB); “Maja e Jezercoë” / “Albania
leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB);
“Korab” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).
Notes. Kabak (2003: 418) has omitted the species for Albania. Here it is confirmed for the
country with first detailed records.
Subtribe DROMIINA Bonelli, 1810
67. Genus Dromius Bonelli, 1810
Notes. New genus for the fauna of Albania.
Subgenus Dromius s.str.
Notes. New subgenus for the fauna of Albania.
67.1. Dromius quadrimaculatus (Linnaeus, 1758)
Material studied. “Elbasan, Alb. 19.IX.18. Dr. H. Priesner”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
68. Genus Microlestes Schmidt-Göbel, 1846
68.1. Microlestes corticalis (L. Dufour, 1820) [= escorialensis (Brisout de Barneville, 1885)]
References. Microlestes plagiatus escorialensis: Holdhaus, 1904: 335 (Velipoja; Vlorë). Microlestes
corticalis escorialensis: Schatzmayr, 1943b: 112 (Lushnje). Microlestes corticalis: Hieke & Wrase, 1988:
160; Kabak, 2003: 422.
Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB).
68.2. Microlestes fissuralis (Reitter, 1901)
References. Microlestes fissuralis: Holdhaus, 1904: 332 (Velipoja; “Ardenica” near Vlorë); Csiki,
1940: 221 (Gjalicë Mt.); Horion, 1941: 343; Hieke et al., 1986: 190 (Kula e Lumës near Kukës);
Hieke & Wrase, 1988: 159; Kabak, 2003: 422.

125. 124 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica”
/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”,
4 s. (MNHUB); Librazhd Prenjas, 750 m, 7.V.1995, 1 s., L. Penev leg. (NMNHS).
68.3. Microlestes luctuosus Holdhaus, 1904
68.3.1. Microlestes luctuosus luctuosus Holdhaus, 1904
References. Microlestes exilis luctuosus: Holdhaus, 1904: 331 (Berat). Microlestes luctuosus: Hieke &
Wrase, 1988: 159. Microlestes apterus: Wrase, 1991: 18.
Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Kjori” / “Albania leg.
Bischoff 1932”, 2 s. (MNHUB); “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s.
(MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. According to Wrase (1991) the material cited as M. exilis Schmidt-Göbel, 1846 and M.
exilis luctuosus (Apfelbeck, 1904: 330-331) may concern either M. apterus or M. luctuosus. I refer
it provisionally to the latter species, which was proved to inhabit the country. Kabak (2003:
423) has omitted or M. luctuosus for Albania. Here, the species is confirmed for the country.
68.4. Microlestes maurus (Sturm, 1827)
68.4.1. Microlestes maurus maurus (Sturm, 1827)
References. Microlestes maurus: Holdhaus, 1904: 332 (Velipoja; Vlorë); Hieke & Wrase, 1988:
159; Kabak, 2003: 423.
68.5. Microlestes minutulus (Goeze, 1777)
References. Blechrus minutulus: Apfelbeck, 1909: 298 (Vlorë; Berat).
Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Mal i Krujes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the
country.
68.6. Microlestes negrita (Wollaston, 1854)
68.6.1. Microlestes negrita apfelbecki Holdhaus, 1904
References. Microlestes minutulus apfelbecki: Holdhaus, 1904: 330 (Velipoja). Microlestes negrita:
Hieke & Wrase, 1988: 158.
Material studied. “Cafa Liqenit Iballe” / “Albania leg. Mueller 1939”, 1 s. (MNHUB).
Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the
country.
68.7. Microlestes plagiatus (Duftschmid, 1812)
References. Microlestes plagiatus: Apfelbeck, 1907a: 495 (Buenë River); Csiki, 1940: 221 (Korab Mt.).
Notes. Kabak (2003: 423) has omitted the species for Albania. Actually it needs further
confirmation for the country.

126. COLEOPTERA: CARABIDAE: HARPALINAE 125
69. Genus Paradromius Fowler, 1887
Subgenus Manodromius Reitter, 1905
69.1. Paradromius linearis (Olivier, 1795)
69.1.1. Paradromius linearis linearis (Olivier, 1795)
References. Dromius linearis: Hieke & Wrase, 1988: 155.
Material studied. “Vorra, Alb. 25.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Bushek juni” /
“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durres juni” / “Albania leg. Bischoff 1932”, 2
s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranës” /
“leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Lum i Tiranës” / “Albania leg. Bischoff 1936”,
3 s. (MNHUB); Tiranë Park, 8.II.1998, 2 s., R. Ward leg. (CMNH).
Notes. Kabak (2003: 424) has omitted the species for Albania. Here P. linearis is confirmed for
the country with first detailed records.
70. Genus Philorhizus Hope, 1838
subgenus Philorhizus s.str.
70.1. Philorhizus notatus (Stephens, 1827)
References. Dromius notatus: Hieke & Wrase, 1988: 156.
Material studied. “Oboti Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM); “Tomor” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. Kabak (2003: 426) has omitted the species for Albania. Here it is confirmed for the
country with first detailed records.
70.2. Philorhizus sigma (P. Rossi, 1790)
References. Dromius sigma: Apfelbeck, 1904: 337 (Velipoja); Hieke & Wrase, 1988: 157; Kabak,
2003: 426.
Subtribe LEBIINA Bonelli, 1810
71. Genus Lebia Latreille, 1802
Subgenus Lamprias Bonelli, 1810
Notes. New subgenus for the fauna of Albania.
71.1. Lebia cyanocephala (Linnaeus, 1758)
71.1.1. Lebia cyanocephala cyanocephala (Linnaeus, 1758)
Material studied. “Skutari Drin”, 1 s. (HNHM); “Tirane” / “leg. Bischoff Albania 1934”, 1 s.
(MNHUB).

127. 126 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. The mention of Hieke & Wrase (1988: 153) for Albania is apparently result of a
following the record “Prisren” (Apfelbeck, 1904: 323). However, this locality is situated in the
Kosovo Region of Serbia. Kabak (2003: 427) has most likely followed one of these sources,
too. Since there is no previous reliable record from Albania, L. cyanocephala is treated here as
new species for the fauna of Albania.
Subgenus Lebia s.str.
71.2. Lebia cruxminor (Linnaeus, 1758)
71.2.1. Lebia cruxminor cruxminor (Linnaeus, 1758)
References. Lebia cruxminor: Hieke et al., 1986: 190 (North Albanian Alps near Theth; Kula e
Lumës near Kukës); Hieke & Wrase, 1988: 153.
Material studied. “Skutari Drin”, 1 s. (HNHM); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”,
1 s. (NMW); “Cucale Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Kjori” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albanien-Exp.
DEI Nordalban. Thethi, 1.-4.VIII.61”, 1 s. (MNHUB).
Notes. Kabak (2003: 428) has omitted the species for Albania. Here L. cruxminor is confirmed
for the country.
71.3. Lebia humeralis Dejean, 1825
References. Lebia humeralis: Hieke & Wrase, 1988: 154.
Material studied. “Albania Djakova” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban.
Exped. Gjalica Ljums 17-26.VI.’18”, 1 s. (NMW); “Lum i Tiranës” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. Kabak (2003: 429) has omitted the species for Albania. Here it is confirmed for the
country with first detailed records.
71.4. Lebia scapularis (Geoffroy, 1785)
71.4.1. Lebia scapularis scapularis (Geoffroy, 1785)
References. Lebia scapularis: Hieke & Wrase, 1988: 154.
Material studied. “Serandë Piqeros” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. Kabak (2003: 429) has omitted the species for Albania. Here L. scapularis is confirmed
for the country with a first detailed record.
71.5. Lebia trimaculata (Villers, 1789)
Material studied. 12 km NE Erseka, 900 m, 12.V.1995, 2 s., L. Penev leg. (NMNHS).
Notes. New species for the fauna of Albania.

128. COLEOPTERA: CARABIDAE: HARPALINAE 127
Subtribe LIONYCHINA Jeannel, 1948
72. Genus Apristus Chaudoir, 1846
72.1. Apristus reticulatus Schaum, 1857
Material studied. “Mesi Kiri-Ufer” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. New species for the fauna of Albania.
72.2. Apristus subaeneus Chaudoir, 1846
References. Apristus subaeneus: Apfelbeck, 1907a: 495 (Liqen i Shkodrës); Müller, 1921a: 139;
Müller, 1924: 139; Hieke & Wrase, 1988: 161.
Material studied. “Skutari Drin”, 1 s. (NMW); “Skutari Vraka”, 6 s. (HNHM); “Skutari Vraka” /
“Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW).
Notes. Kabak (2003: 431) has omitted the species for Albania. Here it is confirmed for the country.
73. Lionychus Wissmann, 1846
Notes. New genus for the fauna of Albania.
Subgenus Lionychus s.str.
Notes. New subgenus for the fauna of Albania.
73.1. Lionychus maritimus Fairmaire, 1862
Material studied. “Himara juni” / “Albania leg. Bischoff 1932”, 17 s. (MNHUB).
Notes. New species for the faunas of the Balkan Peninsula and Albania.
73.2. Lionychus quadrillum (Duftschmid, 1812)
Material studied. “Medua, Alb. 10.5.914”, 1 s. (HNHM).
Notes. New species for the fauna of Albania.
74. Genus Syntomus Hope, 1838
74.1. Syntomus impressus (Dejean, 1825)
74.1.1. Syntomus impressus impressus (Dejean, 1825)
References. Metabletus impressus: Apfelbeck, 1907a: 495 (Buenë River). Syntomus impressum sic!:
Hieke & Wrase, 1988: 158.
Material studied. “Albanien Avlona v. Oertzen.”, 2 s. (MNHUB); Saranda District, Ljukove,
3-4.VI.1994, 5 s., G. Blagoev leg. (NMNHS).
Notes. Kabak (2003: 432) has omitted the species for Albania. Here it is confirmed for the
country.

129. 128 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
74.2. Syntomus obscuroguttatus (Duftschmid, 1812)
References. Metabletus obscuroguttatus: Apfelbeck, 1904: 327 (Vlorë; Velipoja); Apfelbeck, 1909:
298 (Vlorë). Syntomus obscuroguttatus: Hieke & Wrase, 1988: 157; Kabak, 2003: 433.
Material studied. “Oboti Bojana”, 4 s. (HNHM); “Skutari Bojana”, 1 s. (HNHM); “Logara”
/ “leg. Bischoff Albania 1934”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg.
(NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Radona,
Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 15.II.1998, 320 s., from
grass and soil around tree trunks, R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2
m., D. Antonelli leg. (cAVT).
74.3. Syntomus pallipes (Dejean, 1825)
References. Metabletus pallipes: Apfelbeck, 1909: 298 (Vlorë). Syntomus pallipes: Hieke & Wrase,
1988: 157.
Material studied. “Okol Shkalla” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Notes. Kabak (2003: 433) has omitted the species for Albania. Here it is confirmed for the country.
74.4. Syntomus truncatellus (Linnaeus, 1761)
References. Metabletus truncatellus: Csiki, 1940: 221 (Korab Mt.). Syntomus truncatellus: Hieke &
Wrase, 1988: 158.
Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania
Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Korab” / “Albania leg. Bischoff
1935”, 3 s. (MNHUB).
Notes. Kabak (2003: 433) has omitted the species for Albania. Here it is confirmed for the country.
Tribe LICININI Bonelli, 1810
Subtribe LICININA Bonelli, 1810
75. Genus Badister Clairville, 1806
Subgenus Badister s.str.
75.1. Badister bullatus (Schrank, 1798)
Material studied. “Scutari Bojana”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”,
1 s. (NMW).
Notes. New species for the fauna of Albania.
75.2. Badister unipustulatus Bonelli, 1813
References. Badister unipustulatus: Apfelbeck, 1904: 163 (Velipoja); Murraj, 1968: 33 (Tiranë);
Hieke & Wrase, 1988: 151; Baehr, 2003c: 440.

130. COLEOPTERA: CARABIDAE: HARPALINAE 129
Material studied. “Oboti Bojana”, 2 s. (HNHM).
Notes. The record of Murraj (1968) is included here with reservation due to misidentifications
found in his collection.
Subgenus Baudia Ragusa, 1884
75.3. Badister dilatatus Chaudoir, 1837
References. Badister dilatatus: Apfelbeck, 1904: 164 (Velipoja); Hieke & Wrase, 1988: 151;
Baehr, 2003c: 441.
75.4. Badister peltatus (Panzer, 1796)
75.4.1. Badister peltatus peltatus (Panzer, 1796)
References. Badister peltatus peltatus: Baehr, 2003c: 441.
76. Genus Licinus Latreille, 1802
Subgenus Licinus s.str.
76.1. Licinus cassideus (Fabricius, 1792)
76.1.1. Licinus cassideus cassideus (Fabricius, 1792) [= latus Dejean, 1831]
References. Licinus cassideus latus: Magistretti, 1965: 263; Hieke & Wrase, 1988: 150. Licinus cassideus
cassideus: Baehr, 2003c: 442.
Material studied. “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” /
“Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Tomor-Tojk juli 1932” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB);
“Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4
s. (MNHUB); “Boge Aufstieg z. Rab” / “Albania leg. Mueller 1939”, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District),
18.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. Csiki (1931: 911-912) treated dohrni, graecus and submarginatus as synonyms of cassideus.
Here, I follow the arrangement of Baehr (2003c: 442). In spite of that, a revision of the
Balkan taxa close to L. cassideus is needed. First detailed records for Albania.
76.1.2. Licinus cassideus dohrni Fairmaire, 1863
References. Licinus cassideus dohrhi: Fairmaire, 1863: 131 (Albania, loc. typ.); Reitter, 1900: 150;
Apfelbeck, 1904: 162; Baehr, 2003c: 442.
Material studied.“MajaePapingut”/“Albania1936leg.Ramadancoll.Bischoff”, 1s. (MNHUB).
76.2. Licinus graecus Apfelbeck, 1901 [= submarginatus J.R. Sahlberg, 1903]
References. Licinus graecus: Apfelbeck, 1901a: 431 (Berat); Apfelbeck, 1909: 298 (Berat). Licinus
cassideus graecus: Apfelbeck, 1904: 162 (Berat). Licinus submarginatus: Schatzmayr, 1943b: 100
(Tërpan); Baehr, 2003c: 442.

131. 130 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Apfelbeck published two descriptions of L. graecus. First he described L. graecus
(Apfelbeck, 1901a: 431), and after that L. cassideus graecus (Apfelbeck, 1904: 162). Incorrectly
the second taxon has been accredited to be the valid, which create a future complication.
In 1904, Apfelbeck (1904: 399, 402 note 1) synonymized L. submarginatus J.R. Sahlberg,
1903 with L. cassideus graecus Apfelbeck, 1904, not with L. graecus Apfelbeck, 1901. The
only explanation of this fact is that this outstanding specialist seemed to have forgotten
his earlier description. Later, the subsequent authors (Schatzmayr, 1943b; Baehr, 2003c)
followed this view, but treated L. submarginatus as valid species, and L. cassideus graecus
Apfelbeck, 1904 – for its synonym. I believe, however, that both Licinus - species described
by Apfelbeck are identical based on identical type localities. According to the rules of
ISZN (1999) an original description of a taxon is the oldest one. Therefore, the following
nomenclatorial act is proposed: Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn.
nov. of Licinus (Licinus) graecus Apfelbeck, 1901.
76.3. Licinus silphoides (P. Rossi, 1790)
References.Licinussilphoides:Apfelbeck,1909:298(Vlorë-Fieri);Schatzmayr,1943b:100(Karbunara
e Madhe; Stan-Karbunarë); Focarile, 1959: 76; Hieke & Wrase, 1988: 150; Baehr, 2003c: 442.
Subgenus Neorescius Bedel, 1906
76.4. Licinus oertzeni Reitter, 1889
76.4.1. Licinus oertzeni oertzeni Reitter, 1889
References. Licinus oertzeni: Apfelbeck, 1907a: 499, 501,503 (Mal i Shenjt Mt.: top region, 1500 m).
76.4.2. Licinus oertzeni albanicus J. Müller, 1932
References. Licinus oertzeni albanicus: Müller, 1932: 209 (Tomor Mt. near Berat, loc. typ.;
Northwestern Greece, “Buč-Gebirge”, loc. typ.); Baehr, 2003c: 443.
Material studied. “Merdita Orochi”, 1 s. (HNHM); “Maja e Tartarit” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” /
“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Çafa Kjores” / “Albania leg. Bischoff 1935” /
“Licin. Oertzeni albanicus m det. J. Müller”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shen
Ili” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania
leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. The distinction of the two subspecies of L. oertzeni s.l. is very easy due to their clear
external differences. Probably both taxa deserve species status.
Tribe ODACANTHINI Laporte, 1834
Notes. New tribe for the fauna of Albania.
77. Genus Odacantha Paykull, 1798
Notes. New genus for the fauna of Albania.

132. COLEOPTERA: CARABIDAE: HARPALINAE 131
Subgenus Odacanta s.str.
Notes. New subgenus for the fauna of Albania.
77.1. Odacantha melanura (Linnaeus, 1767)
Material studied. “Skutari Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM).
Notes. New species for the fauna of Albania.
Tribe OODINI LaFerté-Sénectère, 1851
78. Genus Oodes Bonelli, 1810
78.1. Oodes gracilis A. Villa & G.B. Villa, 1833
References. Oodes gracilis: Apfelbeck, 1904: 159 (Velipoja); Focarile, 1959: 75; Hieke & Wrase,
1988: 150; Bousquet, 2003c: 445.
Material studied. Shkodër District, Mali Tarabosh Mt., 100 m, 24.XI.2000, 3 m., limestone
terrain, A. Zhalov leg. (NMNHS).
78.2. Oodes helopioides (Fabricius, 1792)
78.2.1. Oodes helopioides helopioides (Fabricius, 1792)
References. Oodes helopioides: Apfelbeck, 1904: 159 (Velipoja near Lezhë); Hieke & Wrase,
1988: 150; Bousquet, 2003c: 445.
Tribe OMPHREINI Ganglbauer, 1891
79. Genus Omphreus Dejean, 1828
Notes. This genus needs revision. The author’s examinations revealed only a small number of
specimens of the most species and these in bad condition (excluding O. morio s.l.) in the largest
Central European collections (DEI; HNHM; NMW). Contrary, in MNHUB the collection of
Omphreus is better represented.
Subgenus Omphreus s.str.
79.1. Omphreus lonai Winkler, 1933
References. Omphreus lonai: Winkler, 1933b: 118 (Llogora, 1200 m, southeast of Vlorë, loc.
typ.); Bousquet, 2003d: 446.
Material studied. “Llogora” / “Albania leg. Bischoff 1935”, 1 m. (MNHUB); “Llogora” /
“Albania leg. Bischoff 1936”, 3 f. (MNHUB).

133. 132 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
79.2. Omphreus morio Dejean, 1828
79.2.1. Omphreus morio albanicus Apfelbeck, 1906
References. Omphreus albanicus: Apfelbeck, 1906: 1666-1667 (Merdita: Mal i Shenjt Mt., loc.
typ.; Zepë Mt., loc. typ.); Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck,
1918c: 160, 165 (Mal i Shenjt Mt.; Zepë Mt.; North Albanian Alps, 1300-2000 m); Hieke,
1981: 82. Omphreus morio: Anonymous, 1907: 286. Omphreus morio albanicus: Winkler, 1933b: 117
(Mal i Dajtit Mt.; Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Peshtrik Mt.); Drovenik & Peks,
1994: 75 (Peshtrik Mt.); Drovenik & Peks, 1999: 87 (Peshtrik Mt.).
Material studied. “Merdita M. Scheit” / “Coll. Apfelbeck”, 4 s. (HNHM); “Merdita M. Scheit”,
3 s. (NMW); “Merdita Zebia”, 6 s. (HNHM; NMW; MNHUB); “Merdita Munela”, 1 s. (HNHM);
“Oroši (Lat.) Alban. spt.” / “Zebja”, 2 s. (DEI; MNHUB); “Merdita Munela” / “Wgth., 1906
Merdita”, 1 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Alban.
Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exped. Gjalica Ljums 17.-26.VI’18”,
2 s. (NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 2 m., 1 f. (MNHUB); “Mal i
Krujes” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB); “Gjallica” / “Albania leg. Bischoff
1936”, 1 m. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 m. (MNHUB); “Mal
i Krujes” / “Albania leg. Bischoff 1939”, 1 f. (MNHUB); Reshen District, near Merkurth,
920-980 m, 11.VI.1993, 1 s., grassland, P. Beron & B. Petrov leg. (NMNHS); Mal i Dajtit Mt.,
20 km east of Tiranë, 1000 m, 9.V.1995, 3 s., L. Penev leg. (NMNHS).
Notes. Bousquet (2003d: 446) has omitted O. morio for Albania. Here the species is confirmed
for the country.
79.2.2. Omphreus morio malissorum Winkler, 1933
References. Omphreus morio malissorum: Winkler, 1933b: 117 (North Albanian Alps: Maja e
Jezercë, loc. typ.; Starca; Çukall Mt.).
Material studied. “Maja e Jezerce Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1
syntype (NMW); “Maja e Jezercoë” / “Albania leg. Bischoff 1931”, 1 syntype (MNHUB);
“Cukale Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1 syntype (NMW); “Cucale
Geb.” / “Albania leg. Bischoff 1931”, 3 syntypes (MNHUB); “Maja e Jezercoë” / “Albania
leg. Bischoff 1935”, 2 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Cucale” / “Albania leg. Bischoff 1936”, 9 s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff
1936”, 3 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj”
/ “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg.
1939”, 3 s. (MNHUB); “M. Raba Massiv” / “Albania Mueller leg. 1939”, 6 s. (MNHUB);
North Albanian Alps, 1400-1800 m, 1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).
79.3. Omphreus ravasinii J. Müller, 1923
References. Omphreus ravasinii: Müller, 1923: 111-112 (Tomor Mt., alpine zone, loc. typ.);
Winkler, 1933b: 118 (Tomor Mt.); Bousquet, 2003d: 446.
Material studied. “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff 1932”, 1 f. (MNHUB).
! Omphreus ravasinii ssp.
Material studied. “Korab” / “Albania leg. Bischoff 1936”, 1 f. (MNHUB).
Notes. It seems this specimen belong to a vicarious form close to O. ravasinii. Unfortunately
its abdomen is destroyed and further material is needed to specify its right status.

134. COLEOPTERA: CARABIDAE: HARPALINAE 133
Tribe PANAGAEINI Bonelli, 1810
80. Genus Panagaeus Latreille, 1802
Subgenus Panagaeus s.str.
80.1. Panagaeus cruxmajor (Linnaeus, 1758) [= schaumi Ganglbauer, 1892]
References. Panagaeus cruxmajor ab. schaumi: Apfelbeck, 1904: 153 (Velipoja near Lezhë). Panagaeus
cruxmajor: Hieke & Wrase, 1988: 152; Baehr, 2003d: 447.
Material studied. “Scutari Drin”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM).
Tribe PLATYNINI Bonelli, 1810
81. Genus Agonum Bonelli, 1810
Subgenus Agonum s.str.
81.1. Agonum afrum (Duftschmid, 1812)
Material studied. Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 7 s., shaded lake edge
under leaves and wet, rotten wood, R. Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
81.2. Agonum carbonarium Dejean, 1828
81.2.1. Agonum carbonarium carbonarium Dejean, 1828 [= alpestre (Heer, 1838)]
References. Agonum alpestre: Schmidt, 1995: 168 (Korab Mt.).
Notes. Bousquet (2003e: 448 sub A. alpestre) has omitted A. carbonarium for Albania. The
species occurrence in the central area of the Balkans is interesting from the biogeographic
point of view since the Albanian locality is distant from the main Western European range of
the species, as well as from its Asian shelters.
81.3. Agonum duftschmidi J. Schmidt, 1994 [= moestum (Duftschmid, 1812)]
References. Platynus viduus var. moestus: Apfelbeck, 1904: 292 (Velipoja). Platynus viduus moestus:
Apfelbeck, 1907a: 495 (Liqen i Shkodrës). Platynus moestus: Apfelbeck, 1909: 297 (Vlorë). Agonum
moestum: Hieke & Wrase, 1988: 86. Agonum duftschmidi: Bousquet, 2003e: 450.
81.4. Agonum extensum Ménétriés, 1849 [= binaghii Burlini, 1939]
References. Agonum monachum var. binaghii: Burlini, 1939: 60 (Vlorë). Agonum extensum: Schatzmayr,
1943b: 111 (Skrofotina).
Material studied. “Apfelb. Avlona”, 1 s. (HNHM); “Myzeqe Ebene” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB).
Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for the country.

135. 134 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
81.5. Agonum gisellae Csiki, 1931 [= angustatum Dejean, 1828]
References. Agonum angustatum: Hieke & Wrase, 1988: 86; Schmidt, 1994: 27 (Buenë River
near Shkodër). Agonum gisellae: Bousquet, 2003e: 450.
81.6. Agonum hypocrita (Apfelbeck, 1904)
References. Agonum hypocrita: Csiki, 1940: 218 (Korab Mt.).
Material studied. “Scutari l. See-Ufer”, 3 s. (HNHM); “Murrsi-Kara Alibey” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Bushnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Mamurash” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB).
Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for
the country.
81.7. Agonum longicorne Chaudoir, 1846 [= holdhausi (Apfelbeck, 1904)]
References. Platynus holdhausi: Apfelbeck, 1904: 294 (Velipoja); Apfelbeck, 1907a: 495 (Buenë
River). Agonum holdhausi: Hieke & Wrase, 1988: 87. Agonum longicorne: Schmidt, 1994: 32 (Buenë
River near Oboti); Bousquet, 2003e: 450.
Material studied. “Latif Pulaj”, 2 s. (HNHM); “Oboti Wald” / “Apfelb. Alban. spt. IV.V.’05”,
1 paralectotype of Platynus holdhausi (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1938”,
1 s. (MNHUB).
Notes. In NMW I found a paralectotype not listed by Schmidt (1994: 31) among the type
material of A. longicorne, though the author included the record of this material (Oboti) in his
list of species distribution.
81.8. Agonum marginatum (Linnaeus, 1758)
References. Agonum marginatum: Hieke & Wrase, 1988: 85.
Material studied. “Skutari Vraka” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Maja e Konispol
Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff
1936”, 2 s. (MNHUB); Tiranë Park, 8.II.1998, shaded lake edge under leaves and wet, rotten
wood and 22.II.1998, from grasses along small sluggish stream, 22 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë
and Gjirokastër), 350 m, 23.IV.2003, 1 m., P. Crucitti & C. Cavalieri leg. (cAVT).
Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for
the country with first detailed records.
81.9. Agonum muelleri (Herbst, 1784)
Material studied. “Skutari l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Alban.
Vermosa Penther”, 3 s. (NMW).
Notes. The mention of Hieke & Wrase (1988: 85) for Albania is apparently a result of following
the record “Prisren” (Apfelbeck, 1904: 290 sub Platynus muelleri). However, currently this locality

136. COLEOPTERA: CARABIDAE: HARPALINAE 135
is situated in the Kosovo Region of Serbia. Bousquet (2003e: 451) has most likely followed
one of these sources, too. Since there is no previous reliable record from Albania, A. muelleri
is treated here as new species for the fauna of Albania.
81.10. Agonum permoestum Puel, 1938
References. Agonum permoestum: Puel, 1938: 194 (Velipoja); Schmidt, 1994: 25 (Buenë River
near Shkodër; Bushnesh; Mamurash; Murrsi-Kara Alibey; Buenë River near Oboti; Rushkull;
Liqen i Shkodrës; Sukthi near Gurse); Bousquet, 2003e: 451.
Material studied. “Scutari l. See-Ufer”, 1 s. (HNHM); “Skutari l. See-Ufer” / “Apfelb. Alban.
spt. IV.V.’05”, 8 s. (NMW); “Oboti Bojana”, 1 s. (HNHM).
81.11. Agonum sexpunctatum (Linnaeus, 1758)
References. Agonum sexpunctatum: Csiki, 1940: 218 (Korab Mt.); Hieke et al., 1986: 189 (Lurë
e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 84.
Material studied. “Alban. Vermosa Penther”, 1 s. (NMW); “Ricavac 1300 m, 25.VI.-2.VII.” /
“Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Albania Tirana” / “3.VI.1958 leg. Xh.
Murraj”, 3 s. (HNHM).
Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for
the country.
81.12. Agonum sordidum Dejean, 1828
81.12.1. Agonum sordidum sordidum Dejean, 1828
References. Platynus sordidus: Apfelbeck, 1904: 290 (Velipoja; Vlorë); Apfelbeck, 1909: 297
(Vlorë). Agonum sordidum: Schatzmayr, 1943b: 111 (Stan-Karbunarë); Hieke et al., 1986: 189
(Poliçan near Tomor Mt.); Hieke & Wrase, 1988: 85.
Material studied. “Elbasan, Alb. IX… Dr. H. Priesner”, 1 s. (NMW); “Liquen i Butrintes
Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff
1935”, 2 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis
Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB).
Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for the country.
81.13. Agonum viridicupreum (Goeze, 1777)
80.13.1. Agonum viridicupreum viridicupreum (Goeze, 1777)
References. Platynus viridicupreus: Apfelbeck, 1904: 290 (Velipoja). Agonum viridicupreum: Hieke
et al., 1986: 189 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 85.
Material studied. “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);
“Tiranë” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania
leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s.
(MNHUB); “Han-Hot” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Sukthi bis Gurse” /
“Albania leg. Bischoff 1938”, 1 s. (MNHUB); “Cafa Liqenit Iballe” / “Albania Mueller leg.
1939”, 5 s. (MNHUB); Tiranë Park, 11.I.1998 / 1.III.1998 / 8.III.1998, 4 s., shaded lake edge
under leaves and wet, rotten wood, R. Ward leg. (CMNH).

137. 136 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Europhilus Chaudoir, 1859
81.14. Agonum antennarium (Duftschmid, 1812)
References. Platynus antennarius: Apfelbeck, 1904: 295 (Velipoja). Agonum antennarium: Hieke
& Wrase, 1988: 87.
Material studied. “Mustajbeg Velipoja Albania”, 2 s. (MNHUB); “Maja Rogamit” / “Albania
leg. Bischoff 1935”, 11 s. (MNHUB); Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 8 s.,
shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH).
Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for the country.
81.15. Agonum thoreyi Dejean, 1828
81.15.1. Agonum thoreyi thoreyi Dejean, 1828
References. Platynus thoreyi: Apfelbeck, 1904: 296 (Velipoja). Agonum thoreyi: Hieke & Wrase,
1988: 87; Bousquet, 2003e: 453.
82. Genus Anchomenus Bonelli, 1810
Subgenus Anchomenus s.str.
82.1. Anchomenus dorsalis (Pontoppidan, 1763)
References. Platynus dorsalis: Apfelbeck, 1904 (Vlorë; Berat; Velipoja); Apfelbeck, 1909: 297
(Vlorë; Berat); Hieke & Wrase, 1988: 83. Platynus dorsale: Schatzmayr, 1943b: 111 (Lushnje);
Hieke et al., 1986: 189 (North Albanian Alps near Theth).
Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alban. Vermos Penther”,
3 s. (NMW); “Skutari Kolcekai” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);
“Bushek bei Elbasan” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bushek juni” / “Albania
leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Lum i Tiranes” / “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Liquen i Butrintes Vivari” /
“leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 12 s.
(MNHUB); “Murrsi-Kara Ali Bey” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB); “Piqeros”
/ “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”,
1 s. (MNHUB); “Am Wege von Murthi nach Konispol” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Cafa e Ceromis Iballe”
/ “Albania Mueller leg. 1939”, 2 s. (MNHUB); “Albanien Tirana Dajti-Vorberge
3.V.39…Mueller”, 1 s. (MNHUB); “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s.
(HNHM); Sauk, 14.III.1961, 1 f. (MNST); Rhesen District, Merkurth, 11.VI.1993, 1 s., P.
Beron & B. Petrov leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B.
Petrov, P. Stoev leg. (NMNHS); Ljukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS).
Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near
Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., 1 f., P.

138. COLEOPTERA: CARABIDAE: HARPALINAE 137
Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,
1 s., leg. M. Komnenov (cSH).
Notes. Bousquet (2003e: 455) has omitted the species for Albania. Here it is confirmed for
the country.
83. Genus Olisthopus Dejean, 1828
83.1. Olisthopus fuscatus Dejean, 1828
References. Olisthopus fuscatus: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88.
Material studied. “Alban. Vermosa Penther”, 2 s. (NMW); “Ibe Erzen” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda District, Lyukove, 5.V.1994, 1 s., P.
Stoev leg. (NMNHS); Saranda District, Lyukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS);
Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).
Notes. Bousquet (2003e: 461) has omitted the species for Albania. Here it is confirmed for
the country.
83.2. Olisthopus glabricollis (Germar, 1817)
References. Olisthopus glabricollis: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88.
Material studied. “Alban. Exp. Kula Ljums 18-28.V.’18”, 4 s. (NMW); “Alban. Exp.’18. Kruma
5.VI.”, 4 s. (NMW); “Burelle” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene”
/ “Albania leg. Bischoff 1936”, 5 s. (MNHUB).
Notes. Bousquet (2003e: 461) has omitted the species for Albania. Here it is confirmed for
the country.
? Olisthopus rotundatus (Paykull, 1790)
? Olisthopus rotundatus rotundatus (Paykull, 1790)
References. Olisthopus rotundatus: Horion, 1941: 310; Hieke & Wrase, 1988: 88. Olisthopus
rotundatus rotundatus: Bousquet, 2003e: 461.
Notes. Horion (1941: 310) and Hieke & Wrase (1988) noted this species for Albania apparently
following the record “Prisren” (Apfelbeck, 1904: 287). Actually this locality is situated in the
Kosovo Region of Serbia. Most probably, Bousquet (2003e: 461) has followed the above
sources. For the time being P. rotundatus is regarded as questionable (possible but yet not
proven) for Albania.
84. Genus Paranchus Lindroth, 1974
Notes. New genus for the fauna of Albania.
84.1. Paranchus albipes (Fabricius, 1796)
Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW).
Notes. New species for the fauna of Albania.

139. 138 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
85. Genus Platynus Bonelli, 1810
Subgenus Batenus Motschulsky, 1865
85.1. Platynus scrobiculatus (Fabricius, 1801)
85.1.1. Platynus scrobiculatus turcicus Apfelbeck, 1904
References. Platynus scrobiculatus turcicus: Schmidt, 1996: 347.
Material studied. “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze Penther’14”,
5 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 26 s. (MNHUB).
Notes. Apfelbeck (1904: 288) described P. scrobiculatus turcicus from Bosnia Herzegovina. In the
same year, Csiki (1904) described P. serbicus Csiki, 1904 from Southeast Serbia. Recently, Bousquet
(2003e: 463) treated both taxa as different species. However, judging by the description of these
taxa and based on research on different populations from the Eastern Balkans (Guéorguiev &
Muilwijk, 2001), I consider the diagnostic characters indicated by Apfelbeck and Csiki reflect
rather geographical variations than being of specific importance. On account of that, P. turcicus
is treated here as conspecific with P. scrobiculatus (Fabricius, 1801): Platynus (Batenus) scrobiculatus
turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus Apfelbeck, 1904. Recently, we
also treated Csiki’s taxon as subspecies of P. scrobiculatus, omitting formal change in rank
(Guéorguiev & Muilwijk, 2001). Here this act is introduced: Platynus (Batenus) scrobiculatus serbicus
Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904. The Albanian populations are
provisionally assigned to the ssp. turcicus. Bousquet (2003e: 463 sub Platynus turcicus) has omitted
the species for Albania. Here it is confirmed for the country with first detailed records.
Subgenus Platynus s.str.
85.2. Platynus assimilis (Paykull, 1790)
References. Platynus assimilis: Apfelbeck, 1904: 296 (Vlorë; Berat; Velipoja); Hieke & Wrase, 1988: 84.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 19 s.
(NMW); “Bushnesh” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” / “Albania leg.
Bischoff 1935”, 5 s. (MNHUB); “Pylla Thumana” / “Albania leg. Bischoff 1935”, 10 s. (MNHUB).
Notes. Bousquet (2003e: 464) has omitted the species for Albania. Here it is confirmed for the country.
Tribe PTEROSTICHINI Bonelli, 1810
86. Genus Abax Bonelli, 1810
Subgenus Abacopercus Ganglbauer, 1891
86.1. Abax carinatus (Duftschmid, 1812)
86.1.1. Abax carinatus carinatus (Duftschmid, 1812) [= latus (Dejean, 1828)]
References. Abax carinatus var. latus: Apfelbeck, 1904: 216 (Velipoja). Abax carinatus: Hieke &
Wrase, 1988: 79. Abax carinatus carinatus: Bousquet, 2003f: 469.
Material studied. “Velipoja Albania”, 1 s. (HNHM); 18 s. from different localities (MNHUB).

140. COLEOPTERA: CARABIDAE: HARPALINAE 139
Subgenus Abax s.str.
86.2. Abax ovalis (Duftschmid, 1812)
References. Abax ovalis: Hieke & Wrase, 1988: 79; Bousquet, 2003f: 470.
Material studied. “Koritnik” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Shkëlzën” /
“Albania leg. Bischoff 1935”, 4 s. (MNHUB).
Notes. First detailed records for Albania.
87. Genus Molops Bonelli, 1810
Notes. The species recognition in this group stay difficult due to the great external resemblance
of the adults that often resulted in impossibility to distinguish the females of closely related
species. Besides, a great similarity and variations in the shape of the aedeagus in close species
occur. The current systematics of the genus must not be treated as definitive (Mlynář, 1977).
Except the “spartanus” group, all the other groups of species of Molops s.l. display North
Aegean origin. The type material used by Müller (1917; 1918a) to describe five Albanian taxa
of Molops s.str. is still preserved in NMW; the same concerns the type material published by
Apfelbeck (1918c).
Subgenus Molops s.str.
Species group “robustus”
87.1. Molops malshentianus Apfelbeck, 1917 [= maderi J. Müller, 1933]
References. Molops malshetianus: Apfelbeck, 1917: 224 (Merdita: Mal i Shenjt Mt. near Oroshi,
loc. typ.); Apfelbeck, 1918b: 88-89 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Apfelbeck,
1930: 167-168 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Mlynář, 1977: 144 (Mal i
Shenjt Mt. near Oroshi). Molops maderi: Müller, 1933: 217 (Kruja Mt., 600 m, loc. typ.); Mařan,
1940c: 192 (Kruja Mt.); Mařan, 1945: 60 (Kruja Mt.); Mlynář, 1977: 39-40 (Kruja Mt.); Turin,
1981: 130; Bousquet, 2003f: 473.
Material studied. “Kruja, Alban., Mader”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff
1932”, 1 f. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 f. (MNHUB); “Mal
i Krujes” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Mal i Krujes” / “Albania leg.
Bischoff 1936”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 f., L.
Penev leg. (NMNHS).
Notes. Apfelbeck published three consecutive descriptions for M. malshentianus (Apfelbeck,
1917; Apfelbeck, 1918b; Apfelbeck, 1930), the second one has been accredited to be the valid.
Actually this is incorrect and the original description must be assigned to the oldest one.
According to Bousquet (2003f: 473) M. malshentianus is synonym of M. maderi, but since the
former has been described earlier, M. malshentianus must be treated as senior synonym: Molops
(Molops) maderi syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917. Notwithstanding,
revision of the types is needed to elaborate this association and to designate lectotypes. The
record from “Maja e Tartarit” should be treated with caution since it is based on a single
female specimen.

141. 140 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
# Molops weiratheri J. Müller, 1930 [= alinae J. Müller, 1932]
References. Molops parnassicola alinae: Müller, 1932: 220 ((“Südalbanien (Epiros)”); Mařan,
1940c: 190, 192.
Notes. Mařan (1940c) specified that the Müller’s record for Albania (Müller, 1932) refers to
Greece. M. weiratheri inhabits the mountains in South Macedonia and North Greece reaching
in the south the massif of Olympus Mt. (Mlynář, 1977: 41-42).
Species group “spartanus”
87.2. Molops spartanus (Schaum, 1862)
? Molops spartanus parnassicola Kraatz, 1875
References. Molops cf. parnassicola: Müller, 1936: 105 (Gjirokastër).
Notes. As Müller (1936) has determinated his material only to species level, the subspecific
treatment of the population near Gjirokastër needs further specification.
87.2.1. Molops spartanus valonensis J. Müller, 1936
References. Molops spec.? parnassicola: Ganglbauer, 1889: 54 (Vlorë). Molops parnassicola:
Apfelbeck, 1904: 217, 220 (Vlorë); Apfelbeck, 1909: 297 (Kanina near Vlorë; Hagion Elias =
Keraun); Müller, 1918a: 7 (Vlorë). Molops parnassicola valonensis: Müller, 1936: 105 (Reza e Kanalit
Mt. near Llogora, loc. typ.; Maja e Tartarit Mt.; Shëndelli Mt.); Mařan, 1940c: 190, 192 (Llogora);
Mařan, 1945: 59 (Vlorë). Molops spartanus valonensis: Mlynář, 1977: 47-48 (Reza e Kanalit Mt.
near Llogora; Llogora; Maja ë Qori; Cafa ë Qori); Bousquet, 2003f: 475.
Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Llogora, Albania mer.”, 4 s.
(MNHUB); “Maj’e Tartarit Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);
“Reza e Kanalit Logara, Albania m.” / “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara”
/ “Albania leg. Bischoff 1932”, 158 s. (MNHUB); “Maja e Qori juni 1932” / “Albania leg.
Bischoff 1932”, 10 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 36 s.
(MNHUB); “Mato-Hasanaj juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen-Ili
juni” / “Albania leg. Bischoff 1932”, 35 s. (MNHUB); “Logara” / “leg. Bischoff Albania
1934”, 48 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 30 s. (MNHUB); Ibë,
24.IV.1962, 1 m. (MNST).
Notes. Before the description of this subspecies, all records from South Albania had been
attributed to M. spartanus parnassicola Kraatz, 1875. Bousquet (2003f: 475) indicated an incorrect
page of the original description of ssp. valonensis.
Species group “simplex”
87.3. Molops simplex Chaudoir, 1868
# Molops simplex simplex Chaudoir, 1868
References. Molops simplex simplex: Bousquet, 2003f: 475.
Notes. Mlynář (1977: 54) noted that the older data for this subspecies from Albania concerns
M. simplex meridionalis, because the nominotypical subspecies inhabits regions removed from
the borders of the country.

142. COLEOPTERA: CARABIDAE: HARPALINAE 141
87.3.1. Molops simplex meridionalis Mlynář, 1977
References. Molops simplex: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m);
Müller, 1918a: 7 (Maranajt Mt.; Rapshë). Molops simplex meridionalis: Mlynář, 1977: 55; Bousquet,
2003f: 474.
Notes: The subspecies occurs only in the marginal northwestern part of the country.
Species group “alpestris”
87.4. Molops alpestris (Dejean, 1828)
87.4.1. Molops alpestris imitator J. Müller, 1917
References. Molops alpestris: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576 m);
Hieke & Wrase, 1988: 81. Molops alpestris imitator: Müller, 1917: 347-348 (Rapshë, loc. typ.); Müller,
1918a: 11 (North Albanian Alps: Rapshë); Mlynář, 1977: 80-81 (Rapshë); Hieke, 1981: 81.
Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 3 syntypes
(NMW); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron & B. Petrov leg.
(NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 8 s., P. Beron & B. Petrov
leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron & B.
Petrov leg. (NMNHS).
Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here, it is confirmed for
the country.
Species group “elatus”
87.5. Molops parreyssi Kraatz, 1875
87.5.1. Molops parreyssi lonae J. Müller, 1921
References. Molops parreyssi: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m);
Klaptocz, 1910: 399 (Theth); Müller, 1918a: 7 (Maranajt Mt.). Molops parreyssi lonae: Müller,
1921b: 28-30 (Çukall Mt., loc. typ.); Mlynář, 1977: 96 (Çukall Mt.: cave “Kunor Vukai”);
Bousquet, 2003f: 473.
Material studied. “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Cucale
Albania sept.” / “leg. Bischoff 1931”, 4 s. (NMW); nearly 100 s. from locality “Cucale”
(MNHUB); 4 s. from locality “Maranay” (MNHUB).
Notes. The species occurs only in the marginal northwestern parts of Albania.
Species insertae sedis
87.6. Molops merditanus Apfelbeck, 1906
87.6.1. Molops merditanus merditanus Apfelbeck, 1906
References. Molops merditanus: Apfelbeck, 1906: 1664 (Merdita: Mal i Shenjt Mt. near Oroshi,
loc. typ.; Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1907a: 499, 503 (Mal i Shenjt Mt.:
top region, 1500 m); Müller, 1918a: 10 (Merdita: Mal i Shenjt Mt. near Oroshi; Merdita: Zepë
Mt. near Fandi); Müller, 1930 (Merdita). Molops merditanus merditanus: Mlynář, 1977: 106-107
(Mal i Shenjt Mt. near Oroshi; Zepë Mt.); Bousquet, 2003f: 473.

143. 142 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Merdita M. Scheit”, 1 paralectotype (DEI); “Merdita M. Scheit” / “Apfelb.
Alban. spt. IV.V.’05” / “Molops merditanus typ. † Apfelb.”, 1 f. (NMW); “Merdita Zebia”, 2
paralectotypes (DEI); “Merdita Zebia” / “Molops merditanus typ. ‡ Apfelb.”, 1 f. (NMW).
Notes. Mlynář (1977: 106) noted that the lectotype of M. merditanus is in the collection of
HNHM, but I could not find it there.
87.6.2. Molops merditanus intermedius J. Müller, 1917
References. Molops obtusangulus intermedius: Müller, 1917: 348-349 (Montenegro: Podgorica,
loc. typ.; Albania / Montenegro border: Ricavac, loc. typ; Albania: Vermosh, loc. typ). Molops
merditanus intermedius: Mlynář, 1977: 107-108 (Ricavac; Vermosh); Drovenik & Peks, 1994: 71
(Ricavac); Drovenik & Peks, 1999: 83 (Ricavac); Bousquet, 2003f: 473.
Material studied. “Ricavac juni” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW);
“Vermosa 10-21.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW).
Notes. Each of the three localities mentioned in the original description (Müller, 1917) could
be a potential “locus typicus”, however a lectotype designation still has to be done.
Species group “curtulus”
Notes. This taxonomically difficult species complex is treated here differently than previously
(Mlynář, 1977):
Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri
reiseri Apfelbeck, 1904
Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri
corallipes J. Müller, 1936
Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri
pseudoalpestris Mlynář, 1977
Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri
victoris J. Müller, 1918
Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J.
Müller, 1917
87.7. Molops albanicus Apfelbeck, 1904
References. Molops albanicus: Apfelbeck, 1906: 1665 (Mal i Shenjt Mt.; Zepë Mt.); Apfelbeck,
1907a: 499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 8 (Merdita: Mal i Shenjt
Mt.; Merdita: Zepë Mt.); Müller, 1930: 124 (Merdita: Zepë Mt.); Mlynář, 1977: 118, 120 (Mal
i Shenjt Mt.; Zepë Mt.; Oroshi); Bousquet, 2003f: 472.
Material studied. “Merdita Zebia”, 9 s. (DEI; HNHM; NMW); “Merdita M. Scheit” / “Coll.
Apfelbeck”, 1 m. (HNHM); “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 21 s. (DEI;
NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 3 s. (NMW); “Alban. Exped. Pashtrik’18
4-17.7”, 1 s. (NMW); 9 s. from locality “Merdita” (MNHUB); 33 s. from locality “Llogora”
(MNHUB); 11 s, from locality “Mal i Shenjit” (MNHUB); 3 s. from locality “Iballe” (MNHUB);
1 s. from locality “Mal i Gjer” (MNHUB); 3 s. from locality “Shen-Ili” (MNHUB).
Additional material included (studied A. Vigna Taglianti). Valbona, Hellit Kuq Mt., south
slope, 930 m, 6.VII.1993, 1 m., L. Latella leg. (cAVT).

144. COLEOPTERA: CARABIDAE: HARPALINAE 143
Notes. Apfelbeck (1904: 222) described this species basing on a single male from the type
locality “Prizren”. This place is situated in the Kosovo Region of Serbia. The binomen Molops
transfuga Apfelbeck (Müller, 1930: 124; Mlynář, 1977: 120) is nomen nudum.
87.8. Molops apfelbecki Ganglbauer, 1891
Notes. In the past, M. apfelbecki and M. reiseri have been treated as separate species. Probable
cause for this state is that Apfelbeck (1904: 223), while describing M. reiseri, missed to compare
it with M. apfelbecki. Later, Mlynář (1977) similarly did not compare the aedeagus of both species,
but noticed the resemblance in their female genital structures. The present author has studied
the aedeagus of M. reiseri holdhausi (syntype, NMW), M. reiseri (lectotype, HNHM), M. reiseri
victoris (syntype, NMW; specimens from Koprivnik Mt. and Shkëlzen Mt., HNHM) and M. apfelbecki
(lectotype, HNHM) and found that M. reiseri holdhausi is different in the shape of the median
lobe from the other three taxa. In fact, the last taxa have identical aedeagus. Therefore, M. reiseri
reiseri and M. apfelbecki apfelbecki are considered as geographical races of single species, and M.
holdhausi – as valid species which occurs together with M. apfelbecki s.l.
87.8.1. Molops apfelbecki victoris J. Müller, 1918
References. Molops victoris: Müller, 1918a: 1-3, 6 (Albania / Montenegro border: North Albanian
Alps, 1300-2000 m, loc. typ.). Molops reiseri victoris: Mlynář, 1977: 114-115 (North Albanian
Alps; Shkëlzen Mt.).
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” /
“victoris n. det. J. Müller”, 2 m. syntypes of Molops victoris (NMW); “Albania Skölsen” / “Akad.
Balk. Exp. Csiki 1917”, 1 m. with extracted penis (HNHM); “Maja Peis” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 5 s. (MNHUB); “Shkëlzën”
/ “Albania leg. Bischoff 1936”, 22 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Bischoff,
Mueller 1939” / “Type” / “Apfelbecki sbsp. Bischoffi m. det. Müller”, 1 s. (MNHUB); “M. Raba
Massiv” / “Albania leg. Bischoff, Mueller 1939” / “Type”, 5 s. (MNHUB); “Maja e Dragomirit”
/ “Albania leg. Bischoff, Mueller 1939” / “M. Apfelbecki sbsp. Bischoffi m. det. G. Müller 1940”,
1 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Bischoff, Mueller 1939”, 5 s. (MNHUB).
Additional material studied. Bosnia Herzegovina / Montenegro. “Apfelbeck Volujak” /
“Coll. Apfelbeck”, 1 m. lectotype and 3 paralectotypes of Molops apfelbecki (HNHM).
Montenegro. “Vunšaj, Cafa glava, Alban. Penther” / “reiseri Apf.”, 1 m. of Molops apfelbecki
victoris (NMW). Serbia (Kosovo Region). “Montes Koprivnik” / “Akad. Balk. Exp. Csiki 1917”,
1 m. of Molops apfelbecki victoris (HNHM). Serbia. “Reiser 1898 Kopaonik” / “Coll. Apfelbeck”,
1 m. lectotypus and 2 paralectotypes of Molops reiseri (HNHM); “Reiser 1899 Kopaonik” /
“reiseri Apf. B& typus” / “type”, 1 m. of Molops apfelbecki reiseri (NMW); “Reiser 1899 Kopaonik”
/ “reiseri Apf. Serb. turk. Grence” / “type”, 1 m., 1 f. of Molops apfelbecki reiseri (NMW); “Reisser
1899 Kopaonik”, 1 m., 1 f. of Molops apfelbecki reiseri (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. Müller (1930: 126) revised the material of Apfelbeck (1918c: 166 sub M. reiseri from
Vunšaj, Cafa glava) and determinated it as M. victoris. However, Bousquet (2003f: 474) has omitted
M. reiseri victoris (currently M. apfelbecki victoris, see above) for Albania in spite of the fact that
Mlynář (1977: 115) gave a detailed record. Here the subspecies is confirmed for the country. The
trinomen M. apfelbecki bischoffi G. Müller 1940, found on labels of two specimens, is nomen nudum.

145. 144 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
87.9. Molops curtulus Ganglbauer, 1891
87.9.1. Molops curtulus cakorensis Mlynář, 1977
References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps). Molops curtulus:
Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Müller, 1918a: 10 (Maranajt
Mt.; North Albanian Alps: Rapshë, Grebeni, Vermosh, Ricavac); Csiki, 1940: 217 (Shkëlzen
Mt.). Molops curtulus cakorensis: Mlynář, 1977: 111.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s.
(NMW); “Vermosa Penther”, 1 s. (NMW); “13.15.VI.14 Vermosa ca 1000 m”, 1 s. (NMW);
“Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Rapsa
10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 8 s. (NMW); “Grebeni 16.VI.” / “Alban.
montenegr. Grenze, Penther’14”, 5 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban.
montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Rogamit” / “Albania leg. Bischoff
1935”, 3 f. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 9 s. (MNHUB); “Boge
Aufstieg z. Rab” / “Albania leg. Bischoff, Mueller 1939”, 1 s. (MNHUB); North Albanian
Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS).
Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here it is confirmed for
the country.
87.9.2. Molops curtulus winklerianus J. Müller, 1936
References. Molops winklerianus: Müller, 1936: 103 (Tomor Mt.: Kulmak, loc. typ.; Shëndelli
Mt. near Tepelenë; Llogora); Molops curtulus winklerianus: Mlynář, 1977: 111 (Tomor Mt.: Kulmak;
Llogora); Bousquet, 2003f: 472.
Material studied. “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW).
Notes. Mlynář (1977: 111) stated in a footnote that the populations of M. curtulus from Central
Albania may belong to a separate race, so further investigations of material from this region
of the country will reveal more details for elucidation that statement.
87.10. Molops holdhausi J. Müller, 1917
References. Molops holdhausi: Müller, 1917: 352-353 (Albania / Montenegro border: North
Albanian Alps, 1300-2000 m, loc. typ.); Müller, 1918a: 6 (North Albanian Alps); Müller, 1930:
126 (North Albanian Alps, Grebeni near Vermosh). Molops reiseri: Apfelbeck, 1918c: 166 (North
Albanian Alps, Grebeni near Vermosh). Molops reiseri holdhausi: Mlynář, 1977: 112, 114 (North
Albanian Alps).
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” /
“holdhausi n. det. J. Müller”, 1 m. syntype of Molops holdhausi (NMW); “Grebeni, Vermoša
Alban., Penther 16 Juni, am Schnee” / “reiseri Apf.”, 2 m. (NMW).
Additional material studied. Montenegro. “Durmitor”, 4 s. (HNHM); “Montenegro
Durmitor, Penther’04”, 1 m., 1 f. (NMW); “Jugoslavia mer. Prokletija Gusinje Matìjka VI.1937”,
1 m., 1 f. (MNHUB); “Jugoslavia mer. Prokletja Gusinje Kodym 20.VI.1937”, 2 s. (HNHM).
Notes. Müller described M. holdhausi and M. victoris as separate species (Müller 1917; Müller
1918a). Mlynář (1977: 112-113) neglected the fact that M. reiseri holdhausi and M. reiseri victoris
occur sympatrically and expressed doubt about the status of both taxa due to the lack of
enough specimens at hand (Mlynář (1977: 115). The present author compared the penis
structures of several males, previously determined as ssp. holdhausi or ssp. victoris, and ascertained

146. COLEOPTERA: CARABIDAE: HARPALINAE 145
the first is good species - Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops)
reiseri holdhausi J. Müller, 1917. Bousquet (2003f: 474) has omitted M. holdhausi for Albania
though Müller (1930) gave a detailed record. Here the species is confirmed for the country.
87.11. Molops plurisetosus J. Müller, 1918 [= malissorum Apfelbeck, 1918]
References. Molops apfelbecki plurisetosus: Müller, 1918a: 3-4, 11 (Albania / Montenegro border:
North Albanian Alps, 1200-2000 m, loc. typ.; Albania / Montenegro border: Ricavac, loc. typ.;
Albania: Vermosh, loc. typ.). Molops malissorum: Apfelbeck, 1918c: 160, 167-168 (Montenegro:
North Albanian Alps, Vusanje, loc. typ.; Albania: pastures over Theth). Molops plurisetosus: Mlynář,
1977: 120-121 (North Albanian Alps; Ricavac; Vermosh).
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4
syntypes of Molops plurisetosus (NMW); “Ricavac Juni” / “Alban. montenegr. Grenze, Penther’14”,
1 syntype of Molops plurisetosus (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze,
Penther’14”, 2 syntypes of Molops plurisetosus (NMW); North Albanian Alps, Theth, 800-1200 m,
28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes,
2400-2550 m, 29.V.1993, 3 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps,
Bogë, 1800-1900 m, 20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian
Alps, Bogë, cave “No 25”, 23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Shkodër
District, Kozuja, 1800 m, 27.VIII.1996, 1 m., 1 f., T. Ivanova leg. (NMNHS).
Additional material studied. Montenegro. “Vunšaj, Prokletija-Geb., Albania Dr. Penther” /
“Molops malissorum n. sp. Apf.”, 2 syntypes of Molops malissorum (NMW).
Notes. Bousquet (2003f: 474) has omitted the species for Albania. Here it is confirmed for the
country. Currently, the exact publishing dates of the descriptions of M. plurisetosus and M. malissorum
were not found, so the problem with the priority of one of the two names is pending. Each of
the different localities mentioned in the original descriptions of M. plurisetosus (Müller, 1918a)
could be a potential “locus typicus”, however lectotype designation still has to be done.
Species group “rufipes”
87.12. Molops osmanilis Apfelbeck, 1904
[Molops osmanilis osmanilis Apfelbeck, 1904]
Notes. The record from “Prisren” (Apfelbeck, 1904: 229) refers to the Kosovo Region of Serbia,
so perhaps Bousquet (2003f: 473) has followed this source when listing M. osmanilis osmanilis for
Albania. The nominotypical subspecies is treated here as likely to be found in the country.
87.12.1. Molops osmanilis tomoricensis J. Müller, 1923
References. Molops tomoricensis: Müller, 1923: 109-110 (Tomor Mt., alpine zone, loc. typ.). Molops
osmanilis tomoricensis: Mlynář, 1977: 123 (Tomor Mt.: Tomor, Abbas Ali); Bousquet, 2003f: 473.
Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 5 s. (NMW;
MNHUB); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);
“Tomor: Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor:
Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 3 (NMW); “Tomor: Tomor Alban.
mer.” / “Winkler, Lona, Bischoff, V.1931”, 7 s. (NMW); more than 90 s. from Tomor Mt.
(MNHUB); “Mal i Dajtit Tirane” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara

147. 146 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit Tirane” / “leg. Bischoff
Albania 1934”, 1 s. (MNHUB); “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff”,
13 s. (MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s.
(MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 2 s. (MNHUB); “AL (Tirana) 8 km
N psene E Tirana 1450 m. 30.IV.1993 leg. H. & S. Peks”, 2 s. (MNHUB).
Notes. All the specimens examined from Albania are put to this race more or less provisionally.
However, for instance specimens from Southwest Albania (localities “Logara” and “Maja e
Dhembelit”) have a shape of the pronotum like spp. osmanilis, whereas these from Central
Albania (localities “Mal i Dajtit” and “Mal i Krujes”) – like ssp. tomoricensis.
87.13. Molops rufipes Chaudoir, 1843
87.13.1. Molops rufipes rufipes Chaudoir, 1843
References. Molops rufipes: Rambousek, 1912: 93; Mlynář, 1977: 124; Hieke, 1981: 81; Hieke &
Wrase, 1988: 82. Molops rufipes rufipes: Mlynář, 1977: 124 (Mal i Dajtit Mt.).
Material studied. “Mal i Daitit b. Tirana, Alb. C.” / “leg. Bischoff 1931 coll. A. Winkler”, 2
s. (NMW); “Tomor: Tomor Alban. mer.”, 1 m. (MNHUB); “Tomor-Buranj juli 1932” / “Albania
leg. Bischoff 1932”, 1 m. (MNHUB); more than 50 s. from locality “Iballe” (MNHUB); more
than 50 s. from locality “Mal i Dajtit” (MNHUB); Tiranë, 26.VII.1965, 1 f. (MNST); Mal i
Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 4 m., 3 f., L. Penev leg. (NMNHS).
Notes. Bousquet (2003f: 474) has omitted the subspecies for Albania. Here it is confirmed
for the country. The material of Rambousek (1912) was not revised and therefore it is placed
in the nominotypical race provisionally.
# Molops rufipes klisuranus Apfelbeck, 1902
References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps).
Notes. Two wrong records (Müller, 1930: 125-126, revised as M. curtulus s.l.) for this taxon
have been recorded from the Western Balkan (South Serbia, Sandzak: Raška near Novi Pazar,
Apfelbeck, 1904: 222; North Albanian Alps, Apfelbeck, 1918c: 166). Therefore the taxon is
removed from the list of the Albanian Carabidae.
87.13.2. Molops rufipes munelensis Apfelbeck, 1906
References. Molops sturanyi munelensis: Apfelbeck, 1906: 1665-1666 (Munella Mt., loc. typ.);
Müller, 1918a: 9 (Merdita: Munella Mt.). Molops rufipes munelensis: Mlynář, 1977: 128, 130 (Munella
Mt.; Munella Mt.: Oroshi; Berishe); Bousquet, 2003f: 474.
Materialstudied.“Orosi”,5s.(HNHM);“Oroši(Ptr.)Alban.spt.”/“Munela”/“Coll. Apfelbeck”,
1 lectotype (HNHM); “Merdita Munela”, 3 paralectotypes (HNHM); “Merdita Munela”, 9 s. (DEI;
MNHUB); “Merdita Munela Apfb.”, 4 s. (MNHUB); “Merdita” / “Apfb.”, 2 s. (DEI); “Merdita
Munela” / “Wgth. 1906 Merdita”, 21 s. (NMW); “Merdita Zebia”, 9 s. (NMW); “Alban. Exp.’18
Pashtrik, 29.V.-4.VI.”, 9 s. (NMW); “Alban. Exp.’18 Pashtrik, 4-17.7”, 5 s. (NMW); “M. Mulecifit”
/ “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Berishe” / “Albania leg. Mueller 1939”, 6 s.
(MNHUB); “AL (Shkodar). Paß SE Puke 850 m 2.V.1993” / “leg. H. & S. Peks”, 1 m. (MNHUB).
[Molops rufipes pseudoosmanilis Mlynář, 1977]
Notes. Since this subspecies was hitherto recorded only from the Macedonian section of the
Korabi Mt., it is expected to live in Albania, too.

148. COLEOPTERA: CARABIDAE: HARPALINAE 147
87.13.3. Molops rufipes steindachneri Apfelbeck, 1908
References. Molops rufipes steindachneri: Mlynář, 1977: 128 (Korab Mt.).
Notes. Bousquet (2003f: 474) has omitted this subspecies for Albania. Actually its occurrence in
the country needs confirmation. Apfelbeck published three consecutive descriptions of this taxon
(Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid.
87.13.4. Molops rufipes sturanyi Apfelbeck, 1906
References. Molops sturanyi: Apfelbeck, 1906: 1665 (Merdita: Mal i Shenjt Mt., loc. typ.; Merdita:
Munella, loc. typ.; Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 497, 499, 503 (Maranajt Mt.:
top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 9 (Merdita:
Zepë Mt.; Merdita: Mal i Shenjt Mt.; Maranajt Mt.); Müller, 1930: 123 (Gjalicë Mt.). Molops
steindachneri: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Molops rufipes sturanyi: Mlynář, 1977: 128
(Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 474.
Material studied. “Albania” / “Merdita”, 2 s. (DEI); “Merdita M. Schèit” / “Apfelb. Alban.
spt. IV.V.’05”, 1 lectotype (HNHM); “Merdita Zebia”, 4 paralectotypes (HNHM); “Merdita
Zebia”, 6 s. (DEI; MNHUB); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s.
(HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM);
“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 11 s. (HNHM); “Alban. Exp. 1918
Gjalica Ljums 17-26.VI.”, 49 s. (NMW); 15 s. from locality “Bjeshka Matroxh” (MNHUB); 20
s. from localaity “Derzë” (MNHUB); more than 300 s. from locality “Gjallica” (MNHUB);
more than 500 s. from locality “Korab” (MNHUB); 2 s. from locality “Taja” (MNHUB).
Notes. The record from the Maranajt Mt. (Müller, 1918a: 9) has to be treated cautiously
because it is the only record of this species north of the Drin River.
Species group “piceus”
87.14. Molops piceus (Panzer, 1793)
87.14.1. Molops piceus balcanicus Mlynář, 1977
References. Molops austriacus: Müller, 1918a: 9 (Rapshë; North Albanian Alps); Horion, 1941: 302.
Molops piceus balcanicus: Mlynář, 1977: 136; Hieke, 1981: 81. Molops piceus: Hieke & Wrase, 1988: 82.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s.
(NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s.
(NMW); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania
leg. Bischoff 1935”, 4 s. (MNHUB).
Notes. Bousquet (2003f: 473) has omitted the species for Albania. Here it is confirmed for
the country.
Subgenus Stenochoromus L. Miller, 1866
87.15. Molops montenegrinus (L. Miller, 1866)
87.15.1. Molops montenegrinus cavernicola J. Müller, 1916
References. Molops montenegrinus cavernicola: Müller, 1916: 81 (Çukall Mt.: cave “Sckamit”, loc.
typ.); Müller, 1918a: 11-12 (Çukall Mt.: cave “Sckamit”; North Albanian Alps: Ricavac, Vermosh);
Bousquet, 2003f: 475.

149. 148 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, more
than 20 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Starka
Alb. sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 6 s. (NMW); “Maja e Jezerce Albania sept.”
/ “leg. Bischoff 1931 coll. A. Winkler”, 2 s. (NMNHS); “Maja e Jezercoë” / “Albania leg.
Bischoff 1933”, 9 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 27 s.
(MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cucale” / “Albania
leg. Bischoff 1936”, 3 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 95 s. (MNHUB);
“Maranaj” / “Albania leg. Bischoff 1937”, 3 s. (MNHUB); “Boge Aufstieg Z. Rab” / “Albania
leg. Mueller 1939”, 2 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 7 s.
(MNHUB); “Maja e Dragomirit” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); North Albanian
Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 2 m., P. Beron & B. Petrov leg. (NMNHS); North
Albanian Alps, Maj’e Redohimes, 1800-2200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg.
(NMNHS); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron &
B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron
& B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Mali-i-Thatë Mt., cave “Shpella e
Haxise, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m,
11.VII.1993, 1 m., 2 f., L. Latella leg. (cAVT).
Notes. All the data from Albania are placed provisionally to the ssp. cavernicola, since no
intraspecific revision of M. montenegrinus s.l. is made. In addition to the material studied above,
in NMW I have seen more specimens, which remained unregistered.
88. Genus Myas Sturm, 1826
Subgenus Myas s.str.
88.1. Myas chalybaeus (Palliardi, 1825) [= rugosicollis Brullé, 1832]
References. Myas chalybaeus: Tschitschérine, 1899: 70; Anonymous, 1907: 284; Hieke & Wrase,
1988: 69; Bousquet, 2003f: 475. Myas rugosicollis: Tschitschérine, 1899: 70.
Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW); “Mal i Dajtit Tiranë” / “Albania
leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB);
“Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania
leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit Tiranë” / “Albania 1936 leg. Ramadan coll.
Bischoff”, 7 s. (MNHUB); “Valbona Çerem” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Berishe” / “Albania Mueller leg. 1939”, 3 s. (MNHUB); “Boge Ober D. Quelle” / “Albania
Mueller leg. 1939”, 3 s. (MNHUB); “Çafa e Ceromis Iballe” / “Albania Mueller leg. 1939”, 1 s.
(MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov
leg. (NMNHS); 5 km Leskoec, 2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Stebleva, cave “Shpella
e Hadzhi i Malit”, 5-8.V.2006, 1 s., A. Zhalov, D. Raychev & J. Tsvetanov leg. (NMNHS).
Notes. First detailed records for Albania.

150. COLEOPTERA: CARABIDAE: HARPALINAE 149
89. Genus Pedius Motschulsky, 1850
89.1. Pedius inquinatus (Sturm, 1824)
References. Pterostichus inquinatus: Apfelbeck, 1904: 257 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);
Schatzmayr, 1930: 198; Horion, 1941: 279; Schatzmayr, 1942: 65; Schatzmayr, 1943b: 110
(Mal i Krujes); Hieke & Wrase, 1988: 71. Pedius inquinatus: Bousquet, 2003f: 478.
Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Lum i Tiranes” / “leg.
Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s.
(MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 22-
31.X.1997, 3 s., R. Ward leg. (CMNH).
89.2. Pedius longicollis (Duftschmid, 1812)
References. Pedius longicollis: Bousquet, 2003f: 478.
90. Genus Poecilus Bonelli, 1810
Subgenus Ancholeus Dejean, 1828
90.1. Poecilus puncticollis (Dejean, 1828)
References. Pterostichus puncticollis: Schatzmayr, 1943b: 110 (Kruja; Skrofotina; Stan-Karbunarë).
Poecilus puncticollis: Hieke & Wrase, 1988: 71; Bousquet, 2003f: 480.
Material studied. “Liquen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);
“Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).
Subgenus Poecilus s.str.
90.2. Poecilus cupreus (Linnaeus, 1758)
90.2.1. Poecilus cupreus cupreus (Linnaeus, 1758)
References. Pterostichus cupreus: Apfelbeck, 1904: 253 (Velipoja); Apfelbeck, 1907a: 495 (Buenë
River); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1930: 184 (Tomor Mt.); Schatzmayr, 1943b:
109 (Kruja; Stan-Karbunarë; Lushnje). Poecilus cupreus: Hieke & Wrase, 1988: 70. Poecilus cupreus
cupreus: Bousquet, 2003f: 482.
Material studied. “Oboti Bojana”, 1 s. (NMW); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”,
1 s. (NMW); “Alban. Exp.’18 Pashtrik 4-14.7.”, 7 s. (NMW); “Albanien Ura Gjolés Umg. Tirana
Marz 1942 Muller Ausbeute”, 1 s. (MNHUB); “Albania Ostrovice” / “7.VII.1959 leg. Xh.
Murraj”, 2 s. (HNHM); Sauk, 14.V.1961, 1 m. (MNST); Sauk, 3.IV.1963, 1 m. (MNST); Saranda
District, Butrinti, 16.IV.1994, 3 s., S. Beshkov leg. (NMNHS); Tiranë Park, 8.II.1998, 2 m.,
shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A.
Paparistos leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT).
Notes. In addition to the material studied, in MNHUB I have seen other more 24 s. from
different localities, which remained unregistered.

151. 150 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
90.3. Poecilus cursorius (Dejean, 1828)
90.3.1. Poecilus cursorius cursorius (Dejean, 1828)
References. Pterostichus cursorius: Schatzmayr, 1930: 187; Schatzmayr, 1943b: 110 (Lushnje).
Poecilus cursorius: Hieke & Wrase, 1988: 71. Poecilus cursorius cursorius: Bousquet, 2003f: 482.
Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Rushkull” / “Albania
leg.Bischoff 1936”,1s.(MNHUB);“SukthibeiGurse”/“Albanialeg.Bischoff 1936”,1s.(MNHUB).
90.4. Poecilus koyi Germar, 1824
90.4.1. Poecilus koyi koyi Germar, 1824
References. Pterostichus koyi dinaricus: Schatzmayr, 1930: 176 (Tomor Mt.); Schatzmayr, 1942:
55 (Tomor Mt.). Pterostichus koyi: Schatzmayr, 1943b: 109 (Tërpan). Poecilus koyi: Csiki, 1940:
216 (Korab Mt.); Hieke et al., 1986: 188 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988:
69. Poecilus koyi koyi: Bousquet, 2003f: 482.
Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 18 s. (NMW);
“Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Albania Mts. Korab” /
“Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Korab 1918 24.31.VII.”, 1 s.
(NMW); “Albania Rapsa Dr. Grabowski”, 1 s. (HNHM); “Albania Zalle-Dardha” / “23.V.1958
leg. Xh. Murraj”, 3 s. (HNHM); Bizë, 9-16.VII.1961, 2 f. (MNST); Mal i Dajtit Mt., 10-11.V.1965,
5 m., 1 f.; Mal i Dajtit Mt., 24.V.1966, 2 f. (MNST); Tiranë, 30.V. 1965, 2 m. (MNST); 2 km N
Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS).
90.5. Poecilus lepidus (Leske, 1785)
90.5.1. Poecilus lepidus lepidus (Leske, 1785)
References. Pterostichus lepidus: Csiki, 1940: 216 (Korab Mt.); Schatzmayr, 1942: 56. Poecilus
lepidus: Hieke & Wrase, 1988: 70. Poecilus lepidus lepidus: Bousquet, 2003f: 483.
Material studied. “Alban. Vermosa Penther”, 4 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW);
“Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania
Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Shkëlzën” / “Albania leg. Bischoff
1936”, 2 s. (MNHUB).
90.6. Poecilus rebeli (Apfelbeck, 1904)
References. Pterostichus rebeli: Apfelbeck, 1904: 254 (Velipoja; Vlorë); Apfelbeck, 1907a: 495
(Buenë River); Scatzmayr, 1930: 184; Schatzmayr, 1942: 59; Bousquet, 2003f: 483.
Material studied. “Velipoja Albania” / “Coll. Apfelbeck”, 1 syntype (HNHM); “Mustajbeg
Velipoja Albania”, 2 syntypes (NMW); “Oboti Albania”, 1 s. (NMW); “Scutari Bojana” /
“Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW).
90.7. Poecilus striatopunctatus (Duftschmid, 1812)
References. Poecilus striatopunctatus: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988:
71; Bousquet, 2003f: 484.
Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

152. COLEOPTERA: CARABIDAE: HARPALINAE 151
90.8. Poecilus versicolor (Sturm, 1824) [= coerulescens (Linnaeus, 1758)]
Material studied. “Alban. Ricavac Penther”, 2 s. (NMW); “Alban. Vermosa Penther”, 4 s.
(NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki
1918”, 1 s. (HNHM); “Albania Mali-Thate” / “17.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM);
“Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 1 s. (HNHM); Golem, 10.VII. 1965, 1 s.
(MNST); Tiranë, 26.VIII.1965, 1 m. (MNST).
Notes. The mention of Hieke & Wrase (1988: 70) for Albania bases apparently on the record
“Prisren” (Apfelbeck, 1904: 255 sub Pterostichus coerulescens). However, currently this locality is
situated in the Kosovo Region of Serbia. Bousquet (2003f: 484) has most likely followed one
of these sources too. Since there is no previous reliable record from Albania, P. versicolor is
treated here as new species for the fauna of Albania.
91. Genus Pterostichus Bonelli, 1810
Subgenus Argutor Dejean, 1821
91.1. Pterostichus cursor (Dejean, 1828)
References. Pterostichus vernalis cursor: Apfelbeck, 1904: 257 (Velipoja). Pterostichus cursor: Hieke
& Wrase, 1988: 73; Bousquet, 2003f: 486.
Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari l. See-Ufer”, 1 s. (NMW); “Skutari
Mesi”, 1 s. (NMW); “Skutari Vraka”, 1 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW); “Liquen
i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
Subgenus Bothriopterus Chaudoir, 1835
91.2. Pterostichus oblongopunctatus (Fabricius, 1787)
91.2.1. Pterostichus oblongopunctatus oblongopunctatus (Fabricius, 1787)
References. Pterostichus oblongopunctatus: Schatzmayr, 1943a: 85; Hieke & Wrase, 1988: 74.
Pterostichus oblongopunctatus oblongopunctatus: Bousquet, 2003f: 487.
Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Maja Rogamit” / “Albania leg. Bischoff
1935”, 2 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).
Notes. First detailed records for Albania.
Subgenus Cheporus Latreille, 1829
91.3. Pterostichus burmeisteri Heer, 1838
91.3.1. Pterostichus burmeisteri burmeisteri Heer, 1838 [= metallicus (Fabricius, 1792)]
References. Pterostichus metallicus: Klaptocz, 1910: 399 (Theth).
Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW);
“Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja
e Jezercë Çafa Ostars” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Shkëlzën” / “Albania

153. 152 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
leg. Bischoff 1935”, 18 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);
“Maja Rogamit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Boge Aufstieg z. Rab” /
“Albania Mueller leg. 1939”, 1 s. (MNHUB); “Boge Çafa Shtegot” / “Albania Mueller leg. 1939”,
1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg. 1939”, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 15.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. Bousquet (2003f: 488) has omitted the species for Albania. Here it is confirmed for the country.
Subgenus Feronidius Jeannel, 1942
91.4. Pterostichus melas (Creutzer, 1799)
91.4.1. Pterostichus melas depressus (Dejean, 1828)
References. Pterostichus melas: Apfelbeck, 1904: 265 (Vlorë; Velipoja); Apfelbeck, 1909: 297
(Vlorë; Berat); Schatzmayr, 1943b: 109 (Kruja; Dukati; Tërpan); Hieke et al., 1986: 189 (Tiranë;
Borshi near Vlorë; Poliçan near Tomor Mt.). Pterostichus melas depressus: Hieke & Wrase, 1988:
76; Bousquet, 2003f: 495.
Material studied. “Mustajbeg Velipoja Albania”, 4 s. (HNHM); “Sudalbanien Kopstein”, 1 s.
(NMW); “Skutari Penther”, 4 s. (NMW); “Hani Grabom mai” / “Alban. montenegr. Grenze,
Penther’14”, 1s.(NMW); “Rapsa10-18.V.”/“Alban.montenegr.Grenze,Penther’14”,1s.(NMW);
“Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 1 s.
(HNHM); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Sauk” /
“16.VI.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Terbac” / “6.VI.1960 leg. Xh. Murraj”, 1 s.
(HNHM); Sauk, 9.III.1961, 1 m. (MNST); Tepelenë, 1-3.VI.1961, 1 m. (MNST); Rodenj, 26-
28.VII.1962, 2 m. (MNST); Kurbneshi, VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Tiranë,
Selit e Vogel, 17.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Saranda District, Ljukove, 5.V.1994, 6 s.,
P. Stoev leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg.
(NMNHS); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997, 5 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 f., A. Paparistos
leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT).
Notes. Besides the material studied above, in MNHUB I have seen other more 63 s. from
Albania, which remained unregistered.
Subgenus Melanius Bonelli, 1810
91.5. Pterostichus atterimus (Herbst, 1784)
91.5.1. Pterostichus atterimus atterimus (Herbst, 1784)
References. Pterostichus atterimus: Apfelbeck, 1904: 258 (Velipoja); Schatzmayr, 1943a: 87.
Pterostichus atterimus atterimus: Bousquet, 2003f: 499.
Material studied. “Gjormi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
91.6. Pterostichus elongatus (Duftschmid, 1812)
References. Pterostichus elongatus: Schatzmayr, 1930: 210 (Shkodër); Hieke & Wrase, 1988: 76;
Bousquet, 2003f: 499.

154. COLEOPTERA: CARABIDAE: HARPALINAE 153
Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Mamurash” / “Albania leg.
Bischoff 1936”, 1 s. (MNHUB).
Subgenus Morphnosoma Lutshnik, 1915
91.7. Pterostichus melanarius (Illiger, 1798)
91.7.1. Pterostichus melanarius melanarius (Illiger, 1798)
References. Pterostichus melanarius: Hieke & Wrase, 1988: 75. Pterostichus melanarius melanarius:
Bousquet, 2003f: 499.
Material studied. “Alban. Exp. 1918 Gjallica Ljums 17-26.VI.”, 1 s. (NMW); “Lesh” / “Albania
leg. Bischoff 1937”, 1 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 8-16.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT).
Notes. First detailed records for Albania.
# subgenus Oreoplatysma Jakobson, 1907
# Pterostichus lodosi Heinz, 1977
References. Pterostichus lodosi: Bousquet, 2003f: 505.
Notes. Most probably, Bousquet (2003f: 505) has listed this Caucasian endemic species for
the country by mistake. Therefore, it is removed from the list of the Albanian ground beetles.
Subgenus Parahaptoderus Jeanne, 1969
Notes. New subgenus for the fauna of Albania.
91.8. Pterostichus brevis (Duftschmid, 1812)
91.8.1. Pterostichus brevis brevis (Duftschmid, 1812)
Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” /
“Alban. montenegr. Grenze Penther’14”, 7 s. (NMW).
Notes. New species for the fauna of Albania.
Subgenus Phonias Gosis, 1886
91.9. Pterostichus apfelbecki Csiki, 1908 [= convexiusculus Apfelbeck, 1904]
References. Pterostichus convexiusculus: Apfelbeck, 1904: 261-262 (Velipoja, loc. typ.); Magistretti,
1965: 380; Jeanne, 1988: 74. Pterostichus apfelbecki: Schatzmayr, 1943a: 82 (Velipoja); Bousquet,
2003f: 507. Argutor convexiusculus: Jeanne, 1988: 74.
91.10. Pterostichus diligens (Sturm, 1824)
References. Pterostichus diligens: Hieke & Wrase, 1988: 72; Bousquet, 2003f: 507.
Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Çafa Markofçes” / “Albania leg.
Bischoff 1935”, 2 s. (MNHUB).
Notes. First detailed records for Albania.

155. 154 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
91.11. Pterostichus strenuus (Panzer, 1796)
References. Pterostichus strenuus: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 234; Hieke
& Wrase, 1988: 72; Bousquet, 2003f: 508. Argutor strenuus: Jeanne, 1988: 74.
Material studied. “Skutari Albania”, 3 s. (NMW); “Mustajbeg Velipoja Albania”, 5 s. (HNHM;
NMW); “Buschnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Pyla Thumana” /
“Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Subgenus Platypterinus Reitter, 1908
91.12. Pterostichus ottomanus Apfelbeck, 1908
91.12.1. Pterostichus ottomanus ottomanus Apfelbeck, 1908
References. Pterostichus ottomanus: Schatzmayr, 1943a: 106. Pterostichus ottomanus ottomanus:
Bousquet, 2003f: 508.
Material studies. “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalle
Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).
Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908;
Apfelbeck, 1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. First detailed
records for Albania.
Subgenus Platysma Bonelli, 1810
91.13. Pterostichus niger (Schaller, 1783)
91.13.1. Pterostichus niger niger (Schaller, 1783)
References. Pterostichus niger: Schatzmayr, 1930: 221 (Tomor Mt.); Csiki, 1940: 216 (Kula Ljums).
Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” /
“Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”,
1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Bizë, 27.VII.1966, 1
m. (MNST).
Additional material included (studied A. Vigna Taglianti). Fier, 4.VII.1993, 1 f., L. Latella
leg. (cAVT).
Notes. Bousquet (2003f: 509) has omitted this species for Albania. Here it is confirmed for the
country.
Subgenus Pseudomaseus Chaudoir, 1838
91.14. Pterostichus anthracinus (Illiger, 1798)
91.14.1. Pterostichus anthracinus anthracinus (Illiger, 1798)
References. Pterostichus anthracinus: Apfelbeck, 1909: 297 (Pasha Liman); Hieke & Wrase, 1988:
74. Pterostichus anthracinus anthracinus: Bousquet, 2003f: 510.
Material studied. 36 s. from different localities (MNHUB); “Albania Shkoder” / “25.V.1958
leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 22.II.1998 / 8.III.1998, 6 s., shaded lake edge
under leaves and wet, rotten wood, R. Ward leg. (CMNH).

156. COLEOPTERA: CARABIDAE: HARPALINAE 155
91.14.2. Pterostichus anthracinus biimpressus (Küster, 1853)
References. Pterostichus anthracinus biimpressus: Apfelbeck, 1904: 260 (Velipoja; Vlorë); Bucciareli
& Sopracordevole, 1958: 198 (Mamurash); Bousquet, 2003f: 510.
Notes. Magistretti (1965: 378) noted by mistake that the type locality of ssp. biimpressus is
“Almissa (Albania)”. In fact this Italian name concerns the town of Omiš at the Dalmatian
coast of Croatia.
91.15. Pterostichus gracilis (Dejean, 1828)
91.15.1. Pterostichus gracilis gracilis (Dejean, 1828)
References. Pterostichus gracilis: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 219;
Schatzmayr, 1943a: 84; Hieke & Wrase, 1988: 74. Pterostichus gracilis gracilis: Bousquet,
2003f: 511.
91.16. Pterostichus minor (Gyllenhal, 1827)
91.16.1. Pterostichus minor minor (Gyllenhal, 1827)
References. Pterostichus minor: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 220; Hieke
& Wrase, 1988: 74. Pterostichus minor minor: Bousquet, 2003f: 511.
91.17. Pterostichus nigrita (Paykull, 1790)
Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten
wood, R. Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
Subgenus Pterostichus s.str. [= Aphaon Heyden, 1906, syn. nov.]
Notes. The principal character used to separate Pterostichus s.str. and Aphaon (replacement
name for Phaon Heyden, 1883) is the number of lateral setiferous pores on pronotum
(Heyden, 1883). However, this character is inapplicable to divide the species of the two
subgenera. For instance, Pterostichus fasciatopunctatus (Creutzer, 1799), type species of the
genus and nominotypical subgenus has sometimes 1-3 marginal seta on the lateral border
of pronotum. On the other hand, P. rhilensis Rottenberg, 1874, the type species of Aphaon
also has 1-3 (sometimes 4) such setae. In fact, no other species from Aphaon shares this
character. The remaining four diagnostic features of Aphaon, e.g. transverse keel on the
last visible sternum in the males, number of the marginal pores on the same sternum in
both sexes, reduction of the scutellar stria and sharp basal border of elytra, cannot be of
subgeneric importance since these present also in other species of Pterostichus s.str. For
this reason Aphaon is treated as junior synonym of the latter subgenus and the following
nomenclatorial acts are introduced:
Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment from Pterostichus (Aphaon)
bischoffianus Jedlička, 1936
Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment from Pterostichus (Aphaon) brucki
Schaum, 1859

157. 156 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment from Pterostichus
(Aphaon) latifianus latifianus Apfelbeck, 1906
Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment from Pterostichus (Aphaon)
latifianus strupii J. Müller, 1935
Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment from Pterostichus
(Aphaon) lumensis lumensis Apfelbeck, 1905
Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment from Pterostichus (Aphaon)
lumensis litae Csiki, 1940
Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment from Pterostichus
(Aphaon) lumensis ljubetensis Apfelbeck, 1908
Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment from Pterostichus (Aphaon)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment from Pterostichus (Aphaon) meisteri
Reitter, 1885
Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment from Pterostichus (Aphaon)
merklii (J. Frivaldszky, 1879)
Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment from Pterostichus (Aphaon)
pentheri Apfelbeck, 1918
Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment from Pterostichus (Aphaon)
reiseri Ganglbauer, 1889
Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment from Pterostichus
(Aphaon) rhilensis rhilensis Rottenberg, 1874
Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment from Pterostichus (Aphaon)
rhilensis kourili Mařan, 1933
Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment from Pterostichus (Aphaon)
rhilensis vitosensis Mařan, 1933
Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment from Pterostichus (Aphaon) walteri
Reitter, 1883
91.18. Pterostichus bischoffianus Jedlička, 1936
References. Pterostichus bischoffianus: Jedlička, 1936: 167 (Gjalicë Mt., loc. typ.); Schatzmayr,
1943a: 109 (Gjalicë Mt.); Bousquet, 2003f: 515.
Material studied. “Gjallica” / “Albania leg. Bischoff 1936” / “Typus” / “bischoffianus sp.n.
type det. Ing. Jedlička”, 1 m. syntype (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 2
m. (MNHUB).
91.19. Pterostichus brucki Schaum, 1859
References. Pterostichus brucki: Apfelbeck, 1918c: 166 (North Albanian Alps, Vermosh, 1100 m);
Jedlička, 1936: 167 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 216 (Gjalicë Mt.); Schatzmayr, 1943a:
107 (Koritnik Mt.; Gjalicë Mt.); Hieke, 1981: 80; Hieke & Wrase, 1988: 78; Bousquet, 2003f: 511.
Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);
“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp.’18
Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Gjalica” / “Albania leg. Bischoff 1935”, 1 s. (NMW);
“Koritnik” / “Albania leg. Bischoff 1935”, 8 s. (NMW; MNHUB); “Gjallica” / “Albania leg.

158. COLEOPTERA: CARABIDAE: HARPALINAE 157
Bischoff 1935”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1936”, 1 s.
(MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 32 s. (MNHUB); “Shkalla Harushe” /
“Korab” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
91.20. Pterostichus latifianus Apfelbeck, 1906
91.20.1. Pterostichus latifianus latifianus Apfelbeck, 1906 [= munella Jedlička, 1936]
References. Pterostichus latifianus: Apfelbeck, 1906: 1663 (North Albania, Merdita: Mal Shenjt
Mt. and Munella Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 502 (Mal i Shenjt Mt.: top region,
1500 m); Schatzmayr, 1930: 256; Jedlička, 1936: 168 (Merdita; Dërze); Jedlička, 1939: 52;
Schatzmayr, 1943a: 109 (Merdita; Dërze). Pterostichus malissorum ab. munella: Jedlička, 1936: 169
(Munella Mt., loc. typ.); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Munella Mt.). Pterostichus
latifianus latifianus: Bousquet, 2003f: 485.
Material studied. “Apfelb. Albanien”, 2 syntypes (MNHUB); “Latif Alban. sept.”, 4 s.
(DEI; HNHM); “Latif Munela” / “Latif Alban. spt.”, 1 s. (HNHM); “Latif Orosi” / “latifianus
typus! Apfelb.” / “typus”, 1 f. syntype (NMW); “Latif Orosi”, 2 syntypes (NMW); “Merdita
Mal Shêit” / “latifianus Apf. cotype det. Apflb.”, 1 m. syntype (MNHUB); “Merdita M.
Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (HNHM); “Merdita M. Scheit” / “Coll.
Apfelbeck”, 12 syntypes (HNHM); “Merdita M. Scheit”, 4 s. probably syntypes (NMW);
“Merdita Munela”, 8 syntypes (NMW; MNHUB); “Merdita Munela” / “Apfelb. Albanien”,
1 syntype (MNHUB); “Merdita Zebia”, 4 syntypes (MNHUB); “Merdita Zebia” / “Wgth.
1906 Merdita”, 10 s. (NMW); “Merdita Oroschi”, 10 s. (HNHM; NMW); “Derzë” / “Albania
leg. Bischoff 1935”, 67 s. (MNHUB).
Notes. Jedlička (1939: 52) stated that his P. malissorum ab. munella is identical with P. latifianus,
based on revision of type material, but subsequent authors (Schatzmayr, 1943a; Lorenz, 1998:
261) overlooked his opinion.
91.20.2. Pterostichus latifianus strupii J. Müller, 1935
References. Pterostichus latifianus strupii: Müller, 1935: 156 (Qaf e Stames, northeast of Kruja,
loc. typ.); Schatzmayr, 1943a: 109 (Qaf e Stames); Bousquet, 2003f: 485.
Notes. The ssp. struppi may be a distinct species since its type locality (Qaf e Stames Pass) is
situated in the geographically isolated massif of Skënderbe Mt.
91.21. Pterostichus lumensis Apfelbeck, 1905
91.21.1. Pterostichus lumensis lumensis Apfelbeck, 1905
References. Pterostichus lumensis: Apfelbeck, 1905: 242 (Albania / Kosovo border, Koritnik
Mt., loc. typ.); Apfelbeck, 1907: 637 (Koritnik Mt., loc. typ.); Apfelbeck, 1917: 225, note 1
(Koritnik Mt.); Apfelbeck, 1930: 169, note 1 (Koritnik Mt.); Schatzmayr, 1930: 255; Jedlička,
1936: 167 (Koritnik Mt.; Korab Mt.); Schatzmayr, 1943a: 108 (Koritnik Mt.). Pterostichus latifianus
ssp.?: Schatzmayr, 1930: 256 (Korab Mt.). Pterostichus lumensis lumensis: Drovenik & Peks, 1994:
67 (Koritnik Mt.); Drovenik & Peks, 1999: 78 (Koritnik Mt.); Bousquet, 2003f: 515.
Material studied. “Latif Koritnik”, 2 syntypes (NMW); “Latif Alban. spt.”, 1 s. probably
syntype (HNHM); “Latif Alban. spt.” / “P. lumensis Apfelb. Typen”, 1 s. probably syntype
(HNHM); “Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 3 s. (NMW); “Alban. Exp. Korab 1918
24.-31.VII.”, 16 s. (NMW); “Korab Albania Bischoff ”, 1 s. (NMW); “Bjeshka Matroxh”, 11 s.

159. 158 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
(MNHUB); “Korab” / “Albania leg. Bischoff 1935”, more than 120 s. (MNHUB); “Gjallica”
/ “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”,
more than 120 s. (MNHUB).
Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck,
1907b), and the older one is correctly treated as valid. Two specimens (1 m., 1 f.) labelled
“Latif Alban. sept.” and identified as Pterostichus djumanus Apfelbeck were found in DEI.
They are put provisionally here, since both specimens resemble P. lumensis. Pterostichus djumanus
Apfelbeck is nomen nudum.
91.21.2. Pterostichus lumensis litae Csiki, 1940
References. Pterostichus lumensis litae: Csiki, 1940: 216 (Gjalicë Mt., loc. typ.; Korab Mt., loc.
typ.); Schatzmayr, 1943a: 108 (Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 515.
Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 m. syntype
(HNHM); “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 4 syntypes (HNHM); “Albania
Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 21 syntypes (HNHM).
? Pterostichus lumensis ljubetensis Apfelbeck, 1908 [= chalybaeus Apfelbeck, 1908]
References. Pterostichus lumensis ljubetensis: Schatzmayr, 1930: 255; Bousquet, 2003f: 515.
Notes. Apfelbeck published three descriptions of this taxon (Apfelbeck, 1908; Apfelbeck,
1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. All data in the literature
and collections about ssp. ljubetensis come from the subalpine and alpine regions of the Shar
Planina Mts. that do not extend into Albania. Here, P. lumensis ljubetensis is treated as doubtful
for the fauna of Albania.
91.22. Pterostichus malissorum Apfelbeck, 1905 [= nigripes Jedlièka, 1936]
References. Pterostichus malissorum: Apfelbeck, 1905: 241-242 (Çukall Mt., loc. typ.); Apfelbeck,
1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1907b: 637 (Çukall Mt., loc.
typ.); Schatzmayr, 1930: 255; Jedlička, 1936: 169 (Malissora; Çukall Mt.; Cafa Valbones; Cafa
Pejs; Maja Pejs); Schatzmayr, 1943a: 108 (Çukall Mt.; Malissora; Cafa Valbones; Cafa Pejs;
Maja Pejs); Bousquet, 2003f: 515. Pterostichus latifianus ab. nigripes: Jedlička, 1936: 168 (North
Albanian Alps: Quf e Valbone, loc. typ.); Jedlička, 1939: 53. Pterostichus latifianus strupii (=
nigripes): Schatzmayr, 1943a: 109 (Quf e Valbone).
Material studied. “Latif Alban. sept.”, 4 s. (DEI; HNHM); “Latif Cukali” / “Coll.
Apfelbeck”, 1 m. syntype (HNHM); “Latif Cukali”, 2 syntypes (NMW); “Maranai Apfelb.”,
1 s. (HNHM); “Maranai Apf.”, 1 s. (HNHM); “Apfelb. Albanien”, 1 m. syntype (MNHUB);
“Apfelb. Albanien” / “Alban. spt. Maranaj”, 1 f. probably syntype (MNHUB); “Apfelb. Alban
spt” / “Maranaj Geb.”, 2 s. (HNHM); “Apfelb. Alban. spt. IV.V.’05” / “Maranaj”, 6 s. (NMW);
“Kunor Vukai Zukali Albanien Lona Giugno 1914”, 6 s. (NMW; MNHUB); “Cukale Albania
Bischoff ”, 12 s. (MNHUB); “Cukale-Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB);
“Çafa Stamës” / “leg. Strupi Albania 1934” / “Cotype” / “latifianus a. nigripes n. ab. det. Ing.
Jedlička”, 1 f. designated as syntype of Pterostichus latifianus ab. nigripes Jedlička, 1936
(MNHUB); “Cukale” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Çafa Valbonës” /
“Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 3
s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Maranaj” /
“Albania leg. Bischoff 1937”, 23 s. (MNHUB); “Boge ober D. Quelle” / “Albania leg. Mueller

160. COLEOPTERA: CARABIDAE: HARPALINAE 159
1939”, 1 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 10 s. (MNHUB);
“Maja e Dragomirit” / “Albania leg. Mueller 1939”, 4 s. (MNHUB); North Albanian Alps,
Maj’e Redohimes, 1900-2200 m, 28.V.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); North
Albanian Alps, Maj’e Redohimes, 2200-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg.
(NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov
leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron &
B. Petrov leg. (NMNHS).
Notes. Apfelbeck published two descriptions of this species (Apfelbeck, 1905; Apfelbeck,
1907b), and the older one is correctly treated as valid. Jedlička (1939: 52) stated that his P.
latifianus ab. nigripes Jedlička, 1936 is identical with P. malissorum Apfelbeck, 1905. Later,
Schatzmayr (1943a), however, synonymized P. latifianus ab. nigripes Jedlička, 1936 with P. latifianus
strupii J. Müller, 1935. I support the view of Jedlička because the type locality of the former is
quite close to the known findings of P. malisstorum, whereas P. latifianus strupii lives further in
the south: Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. of Pterostichus
(Pterostichus) malissorum Apfelbeck, 1905.
91.23. Pterostichus pentheri Apfelbeck, 1918
References. Pterostichus pentheri: Apfelbeck, 1918c: 161, 166 (North Albanian Alps, loc. typ.);
Schatzmayr, 1930: 255; Jedlička, 1936: 168 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë
Mt.); Schatzmayr, 1943a: 108 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë Mt.); Bousquet,
2003f: 515.
Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Prokletija Albanien Dr. Penther”, 2
syntypes (NMW); “Prokletija” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes (DEI;
NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes
(NMW); “Maja e Jezercë” / “Albania leg. Bischoff 1931”, 7 s. (MNHUB); “Maja e Jezercë
Çafa Pejs-Çafa Gstars” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Çafa e Markofçes” /
“Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Maja e Jezercë” / “Albania leg. Bischoff 1935”,
12 s. (MNHUB); more than 90 s. from locality “Shkëlzën” (MNHUB); North Albanian Alps,
Maj’e Redohimes, 2400-2550 m, 29.V.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-
Rubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m,
11.VII.1993, 2 m., 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2000 m, 14.VII.1993, 2
m., A. Gobetti leg. (cAVT).
91.24. Pterostichus reiseri Ganglbauer, 1889
References. Pterostichus reiseri: Apfelbeck, 1918c: 160, 166 (North Albanian Alps, Vermosh,
Grebeni, 1100-1700 m); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Grebeni).
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 2 s.
(NMW); “Vermoša Albanien Dr. Penther”, 2 s. (NMW); “Vermosa 20-21.VII.” / “Alban.
montenegr. Grenze, Penther’14”, 1 s. (NMW).
Notes. Bousquet (2003f: 516) has omitted P. reiseri for Albania. Here the species is confirmed
for the country. The record “Albanien: Žljeb plan.” (Jedlička, 1936: 168) refers to the massif
shared among Montenegro and the Kosovo Region of Serbia.

161. 160 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Species insertae sedis
? Pterostichus epiroticus Csiki, 1930 [= corax Ganglbauer, 1889]
References. Pterostichus epiroticus: Schatzmayr, 1943a: 107.
Notes. The record “Peristerigebirge bei Jannina (Albanien)” (Schatzmayr, 1943a) is based on an older
record of Apfelbeck (1904: 267 sub P. corax). Currently this record concerns the territory in
Northwestern Greece. In NMW, besides specimens from Central Greece (Oeta Mt.; Korax Mt.), I
found five recently collected specimens from the region near to Joannina and Metsovon, which is very
close to the Albanian territory. Therefore, P. epiroticus is treated here as likely to be found in Albania.
92. Genus Speluncarius Reitter, 1886
Subgenus Hypogium Tschitschérine, 1900 [= Hypogeobium Tschitschérine, 1903]
92.1. Speluncarius albanicus (Tschitschérine, 1900)
References. Platysma albanicum: Tschitschérine, 1900: 49-51 (“Albanie”, loc. typ.). Pterostichus
albanicus: Apfelbeck, 1904: 236-237; Schatzmayr, 1930: 329. Tapinopterus albanicus: Reitter, 1914:
262; Müller, 1937: 134; Schatzmayr, 1943a: 131; Jeannel, 1950: 157, 159-160. Hypogeobium
albanicum: Jeannel, 1950: 160, 162; Jeannel, 1953: 11. Speluncarius albanicus: Sciaky, 1982: 28;
Casale & Giachino, 1991: 215-216; 219; Bousquet, 2003f: 516.
Material studied. “Albania” / G.C.Champion Coll. B.M.1927-409” / “Platysma (Hypogium)
albanicum m. Typ. Tschitscherin det” / “syntype” (round blue coloured label), 1 f. (BMNH).
Notes. The type locality of S. albanicus remains not yet specified.
Subgenus Speluncarius s.str.
92.2. Speluncarius minutulus J. Müller, 1937
References. Tapinopterus minutulus: Müller, 1937: 132-123 (Shkëlzen Mt., loc. typ.); Schatzmayr, 1943a:
128 (Shkëlzen Mt.); Turin, 1981: 127. Speluncarius minutulus: Jeanne, 1982: 41-42 (Shkëlzen Mt.); Sciaky,
1982: 20; Drovenik & Peks, 1994: 68; Drovenik & Peks, 1999: 79; Bousquet, 2003f: 516.
Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 f. designated as paralectotype
by P.M. Giachino (MNHUB).
Notes. Drovenik & Peks (1994) and Drovenik & Peks (1999) noted S. minutulus for Montenegro
by mistake because it is a local endemic species for Northeast Albania.
93. Genus Stomis Clairville, 1806
Subgenus Stomis s.str.
93.1. Stomis pumicatus (Panzer, 1796)
93.1.1. Stomis pumicatus pumicatus (Panzer, 1796)
References. Stomis pumicatus: Apfelbeck, 1904: 213 (Velipoja); Hieke & Wrase, 1988: 68;
Bousquet, 2003f: 517.

162. COLEOPTERA: CARABIDAE: HARPALINAE 161
Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Bojana
Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB).
93.2. Stomis rostratus (Duftschmid, 1812)
93.2.1. Stomis rostratus rostratus (Duftschmid, 1812)
References. Stomis rostratus: Apfelbeck, 1918c: 166 (North Albanian Alps, Grebeni near
Vermosh, 1100-1800 m); Magistretti, 1965: 362; Bousquet, 2003f: 518.
Material studied. “Grebeni 16.VI.” / “Alban. montenegr. Grenze, Penther’14”, 3 s. (NMW);
“Alban. Vermosa Penther”, 2 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr.
Grenze, Penther’14”, 4 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze,
Penther’14”, 4 s. (NMW); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).
94. Genus Tapinopterus Schaum, 1858
Subgenus Tapinopterus s.str.
Notes. The species from this subgenus are placed here in three groups of species in agreement
with a forthcoming arrangement proposed by P. M. Giachino and L. Picciau (personal
communication).
I. South Aegean groups.
Species group “extensus”
# Tapinopterus extensus (Dejean, 1831)
# Tapinopterus extensus abreshensis (Lutshnik, 1916) [= convexiusculus Apfelbeck, 1904]
References. Pterostichus extensus convexiusculus: Apfelbeck, 1904: 251 (“Turcia – Abresch”, loc.
typ.); Schatzmayr, 1943a: 127.
Notes. According to Schatzmayr (1943a) the type locality of this race (“Abresch”) must be
somewhere between Albania and Palestine. Bousquet (2003f: 519) has noted the subspecies
for Greece. Based on Apfelbeck’ short description (Apfelbeck, 1904: 251 sub Pterostichus extensus
convexiusculus), I suppose that ssp. abreshensis is not subordinate to T. extensus s.l., taking into
consideration also the fact that Apfelbeck (1904) compared the former with T. kaufmanni
Ganglbauer, 1897. Dealing with the taxa of Tapinopteris extensus s.l., Mařan (1939b: 298, 303)
without hesitation separated T. extensus abreshensis from the remaining races of the species.
Only a revision of Apfelbeck’s type (probably in Museum Sarajevo) could establish the
systematic position of the taxon in question. Presently, both the species and subspecies are
removed from the list of the Albanian Carabidae.
94.1. Tapinopterus extensoides Straneo, 1936 [= extensoides Jedlièka, 1936]
References. Tapinopterus extensoides Straneo: Straneo, 1936: 155-156 (Llogora, typ. loc.); Müller,
1937: 132, note 1 (Llogora; cave “Paftali” near Berat; Shëndelli Mt.); Schatzmayr, 1943a: 127
(Llogora; cave “Paftali” near Berat; Shëndelli Mt.). Tapinopterus extensoides Jedlička: Jedlička,
1936: 169 (Llogora, loc. typ.); Turin, 1981: 127. Tapinopterus extensus extensoides “Straneo et

163. 162 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Jedl.”: Mařan, 1939b: 295-299, 302-303 (Llogora). Tapinopterus extensus extensoides Jedlička:
Bousquet, 2003f: 519.
Material studied. “Logara Juni” / “Albania leg. Bischoff 1932”, 1 m. syntype (NMW); “Maja
e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Reza e Kanalit Logara, Alban. m.”
/ “Albania leg. Bischoff 1932”, 1 f. syntype (NMW); “Logara juni” / “Albania leg. Bischoff
1932”, 3 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 7 s. (MNHUB); “Llogora,
Alb. Mer.” / “Albania leg. Bischoff 1935”, 1 f. syntype of extensoides Jedlička, 1936 (MNHUB);
“Llogora” / “leg. Bischoff Albania 1936”, 17 s. (MNHUB); “Shen Ili” / “leg. Bischoff Albania
1936”, 1 s. (MNHUB); “Borshe” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “Shpella Buz
i Lugit Mal i Piqs” / “Borshe” / “Albania leg. Bischoff 1940”, 2 s. (MNHUB); “Albania,
Periferi Tepelenë, a gorge 1.5 km W of Progonat,” / “785 m, 9.X.2004, Z. Fehér, J. Kontschán
& D. Murányi (No. 133)”, 2 f. (HNHM).
Notes. In 1936 Jedlička (1936) and Straneo (1936) described T. extensoides having at disposal
identical specimens collected by Bischoff from the region of Llogora. Later, Müller (1937:
132, note 1) stated that the Straneo’s description has priority over that of Jedlička. This opinion
is correct in spite of Mařan’s statement that both are authors of this taxon (Mařan, 1939b:
302). It is Mařan who first regarded extensoises as subspecies of T. extensus. However, in my
view T. extensoides Straneo is clearly distinct from all the other subspecies of T. extensus by an
evident apomorphic character – 2 (instead 1) posterior supraorbital setiferous pores (see also
Straneo, 1936). The check of all specimens at hand demonstrated that this character is constant
(as only a single female from HNHM has a single left and two right posterior supraorbital
pores) so that it may define a separate species: Tapinopterus (Tapinopterus) extensoides (Straneo,
1936), stat. rest. from Tapinopterus (Tapinopterus) extensus extensoides (Straneo, 1936).
Species group “protensus”
94.2. Tapinopterus protensus (Schaum, 1857)
Notes. Bousquet (2003f: 520) noted that T. protensus aetolicus Ganglbauer, 1889 inhabits
Montenegro, and not Albania. In spite of that Schatzmayr (1943: 122) cited it for the studied
country (see below). Here, the species records and reference data from Albania are referred to
two defined and two undefined subspecies. The occurrence of T. protensus in Montenegro
needs serious confirmations.
94.2.1. Tapinopterus protensus peristericus Apfelbeck, 1901
References. Tapinopterus aetolicus peristericus: Müller, 1935: 153 (Shëndelli Mt. near Tepelenë).
Material studied. “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff”, 2 s.
(MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 6 s. (MNHUB);
“Shen Ili” / “leg. Bischoff Albania 1936”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg.
Bischoff 1937”, 31 s. (MNHUB).
Notes. This subspecies is confined to the area between the valleys of Drina River and Vijosë
River and lives also in Northwestern Greece. Müller (1935: 153-154) noted the identity of
syntypes of T. peristericus Apfelbeck, 1901 with specimens from the Shëndelli Mt. In MNHUB,
I found specimens from Shëndelli Mt. labelled “aetolicus sbsp. peristericus det. J. Müller”, and
others - “aetolicus subsp. nova m. det. J. Müller”.

164. COLEOPTERA: CARABIDAE: HARPALINAE 163
94.2.2. Tapinopterus protensus tomoricensis J. Müller, 1935
References. Tapinopterus aetolicus tomoricensis: Müller, 1935: 153-154 (Tomor Mt., loc. typ.).
Tapinopterus thessalicus tomoricensis: Mařan, 1940b: 59 (Tomor Mt.). Tapinopterus protensus tomoricensis:
Schatzmayr, 1943a: 122 (Tomor Mt.); Bousquet, 2003f: 520.
Material studied. “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 2 syntypes
(NMW); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 8 syntypes (NMW);
40 s. from Tomor Mt. (MNHUB).
! Tapinopterus protensus ssp.1
Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB).
Notes. Josef Müller has noted on the label that the specimen from “Mal i Dajtit” belongs to
an undescribed taxon. This specimen has gigantic body dimensions (21.5 mm) in comparison
with the other races from Albania, but more material is needed for the correct classification
of this local population.
! Tapinopterus protensus ssp.2
References. Tapinopterus aetolicus: Müller, 1935: 154 (Reza e Kanalit Mt. near Llogora).
Tapinopterus protensus aetolicus: Schatzmayr, 1943a: 122 (Reza e Kanalit Mt. near Llogora).
Material studied. “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Logara” / “leg.
Bischoff Albania 1934”, 2 s. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 s.
(MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 7 s. (MNHUB).
Notes. This still not defined race inhabits the mountains west of the valleys of Vijosë River
and Drina River. Müller has identified the specimens from “Llogara” as ssp. aetolicus, and that
from “Maja e Tartarit” as “aetolicus sbsp. valonensis m. det. J. Müller”. Both populations from
Albania may not belong to the ssp. aetolicus (as Müller stated, 1935), if considering the
geographical factor, since the type locality of ssp. aetolicus is the Korax Mt. (Central Greece),
situated nearly 270 km south of the area of Llogora.
II. North Aegean groups.
Species group “balcanicus”
94.3. Tapinopterus bischoffi J. Müller, 1936
References. Tapinopterus bischoffi: Müller, 1936: 106-107 (Mal i Dajtit Mt., loc. typ.); Schatzmayr,
1943a: 127 (Mal i Dajtit Mt.); Turin, 1981: 127; Bousquet, 2003f: 519.
Material studied. “Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Mal
i Dajtit” / “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Mal i Drini” / “leg. Bischoff Albania
1934” / “cotype” / “Tapinopterus bischoffi m. det. J. Müller”, 1 m. (NMW); “Rushkull” / “Albania
leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll. Bischoff”,
5 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes”
/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1939”,
7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); “AL (Kruje)
Paß ca. 5 km ne. Kruje 700-900 m, 1.V.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB); Mal i
Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 m., 2 f., L. Penev leg. (NMNHS).

165. 164 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
# Tapinopterus comita Jedlička, 1935
References. Tapinopterus eomita sic!: Turin, 1981: 127. Tapinopterus comita: Bousquet, 2003f: 519.
Notes. I do not find any exact records in the literature or material in the collection of this
species from Albania, thus the indications of Turin (1981) and Bousquet (2003f) for the country
may be treated as incorrect. T. comita inhabits only the Pelister Mt. (Jedlička, 1935; Drovenik &
Peks, 1994: 68), a massif shared between Macedonia and Greece. The species is removed
from the list of the Albanian Carabidae.
94.4. Tapinopterus detonii Schatzmayr, 1943
References. Tapinopterus detonii: Schatzmayr, 1943b: 110 (Mal i Krujes Mt., 1000 m, loc. typ.);
Turin, 1981: 127; Bousquet, 2003f: 519.
Material studied. “Albania”, 1 s. (DEI).
Notes. Further taxonomic work is needed to demonstrate the differentiation of T. detonii
from T. bischoffi J. Müller, 1936, since the former seems not well defined and may be a subspecies
or synonym of the latter.
94.5. Tapinopterus dochii (Apfelbeck, 1906)
References. Pterostichus dochii: Apfelbeck, 1906: 1663-1664 (Merdita: Mal i Shenjt Mt., loc. typ.;
Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 499, 502 (Mal i Shenjt Mt.: top region, 1500 m).
Tapinopterus dochii: Jedlička, 1935: 186 (Korab Mt.); Müller, 1936: 107 (Korab Mt.); Schatzmayr,
1943a: 126 (Mal i Shenjt Mt.; Zepë Mt.; Munella Mt.); Hieke, 1981: 81; Bousquet, 2003f: 519.
Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05” / “Tapinopterus dochii
Apf.”, 1 f. syntype (NMW); more than 100 s. from localities “Berishe”, “Cucale”, “Maja e
Shpatit”, “Cafa e Sharit”, “Maja Shales Vogel Iballe”, “Korab”, “Munela”, “Derzë”, “Taja”,
“M. Mulecifit”, “Merdita M. Sheit” (MNHUB).
[Tapinopterus miridita (Apfelbeck, 1904)]
Notes. The only records for T. miridita s.l. from Albania are those of Hieke (1981: 81) and
Bousquet (2003f: 520). However, the present author cannot find any exact data in the literature
or collections to prove the species occurrence in Albania. The type locality “Sveti Troice”
(Apfelbeck, 1904: 245), or more correct “Sveta Trojica”, concerns a monastery near the village
of Mušutište, a place situated in the northern slopes of the Shar Planina Mt. (in the Kosovo
Region of Serbia) (see also Jedlička, 1935: 186). Oddly enough, the “loc. typ.” and known
records of the species are quite removed from the region of Mirdita (situated in the south part
of North Albania, south of the Drin River), after which the species was named. T. miridita is
treated here as likely to be found in the country.
[Tapinopterus miridita miridita (Apfelbeck, 1904)]
Notes. The occurrence of this race in the marginal northeastern regions of Albania (the
Koritnik Mt.; the Gjalicë Mt.) is possible.
[Tapinopterus miridita jablanicensis Mařan, 1935]
Notes. This race was described from the Macedonian part of the Jablaniča Mt. (shared between
Albania and Macedonia), but so far the subspecies is not properly cited for Albania.

166. COLEOPTERA: CARABIDAE: HARPALINAE 165
# Tapinopterus miridita jacupicensis Jedlička, 1935
Notes. Bousquet (2003f: 520) cited this subspecies for Albania. However, this taxon is a local
endemic for the massif of the Jakupiča Mt. situated in the central part of Republic of
Macedonia. Probably the author in question has had in mind the spp. jablanicensis when he
mentioned ssp. jacupicensis for Albania. The latter subspecies is here excluded from the list of
the Albanian Carabidae.
94.6. Tapinopterus monastirensis Reitter, 1913
94.6.1. Tapinopterus monastirensis gestroi J. Müller, 1923
References. Tapinopterus gestroi: Müller, 1923: 110-111 (Tomor Mt.: Toik, loc. typ.). Tapinopterus
monastirensis gestroi: Müller, 1936: 107; Mařan, 1939e: 82, 84 (Tomor Mt.: Toik); Schatzmayr,
1943a: 126 (Tomor Mt.); Bousquet, 2003f: 520.
Material studied. “Albania”, 1 s. (DEI); “Tomorica Kulmak” / “Albania 1922 Ravasini,
Lona”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1
s. (NMW); “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);
“Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW); “Tomor-
Tojk juli 1932” / “Albania leg. Bischoff 1932”, 32 s. (MNHUB); “Tomor” / “Albania leg.
Bischoff 1932”, 17 s. (MNHUB).
Tribe SPHODRINI Laporte, 1834
Subtribe ATRANOPSINA Baehr, 1982
95. Genus Platyderus Stephens, 1827
Notes. The present knowledge about the Balkan representatives of this genus poorly
corresponds to the actual status of the species.
Subgenus Platyderus s.str.
95.1. Platyderus dalmatinus L. Miller, 1881
95.1.1. Platyderus dalmatinus dalmatinus L. Miller, 1881
References. Platyderus dalmatinus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme,
300 m, and Vorfai Siperme, 700 m).
Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 m. (NMW).
Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Here, it is
confirmed for the fauna of Albania.
95.2. Platyderus graecus (Reiche & Saulcy, 1855)
References. Platyderus graecus: Apfelbeck, 1909: 297 (Vlorë).
Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Further study is
needed to confirm its occurrence in the country.

167. 166 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
95.3. Platyderus minutus (Reiche & Saulcy, 1855)
95.3.1. Platyderus minutus obtusangulus J. Müller, 1923
References. Platyderus minutus: Apfelbeck, 1907a: 498, 501 (Mal i Shenjt Mt.: top region, 1500
m). Platyderus minutus obtusangulus: Müller, 1923: 112 (Tomor Mt.: Kulmak, loc. typ.); Hovorka
& Sciaky, 2003a: 523.
Material studied. “Albania”, 1 f. (BMNH).
95.4. Platyderus rufus (Duftschmid, 1812)
95.4.1. Platyderus rufus rufus (Duftschmid, 1812)
References. Platyderus rufus: Apfelbeck, 1904: 279 (Velipoja near Lezhë); Hieke & Wrase, 1988:
89. Platyderus rufus rufus: Hovorka & Sciaky, 2003a: 523.
Material studied. Nearly 50 s. from different localities(MNHUB); Rojan, 6.III.1961, 1 f. (MNST).
Subtribe CALATHINA Laporte, 1834
96. Genus Calathus Bonelli, 1810
Subgenus Bedelinus Ragusa, 1885
96.1. Calathus circumseptus Germar, 1824
References. Calathus circumseptus: Hovorka & Sciaky, 2003b: 524.
Material studied. “Albania”, 1 s. (HNHM); “Durazzo”, 1 s. (HNHM); “Apfelb. 1893 Durazzo”,
1 s. (HNHM); “Berat” / “Albania leg. Bischoff 1931”, 4 s. (MNHUB); “Tepelene juni 1932” /
“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Erzen” / “leg. Bischoff Albania 1934”,
1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania
leg. Bischoff 1935”, 3 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);
“Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Saranda District, Qeparo,
4-5.V.1994, 2 s., P. Stoev leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg.
(NMNHS); Kruja Castle, 16.XI.1997, 2 s., R. Ward leg. (CMNH).
Notes. First detailed records for Albania.
Subgenus Calathus s.str.
96.2. Calathus bosnicus Ganglbauer, 1891
References. Calathus bosnicus: Hovorka & Sciaky, 2003b: 524.
Material studied. “Prokletija 1300-2000 m” / “Alban.-montenegr. Grence, Penther’14”, 2 s.
(NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Gjalica Ljums 17-26.VI.”, 1
s. (NMW); “Llogora…”, 1 s. (NMW); 60 s. from localities “Shkëlzën”, “Çafa Markofçes”,
“Maja Pejs”, “Korab”, “Maja e Jezercoë”, “Valbona Çerem” (MNHUB); North Albanian Alps,
Bogë, 1000-1100 m, 5-9.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS).
Notes.FirstdetailedrecordsforAlbania.Therecord“Llogora…”eitherconcernsanothercongener
or is “patria falsa”, since the locality is quite removed in the south from the range of the species.

168. COLEOPTERA: CARABIDAE: HARPALINAE 167
[Calathus distinguendus Chaudoir, 1846]
Notes. According to Battoni & Vereschagina (1984: 141) this species may live in Albania, but
so far this statement is not proved.
96.3. Calathus ellipticus Reitter, 1889
96.3.1. Calathus ellipticus ellipticus Reitter, 1889
Material studied. “Çafa e Dhembellit”, 2 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff
1932”, 26 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB);
“Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg.
Bischoff 1937”, 2 s. (MNHUB).
Notes. New species for the fauna of Albania. After a comparison of type material with
specimens from Albania, D.W. Wrase (personal communication) concluded that they are
conspecific, being in agreement with specimens from Western Greece. A revision of all related
taxa involved (including also C. giganteus and C. korax) is needed to clarify the status of the
Albanian population.
96.4. Calathus fuscipes (Goeze, 1777)
96.4.1. Calathus fuscipes graecus Dejean, 1831 [= latus (Audinet-Serville, 1821) nec Linnaeus, 1758]
References. Calathus fuscipes: Apfelbeck, 1904: 284 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë;
Berat); Csiki, 1940: 217 (Shkëlzen Mt.; Kula Ljums); Csiki, 1943: 214; Schatzmayr, 1943b: 110
(“Poggio Boschetto”; Kruja; Dukati); Battoni & Vereschagina, 1984: 152; Hieke et al., 1986:
189 (Tiranë; Tomor Mt.: Abbas Ali Monastery; Bizë near Shëngjergji; Lurë e Vjetër near Lura
Mt.); Hieke & Wrase, 1988: 89. Calathus fuscipes var. graecus: Wohlberedt, 1909: 127 (around
Liqen i Shkodrës). Calathus fuscipes graecus: Schatzmayr, 1937: 27 (Gjirokastër); Hovorka &
Sciaky, 2003b: 525. Calathus fuscipes var. latus: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Calathus
fuscipes fuscipes: Hovorka & Sciaky, 2003b: 525.
Material studied. More than 180 s. from different localities (MNHUB); L. Selitës, 8-9.VII.1961,
5 s. (MNST); Lurë, 18-25.VII.1961, 2 s. (MNST); Theth, 1-8.VIII.1961, 2 f. (MNST); Ibë,
14.VII.1962, 1 f. (MNST); Shkodër, 15.V.1963, 1 m.; Mal i Dajtit Mt., 10-11.V.1965, 1 m., 1 f.
(MNST); 2 km N Durrës, 24.V.1993, 1 m., 1 f., P. Stoev & D. Zaprjanova leg. (NMNHS);
North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 f., P. Beron & B. Petrov
leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 f., P. Beron & B.
Petrov leg. (NMNHS); Reshen, 11.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Cafasan,
1230 m, 27.VII.1993, 3 m., P. Stoev & D. Zaprjanova leg. (NMNHS); Tiranë, botanical garden,
8.V.1995, 1 m, 1 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt.,
450 m, 11.V.1995, 2 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m,
11.V.1995, 2 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 /
8.II.1998, 5 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., 3 f., A.
Paparistos leg. (cAVT).
Notes. According to A. Vigna Taglianti (personal communication) the populations from Albania
belong to ssp. graecus. Besides the material studied above, in NMW I have seen more specimens,
which remained unregistered.

169. 168 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
96.5. Calathus giganteus Dejean, 1828
References. Calathus giganteus: Apfelbeck, 1904: 282 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);
Schatzmayr, 1937: 2, 6; 21 (Vlorë; Sopoti near Gjirokastër); Hovorka & Sciaky, 2003b: 525.
Material studied. “Llogora juni” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tepelene
juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania
1934”, 1 s. (MNHUB); “Llogora” / “Albania leg.Bischoff 1936”, 3 s. (MNHUB); “Maja e Konispol
Seraqin” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Llogora
Pass, 1025 m, 11.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).
Additional material included (studied S. Hristovski and D.W. Wrase). “Logara” / “Albania,
leg Bischoff 1932”, 1 s. (MNHUB); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,
1 s., leg. M. Komnenov (cSH).
? Calathus glabricollis Dejean, 1828
Material studied. “coll. L.W. Schaufuß” / “Calathus glabricollis Albanien verus”, 1 m. (MNHUB).
Notes. It is not sure if the record above concerns the present territory of Albania, so the
species is treated here as doubtful for the country.
96.6. Calathus korax Reitter, 1889
References. Calathus corax: Schatzmayr, 1937: 2, 7; 21 (Gjirokastër).
Notes. Hovorka & Sciaky (2003b: 525) have omitted C. korax for Albania. Further collections
are needed to confirm its occurrence in the country.
96.7. Calathus ravasinii J. Müller, 1935
96.7.1. Calathus ravasinii ravasinii J. Müller, 1935
References. Calathus ravasinii: Müller, 1935: 152-153 (Tomor Mt.: Kulmak, loc. typ.); Schatzmayr,
1937: 2, 7, 22 (Tomor Mt.); Turin, 1981: 136; Hovorka & Sciaky, 2003b: 526.
Material studied. “M. Toik Alban. mer.”, 1 s. probably syntype (NMW); “Tomorica Kulmak”
/ “Albania 1922 Ravasini, Lona”, 3 syntypes (NMW); “Tomor” / “Albania leg. Bischoff 1932”,
22 s. (MNHUB).
Notes. Schatzmayr (1937: 22) treated C. ravasinii as separate species, but hinted that it resembles
C. macedonicus. Based on the material studied, the present author considers the two taxa are only
well-defined subspecies of one species: Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat.
nov. for Calathus (Calathus) macedonicus Mařan, 1935. Müller (1935) published his description in
Bollettino della Società Entomologica Italiana, volume 67; this issue was printed on 26.II.1935.
On the other hand, Mařan (1935) described C. macedonicus in Sborník Entomologického Odděleni
Národního Musea v Praze, volume 13; this issue was printed on 1.XII.1935. The nominotypical
taxon is easily recognized from ssp. macedonicus by the anteriorly narrowed sides of pronotum.
96.7.2. Calathus ravasinii macedonicus Mařan, 1935
References. Calathus macedonicus: Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 525.
Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM);
“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); “Bjeshka Matroxh” /

170. COLEOPTERA: CARABIDAE: HARPALINAE 169
“Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 23 s.
(MNHUB); “Teja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalla Harushe” / “Korab”
/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
Subgenus Neocalathus Ball & Nègre, 1972
96.8. Calathus albanicus Apfelbeck, 1906
References. Calathus albanicus: Apfelbeck, 1906: 1666 (Merdita: Zepë Mt., loc. typ.); Apfelbeck,
1907a: 500, 503; Schatzmayr, 1937: 2, 8; 36 (Zepë Mt.); Csiki, 1940: 218 (Gjalicë Mt.; Korab
Mt.); Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 526.
Material studied. More than 160 s. with three kinds of labeling: “Albania Galica Lums” /
“Akad. Balk. Exp. Csiki 1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp.
Csiki 1918” (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); more
than 150 s. from the massifs of Shar Planina Mt., Merdita, Munella Mt., Korab Mt., Gjalica
Ljums and others (NMW); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 9 s. (MNHUB);
“Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff
1935”, 22 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Mal i
Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Fusha Korabit” / “Albania leg. Bischoff
1936”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 12 s. (MNHUB); “Berishe”
/ “Albania leg. Mueller 1939”, 2 s. (MNHUB); “Maja Shalles i Vogel-Ibale” / “Albania leg.
Mueller 1939”, 1 s. (MNHUB); “Kunora Drunit Iballe” / “Albania leg. Bischoff 1939”, 1 s.
(MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB).
Notes. This species inhabits the mountain chain Shar Planina Mt. - Korabi Mt. and massifs in
the area of Mirdita, but not the North Albanian Alps, where C. albanicus most probably is
replaced by C. metallicus.
96.9. Calathus ambiguus (Paykull, 1790)
96.9.1. Calathus ambiguus ambiguus (Paykull, 1790)
Material studied. “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”,
3 s. (MNHUB).
Notes. The mention of Hieke & Wrase (1988: 90) for Albania is apparently result of following
the record “Prisren” (Apfelbeck, 1904: 285). Currently this locality is situated in the Kosovo
Region of Serbia. Hovorka & Sciaky (2003b: 526) have most likely followed one of these
sources too. Since there is no previous reliable record from Albania, C. ambiguus is treated here
as new species for the fauna of the country.
96.10. Calathus cinctus Motschulsky, 1850 [= erythroderus Gemminger & Harold, 1868]
References. Calathus mollis f. erythroderus: Schatzmayr, 1943b: 111 (Lushnje). Calathus cinctus:
Aukema, 1990: 138.
Material studied. “Mal i Daitit” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Mal i Krujes”
/ “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”,
7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Mal i Daitit”

171. 170 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
/ “Albania 1936 leg. Ramadan coll. Bischoff ”, 4 s. (MNHUB); Kruja Castle, 16.XI.1997, 1 s.,
R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A.
Paparistos leg. (cAVT).
Notes. Hovorka & Sciaky (2003b: 527) have omitted C. cinctus for Albania. Here the species is
confirmed for the country.
96.11. Calathus erratus (C.R. Sahlberg, 1827)
96.11.1. Calathus erratus erratus (C.R. Sahlberg, 1827)
References. Calathus erratus: Hieke & Wrase, 1988: 90. Calathus erratus erratus: Hovorka &
Sciaky, 2003b: 527.
Material studied. “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Shkëlzën”
/ “Albania leg. Bischoff 1935”, 2 s. (MNHUB).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT).
Notes. First detailed records for Albania. In addition to the material studied above, the present
author has seen several further specimens in NMW, which remained unregistered.
96.12. Calathus melanocephalus (Linnaeus, 1758)
96.12.1. Calathus melanocephalus melanocephalus (Linnaeus, 1758)
References. Calathus melanocephalus: Csiki, 1940: 218 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.);
Hieke et al., 1986: 189 (Bizë near Shëngjergji; Lurë e Vjetër near Lura Mt.); Aukema, 1990:
138. Calathus melanocephalus melanocephalus: Hovorka & Sciaky, 2003b: 527.
Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); 30 s.
from localities “Shkëlzën”, “Bjeshka Matroxh”, “Tomor”, “Korab”, “Koritnik”, “Gjallica”,
“Mal i Shenjit”, “Boge”, “Cucale”, “Taja”, “Llogora”, “Cafa e Sherit Iballe”, “Çafa Markofçes”
(MNHUB); Tomor Mt., 21.IX.1961, 1 m., 1 f. (MNST).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,
900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. The mention of Hieke & Wrase (1988: 91) for Albania refers apparently to the record
“Prisren” (Apfelbeck, 1904: 286). Actually this locality is presently situated in the Kosovo
Region of Serbia. Besides the specimens above, I have seen several further specimens in
NMW, which remained unregistered.
96.13. Calathus metallicus Dejean, 1828
96.13.1. Calathus metallicus prokletijensis Mařan, 1938
References. Calathus metallicus: Hieke & Wrase, 1988: 90. Calathus metallicus metallicus: Hovorka
& Sciaky, 2003b: 527.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 15 s.
(NMW); nearly 100 s. from localities “Shkëlzën”, “Çafa Markofçes”, “Maja e Jezerze”
(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 1 f., P. Beron & B.
Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., P. Beron
& B. Petrov leg. (NMNHS).

172. COLEOPTERA: CARABIDAE: HARPALINAE 171
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
2250 m, 11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).
Notes. First detailed records for Albania. The absence of C. metallicus in the massifs of Shar
Planina and Korab Mt. may be a result of its replacement by C. albanicus there.
96.14. Calathus mollis (Marsham, 1802)
96.14.1. Calathus mollis mollis (Marsham, 1802)
References.Calathus mollis: Apfelbeck, 1904: 285 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë;
Berat); Hieke & Wrase, 1988: 91.
Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB).
Notes. Hovorka & Sciaky (2003b: 528) have omitted C. mollis for Albania. Here the species is
confirmed for the country. C. mollis is exclusively a littoral species and most probably the record
fromBerat(Apfelbeck,1904;1909)concernsC.cinctus(A.VignaTaglianti,personalcommunication).
97. Genus Synuchidius Apfelbeck, 1908
97.1. Synuchidius ganglbaueri Apfelbeck, 1908
References. Synuchidius ganglbaueri: Csiki, 1940: 218 (Gjalicë Mt.; Korab Mt.); Hieke, 1981: 82;
Hovorka & Sciaky, 2003b: 529.
Material studied. 66 s. from three kinds of labeling: “Albania Korab” / “Akad. Balk. Exp. Csiki
1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918” (HNHM);
“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); “Alban. Exped. Gjalica Ljums
17-26.VI.”, 12 s. (NMW); “Alban. Exped. Korab 1918 24-31..VII.”, 19 s. (NMW); “Jablanica coll.
Lichtneckert”, 2 s. (HNHM); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB);
“Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB).
Notes.Apfelbeckpublishedthreeconsecutivedescriptionsforthesubgenusandspecies(Apfelbeck,
1908; Apfelbeck, 1917; Apfelbeck, 1930), and the oldest ones are correctly treated as valid.
Subtribe SPHODRINA Laporte, 1834
98. Genus Laemostenus Bonelli, 1810
Subgenus Antisphodrus Schaufuss, 1865
98.1. Laemostenus cavicola (Schaum, 1858)
98.1.1. Laemostenus cavicola albanicus J. Müller, 1914
References. Laemostenus cavicola albanicus: Müller, 1914: 1005-1006 (Çukall Mt.: cave “Scpedha n’
kunor Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs n’ gur t’barzodhs”, loc. typ.); Pretner,
1978: 135, 148 (caves in Çukall Mt.; near Shkodër); Genest & Juberthie, 1994: 593 (Çukall Mt.:
cave “n’Kunor Vukai”, cave “Nre Gegs n’gur t’barzodhs”); Casale, 1988: 708, 716, 720 (Çukall
Mt.); Casale, 2003: 534. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 700-
1400 m). Laemosthenes cavicola: Jeannel, 1937b: 89; Magistretti, 1965: 439. Laemostenus cavicola modestus:
Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë).

173. 172 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Albanie”, 1 s. (MNHUB); “Albanien”, 1 s. (MNHUB); “Scpedha n’Kunor
Zukali Albanien Lona Giuno 1914”, 1 syntype (DEI); “Prokletija 1300-2000 m” / “Alban.
montenegr. Grenze Penther’14”, 12 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze
Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze
Penther’14”, 3 s. (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze Penther’14”, 2 s.
(NMW); “Albania Rapsa Dr. Grabowski”, 6 s. (HNHM); “Shpella Kodra Çarit” / “Cukale”, 3 s.
(MNHUB); “Brigie Hot” / “leg. Bischoff Albania 1934” / “Shpella e Lugstit”, 1 s. (MNHUB).
# Laemostenus cavicola modestus (Schaufuss, 1862)
References. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 700-1400 m).
Laemostenus cavicola modestus: Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë).
Notes.IfollowCasale(2003: 534)whonotedssp. modestus onlyforCroatiaandBosniaHerzegovina.
98.2. Laemostenus elongatus (Dejean, 1828)
98.2.1. Laemostenus elongatus pentheri J. Müller, 1924
References. Laemostenus elongatus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme,
300 m, and Vorfai Siperme, 700 m). Laemostenus elongatus pentheri: Müller, 1924: 70-71 (North
Albanian Alps: state attribution of loc. typ. in original description not specified; Rikavac;
Vermosh; Rapshë); Pretner, 1978: 136, 148, 168 (Rikavac; Vermosh; Rapshë); Casale, 1988:
700, 704 (Okol near Theth: designated as loc. typ.; Maranaj; Vermosh, Rapshë).
Material studied. 49 s. from localities “Maranaj”, “Shkëlzën”, “Maja e Jezercoë”, “Çafa Peis”,
“Cucale”, “Boge”, “Maja e Dragomirit”, “Maja Rogamit” (MNHUB); North Albanian Alps,
Bogë, 1400-1800 m, 1.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District,
Merkuth, 920-980 m, 11.VI.1933, 1 m., grassland, P. Beron & B. Petrov leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
Mali i Thatë Mt., 18.VII.1993, 1 f., L. Latella leg. (cAVT).
Notes. Casale (2003: 534) has omitted the species for Albania. Here it is confirmed for the country.
Subgenus Laemostenus s.str.
98.3. Laemostenus dalmatinus (Dejean, 1828)
Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW).
Notes. New species for the fauna of Albania.
98.4. Laemiostenus janthinus (Duftschmid, 1812)
98.4.1. Laemiostenus janthinus janthinus (Duftschmid, 1812)
References. Laemiostenus janthinus: Csiki, 1940: 218 (Gjalicë Mt.).
Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Notes. Casale (2003: 536) has omitted the species for Albania. Here it is confirms for the country.
98.5. Laemostenus venustus (Dejean, 1828)
References. Laemostenus venustus: Apfelbeck, 1904: 275 (Velipoja); Casale, 1988: 476 (Velipoja);
Hieke & Wrase, 1988: 93; Casale, 2003: 536.

174. COLEOPTERA: CARABIDAE: HARPALINAE 173
Material studied. “Mamurash” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mamurash”
/ “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Gjormi” / “Albania leg. Bischoff 1935”, 2 s.
(MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Sukthi bis Gurse”
/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB).
Subgenus Pristonychus Dejean, 1828
98.6. Laemostenus cimmerius (Fischer von Waldheim, 1823)
98.6.1. Laemostenus cimmerius cimmerius (Fischer von Waldheim, 1823)
References. Sphodrus cimmerius: Anonymous, 1907: 285. Prustonychus cimmerius: Magistretti, 1965:
433. Laemostenus cimmerius: Wrase, 1991: 11. Laemostenus cimmerius cimmerius: Casale, 1988: 809
(Durrës); Casale, 2003: 537.
Material studied. Durrës, amphitheater, 12.VI.1993, 1 f., B. Petrov leg. (NMNHS); Tiranë
Vore, 27.VI.1995, 1 m., S. Abadjiev leg. (NMNHS).
98.6.2. Laemostenus cimmerius weiratheri J. Müller, 1932
References. Laemostenus cimmerius weiratheri: Casale, 1988: 809 (M. Sopot); Sakalian & Guéorguiev,
1997: 48; Casale, 2003: 537.
98.7. Laemostenus terricola (Herbst, 1784)
98.7.1. Laemostenus terricola punctatus (Dejean, 1828)
References. Pristonychus terricola: Hieke et al., 1986: 189 (Mali me Gropë). Laemostenus punctatus:
Hieke & Wrase, 1988: 93. Laemostenus terricola punctatus: Casale, 1988: 787 (Mali me Grope,
Dolinen-Gez); Casale, 2003: 539.
Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);
“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tomor-Tojk juli
1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”,
3 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 8 s. (MNHUB); “Vukaj” / “Cucale” /
“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”,
2 s. (MNHUB); “Mal i That” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albanien – Exp.
DEI Mali me Gropë, Dolinengebiet 1350 m, 6.VII.1961”, 1 s. (DEI).
Subtribe SYNUCHINA Lindroth, 1956
99. Genus Synuchus Gyllenhal, 1810
Subgenus Synuchus s.str.
99.1. Synuchus vivalis (Illiger, 1798)
99.1.1. Synuchus vivalis vivalis (Illiger, 1798) [= nivalis (Panzer, 1796)]
References. Synuchus nivalis: Hieke & Wrase, 1988: 88. Synuchus vivalis vivalis: Hovorka & Sciaky,
2003c: 546.

175. 174 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Theth”
/ “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM).
Notes. First detailed records for Albania.
Tribe ZABRINI Bonelli, 1810
Subtribe AMARINA C. Zimmermann, 1831
100. Genus Amara Bonelli, 1810
Subgenus Amara s.str.
100.1. Amara aenea (DeGeer, 1774)
References. Amara aenea: Apfelbeck, 1904: 300 (Vlorë; Berat; Velipoja); Apfelbeck, 1907a:
497, 499 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m);
Apfelbeck, 1909: 297 (Vlorë); Csiki, 1940: 215 (Shkëlzen Mt.; Kula Ljums; Ura i Lopez; “Stičen”
near Gjalicë Mt.; Nangë; Gostilë); Schatzmayr, 1943b: 105 (Dukati; Tërpan); Hieke et al.,
1986: 189 (Tomor Mt.: Ali Abbas Monastery; North Albanian Alps near Theth); Hieke &
Wrase, 1988: 100; Hieke, 2003: 547.
Material studied. Tiranë, 7.IV.1960, 1 s. (MNST); Ibë, 18.V.1960, 1 s. (MNST); Rodeni, 2-
22.VII.1962, 4 s. (MNST); Zooteknica, 16.V.1963, 1 s. (MNST); Theth, 800-1200 m, 28.V.1993,
1 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, cave No 25, 23.VI.1993,
1 s., P. Beron, B. Petrov leg. (NMNHS); Saranda Dhermi, 2.V.1994, 1 s., P. Stoev leg. (NMNHS);
Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg. (NMNHS); 5 km Leskovic,
2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 2 s., S.
Golovach, B. Petrov, P. Stoev leg. (NMNHS); 5 km Saranda, 3.VI.1995, 1 s., G. Blagoev leg.
(NMNHS); Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Lyukove, 5.VI.1995, 1 s., V.
Sakalian leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 12-14.II.1998 / 16-21.II.1998 / 23-
28.II.1998, 56 s., R. Ward leg. (CMNH).
Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona, 900
m, 5.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995, 1 f., D. Antonelli
leg. (cAVT); Rapsh, 15.V.2002, 2 m., 1 f., A. Paparistos leg. (cAVT).
Notes. In addition to the material studied above, the present author has seen numerous further
specimens in HNHM, NMW and MNHUB, which remained unregistered.
100.2. Amara anthobia A. Villa & G.B. Villa, 1833
References. Amara anthobia: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke
& Wrase, 1988: 102; Hieke, 2003: 547.
Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Logara” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Cafa
e Dhembellit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Vlorë District, Llogora Pass,
1050 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS); Dhermi, 100 m, 11.V.1995, 1 s., S. Golovach,
B. Petrov, P. Stoev leg. (NMNHS).

176. COLEOPTERA: CARABIDAE: HARPALINAE 175
100.3. Amara convexior Stephens, 1828
References. Amara convexior: Hieke & Wrase, 1988: 97; Hieke, 2003: 548.
Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).
Notes. First detailed record for Albania.
100.4. Amara curta Dejean, 1828
References. Amara curta: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576 m;
Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 215 (Ura i Lopez; Gjalicë Mt.; Korab Mt.);
Hieke & Wrase, 1988: 98; Hieke, 2003: 548.
Material studied. “Apfelb. Alban. spt.. IV.V.’05”, 3 s. (NMW); “Prokletjia 1300-2000” / “Alban.
Montenegr. Grenze, Penther’14”, 2 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban.
Montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania,
Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Tartarit” /
“Albania leg. Bischoff 1932”, 5 s. (MNHUB); “Berat-Bargulash” / “leg. Bischoff Albania
1934”, 1 s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën”
/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 5 s.
(MNHUB); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron, B. Petrov
leg. (NMNHS); Shkoder, 12.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS).
100.5. Amara eurynota (Panzer, 1796)
References. Amara eurynota: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke
& Wrase, 1988: 101; Hieke, 2003: 549.
Material studied. “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bashtrik” /
“Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff
1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda
District, Qeparo, 4-5.V.1994, 1 s., P. Stoev leg. (NMNHS).
? Amara famelica C. Zimmermann, 1832
Material studied. “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 7 s. (NMW); “Alban. Exp.
1918 Korab, 23-31.VII.”, 1 s. (NMW).
Notes. F. Hieke (personal communication) expressed doubt about the correct identification
of the specimens in NMW.
100.6. Amara familiaris (Duftschmid, 1812)
References. Amara familiaris: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);
Hieke & Wrase, 1988: 101; Hieke, 2003: 549.
Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Logara”
/ “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 1 s.
(MNHUB); “Logara” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

177. 176 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
100.7. Amara lucida (Duftschmid, 1812)
References. Amara lucida: Apfelbeck, 1904: 301 (Berat); Apfelbeck, 1909: 297 (Berat); Csiki,
1940: 215 (Ura i Lopez; Korab Mt.); Hieke & Wrase, 1988: 102; Hieke, 2003: 549.
Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW);
“Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Ura i Lopez” /
“Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Krabe” / “Albania leg. Bischoff 1932”, 1
s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Shen Ili” /
“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor-Burany juli 1932”, 3 s. (MNHUB);
“Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “leg.
Bischoff Albania 1934”, 4 s. (MNHUB); “Mal i Shejit” / “leg. Bischoff Albania 1934”, 1
s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Logara” / “Albania
leg. Bischoff 1936”, 3 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1
s. (MNHUB).
100.8. Amara morio Ménétriés, 1832
100.8.1. Amara morio nivium Tschitschérine, 1900
References. Amara morio nivium: Hieke, 1995: 288 (Gjalicë Ljums); Hieke, 2003: 550.
Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM;
MNHUB).
100.9. Amara ovata (Fabricius, 1792)
References. Amara ovata: Hieke, 2003: 550.
Material studied. “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Llogora” / “leg.
Bischoff Albania 1936”, 2 s. (MNHUB).
Additional material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (MCG);
“Lona, Albania”, 1 s. (MSNM).
Notes. The mention of Hieke & Wrase (1988: 96) for Albania refers apparently to the record
“Prisren” (Apfelbeck, 1904: 299). However, this locality is situated in the Kosovo Region of
Serbia. Besides the material studied above, I have seen more specimens in NMW, which remained
unregistered. First detailed records for Albania.
100.10. Amara proxima Putzeys, 1866
References. Amara proxima: Hieke & Wrase, 1988: 98; Hieke, 2003: 551.
Material studied. “Shendeli Alban. Mer.” / “Winkler. Lona, Bischoff, V.1931”, 6 s. (NMW;
MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania
leg. Bischoff 1936”, 3 s. (MNHUB); “Maja e Tartarit juni 1937” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB).
Additional material included (studied F. Hieke). “Argiro Kastron, S.Alban”, 2 s. (MCG;
MTKD); “Ismen Ili, Albania”, 1 s. (NMPC).
Notes. First detailed records for Albania.

178. COLEOPTERA: CARABIDAE: HARPALINAE 177
100.11. Amara saphyrea Dejean, 1828
References. Amara saphyrea: Hieke & Wrase, 1988: 96; Hieke, 2003: 551.
Material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (NMW).
Notes. First detailed record for Albania.
100.12. Amara similata (Gyllenhal, 1810)
Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten
wood, R. Ward leg. (CMNH).
Notes. New species for the fauna of Albania.
# Amara spreta Dejean, 1831
References. Amara spreta: Csiki, 1940: 215 (Gjalicë Mt.).
Notes. Basing on the most recent information about the species distribution (Hieke, 2003:
551), the Csiki’s identification is most probably incorrect and that the material in question
concerns another congener. F. Hieke (personal communication) expressed similar opinion, so
A. spreta is removed from the list of the Albanian Carabidae.
100.13. Amara tibialis (Paykull, 1798)
References. Amara tibialis: Csiki, 1940: 215 (Korab Mt.); Hieke & Wrase, 1988: 103; Hieke,
2003: 551.
Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).
Subgenus Amarocelia Motschulsky, 1862
100.14. Amara erratica (Duftschmid, 1812)
References. Amara erratica: Csiki, 1940: 216 (Gjalicë Mt.); Hieke & Wrase, 1988: 103; Hieke,
2003: 556.
Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 1 s. (HNHM); “Prokletija 1300-2000
m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Skëlzën” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB).
Subgenus Bradytus Stephens, 1827
100.15. Amara apricaria (Paykull, 1790)
References. Amara apricaria: Hieke et al., 1986: 190 (Borshi near Vlorë; Bizë near Shëngjergji);
Hieke & Wrase, 1988: 106; Hieke, 2003: 553.
Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk.
Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp.’18 Pashtrik, 4-14.7.”, 3 s. (NMW); 28 s. from
localities “Shkalle Harushe-Korab”, “Korab”, “Shkëlzën”, “Mal i Sheniit”, “Mal i Tartarit”,
“Taja”, “Giallica” (MNHUB); “Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 2 s. (HNHM).

179. 178 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
100.16. Amara consularis (Duftschmid, 1812)
Material studied. Mal i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 s. (MNST).
Notes. New species for the fauna of Albania.
Subgenus Celia C. Zimmermann, 1832
100.17. Amara bifrons (Gyllenhal, 1810)
References. Amara bifrons: Csiki, 1940: 216 (Korab Mt.); Hieke & Wrase, 1988: 105; Hieke, 2003: 555.
Material studied. “Ricavac 1300 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1
s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW);
“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp. 1918
Korab 23-31.VII.”, 4 s. (NMW); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);
“Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercë” / “Albania leg.
Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB);
“Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM).
Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,
2250 m, 11.VII.1993, 4 f., L. Latella leg. (cAVT).
100.18. Amara hannemanni Hieke, 1991
References. Amara hannemanni: Hieke, 1991: 1-3 (Shkëlzen Mt., loc. typ.); Hieke, 2003: 556.
Material studied. “Skëlzën” / “Albania leg. Bischoff 1935”, 1 paratype (MNHUB); “Skëlzën”
/ “Albania leg. Bischoff 1936”, 6 paratypes (MNHUB).
100.19. Amara montana Dejean, 1828
References. Amara montana: Hieke, 2003: 557.
Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Shkoder” /
“25.V.1958 leg. Xh. Murraj”, 2 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS);
Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH).
Notes. First detailed records for Albania.
? Amara praetermissa (C.R. Sahlberg, 1827)
References. Amara praetermissa: Hieke & Wrase, 1988: 105.
Materialstudied.“Latif Koritnik”/“Coll.Apfelbeck”,2s.(HNHM);“Latif Koritnik”,3s.(NMW).
Notes. Hieke (2003: 557) has omitted the species for Albania in spite of his previous data
(Hieke & Wrase, 1988: 105). He stated me (personal communication) that the specimens above
might belong to A. hannemanni, so that A. praetermissa is treated here as doubtful for Albania.
Subenus Curtonotus Stephens, 1827
100.20. Amara aulica (Panzer, 1796)
References. Amara aulica: Hieke & Wrase, 1988: 109; Hieke, 2003: 559.

180. COLEOPTERA: CARABIDAE: HARPALINAE 179
Material studied. “Tomor Albanien”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff
1932”, 1 s. (MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
Subgenus Paracelia Bedel, 1899
100.21. Amara dalmatina Dejean, 1828
References. Amara dalmatina: Hieke (in press).
Material included (studied F. Hieke). “SE of Medua, Albania”, 1 s. (MNHUB); “Shendeli,
Albania mer.”, 1 s. (MNHUB).
Notes. First detailed records for Albania.
100.22. Amara quenseli (Schönherr, 1806)
100.22.1. Amara quenseli quenseli (Schönherr, 1806)
References. Amara quenseli: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576
m; Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 216 (Korab Mt.); Holdhaus & Lindroth,
1940: 146-147, 250 (Koritnik Mt.; Korab Mt.); Hieke & Wrase, 1988: 106. Amara quenseli quenseli:
Hieke, 2003: 564.
Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 2 s. (HNHM); “Latif Koritnik”, 8 s.
(NMW); “Merdita Munela” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Prokletija 1300-
2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Prokletija 1300-2000 m” /
“Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Galica Lums” / “Akad. Balk.
Exp. Csiki 1917”, 2 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki
1918”, 3 s. (HNHM); “Korab” / “Albania leg. Bischoff 1935”, 16 s. (MNHUB); “Korab” /
“Albania leg. Bischoff 1936”, 19 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 23
s. (MNHUB).
Subgenus Parapercosia Tschitschérine, 1899
100.23. Amara taurica (Motschulsky, 1844) [= fallaciosa (J. Müller, 1946)]
References. Zabrus fallaciosus: Müller, 1946: 107-110 (South Albania: Shëndelli Mt. near Tepelenë,
loc. typ.); Turin, 1981: 148. Amara taurica: Hieke, 2003: 565.
Notes. Hieke (1995: 35-36) synonymized Zabrus fallaciosus Müller, 1946 with Amara taurica,
noting the type locality of the former as “Patria falsa!”, but later the same has cited A. taurica
for Albania (Hieke, 2003: 565). According to F. Hieke (personal communication) A. taurica is
known from different parts of Turkey and from two localities in Greece, so that the type
locality of Zabrus fallaciosus in Albania should not be considered as doubtful (Hieke, 2005).
Subgenus Percosia C. Zimmermann, 1832
100.24. Amara equestris (Duftschmid, 1812)
100.24.1. Amara equestris equestris (Duftschmid, 1812) [= tomoricensis J. Müller, 1923]

181. 180 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
References. Amara tomoricensis: Müller, 1923: 108-109 (Tomor Mt.: Kulmak, loc. typ.). Amara
equestris: Csiki, 1940: 216 (Korab Mt.); Hieke, 1978: 229, 257 (Tomor Mt.: Kulmak; Koritnik
Mt.; Korab Mt.); Hieke & Wrase, 1988: 108. Amara equestris equestris: Hieke, 2003: 565.
Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);
“Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Korab” / “Albania leg. Bischoff
1935”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Albania
Zalle-Dardha” / “24.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM).
Subgenus Xenocelia Hieke, 2001
100.25. Amara bischoffi Jedlička, 1946
References. Amara municipalis: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.); Jedlička, 1946: 7
Peshtrik Mt. (loc. typ.); Hieke & Wrase, 1988: 104. Amara bischoffi: Turin, 1981: 145; Hieke,
1983: 328-329, 334 (Peshtrik Mt.; Taja); Hieke, 2001: 33 (two localities). Amara municipalis
bischoffi: Hieke, 2003: 557.
Material studied. “Taja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Bashtrik” / “Albania
leg. Bischoff 1936”, 1 syntype (MNHUB).
Notes. According to F. Hieke (personal communication) the records of Csiki (1940) are
problematic since they may concern A. bischoffi, as well as A. messae.
100.26. Amara cursitans C. Zimmermann, 1832 [= properans C. Zimmermann, 1832]
References. Amara properans: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.). Amara cursitans: Hieke
& Wrase, 1988: 104; Hieke, 2001: 50 (five localities); Hieke, 2003: 556.
Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s.
(NMW); “Skëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg.
Bischoff 1936”, 5 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);
“Maja e Dragomit” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB).
100.27. Amara fusca Dejean, 1828 [= complanata Dejean, 1828]
References. Amara complanata: Csiki, 1940: 215 (Gjalicë Mt.). Amara fusca: Hieke & Wrase,
1988: 104; Hieke, 2001: 42 (“Koritza-Ebene”, 1 s. in MNHN); Hieke, 2003: 556.
100.28. Amara messae Baliani, 1924
References. Amara messae: Hieke & Wrase, 1988: 104; Hieke, 2001: 65; Hieke, 2003: 557.
Material studied. “Giallica” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë”
/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cotype” / “Bashtrik” / “Albania leg. Bischoff
1936”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron, B.
Petrov leg. (NMNHS).
Notes. First detailed records for Albania.

182. COLEOPTERA: CARABIDAE: HARPALINAE 181
Subgenus Zezea Csiki, 1929
100.29. Amara chaudoiri Schaum, 1858
100.29.1. Amara chaudoiri incognita Fassati, 1946
References. Amara chaudoiri incognita: Hieke, 1970: 138 (Skrofotino near Vlorë); Hieke, 2003:
567. Amara chaudoiri: Hieke & Wrase, 1988: 94.
# Amara erythrocnema Dejean, 1828
References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina).
Notes. At present the species is treated as local endemic for the island of Corfu (Hieke, 2003:
567). F. Hieke (personal communication) expressed the opinion that the record for A.
erythrocnema from Albania most probably concerns A. reflexicollis. Therefore, the former species
is removed from the list of Albanian Carabidae.
100.30. Amara fulvipes (Audinet-Serville, 1821)
References. Amara fulvipes: Apfelbeck, 1904: 298 (Velipoja); Hieke, 1970: 157 (Velipoja; Lum;
Tiranës; Lum i Tiranës; Mal i Dajtit Mt.); Hieke & Wrase, 1988: 94; Hieke, 2003: 567.
Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane”
/ “leg. Bischoff Albania 1934”, 6 s. (MNHUB).
100.31. Amara reflexicollis Motschulsky, 1844
100.31.1. Amara reflexicollis reflexicollis Motschulsky, 1844
References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina). Amara reflexicollis
reflexicollis: Hieke, 2003: 568.
100.32. Amara tricuspidata Dejean, 1831
References. Amara tricuspidata: Hieke & Wrase, 1988: 94; Hieke, 2003: 568.
Material studied. “Museum Paris Albanie Starova (env. de Koritza) Caporal Vuillaume 1918”
/ “Albanie Armée d’Orient 1916-1918” / “Avril” / “Mai”, 1 s. (MNHUB).
Additional material included (studied F. Hieke). 1 s. labelled as that up above (MNHP).
Notes. First detailed record for Albania.
Subtribe ZABRINA Bonelli, 1810
101. Genus Zabrus Clairville, 1806
Subgenus Pelor Bonelli, 1810
Notes. The species from this subgenus are arranged here in three groups of species with the
aim to delineate the related lineages.

183. 182 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Species group “aetolus”
101.1. Zabrus aetolus Schaum, 1864
[Zabrus aetolus kodymi Mařan, 1940]
Notes. This race has been described from the Montenegrin part of the North Albanian Alps
(Mařan, 1940a: 33), basing on the label data of the material collected by O. Kodym and C.
Purkynì in 1937. The ssp. kodymi may inhabit the Albanian part of the mountain, too.
101.1.1. Zabrus aetolus winkleri J. Müller, 1946
References. Zabrus winkleri: Müller, 1946: 110-111 (Maja e Tartarit Mt., south of Vlorë, loc.
typ.; Maja e Djembellit Mt., east of Gjirokastër; Maja e Policanit Mt., east of Gjirokastër);
Turin, 1981: 148. Zabrus aetolus winkleri: Freude, 1989: 83, 95 (Maja e Tartarit Mt.; Maja e
Dhembellit Mt.; Maja e Policanit Mt.); Serrano & Andújar, 2003: 570.
Material studied. “Argirokastron Sud-Albania”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania
leg. Bischoff 1932”, 10 s. (MNHUB).
101.2. Zabrus albanicus Apfelbeck, 1904
101.2.1. Zabrus albanicus albanicus Apfelbeck, 1904
References. Zabrus albanicus latifianus: Ganglbauer, 1931: 39 (Koritnik Mt.); Mařan, 1939d:
145-146 (Koritnik Mt.); Schatzmayr, 1943b: 108 (Koritnik Mt.); Freude, 1989: 83, 88, 97
(Koritnik Mt.); Drovenik & Peks, 1994: 96 (Koritnik Mt.); Drovenik & Peks, 1999: 111 (Koritnik
Mt.). Zabrus albanicus: Csiki, 1940: 221 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Schatzmayr,
1943b: 108 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke, 1981: 83. Zabrus albanicus albanicus:
Serrano & Andújar, 2003: 570.
Material studied. “Latif Koritnik” / “albanicus typ. Apfelb.”, 12 s. (NMW); “Latif Alban. spt.”,
3 s. (DEI); more than 140 s. from localities “Albania Galica Lums” (or “Gyalica Lums”), “Skölsen”
and “Korab” (HNHM); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4
s. (NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 2 s. (NMW); “Alban. Exped. 1918 Gjalica
Ljums 17-26.VI.”, 20 s. (NMW); “Alban. Exped. Korab 1918 24-31.VII.”, 71 s. (NMW).
Notes. The populations from the Koritnik Mt., as well as the records for these (Ganglbauer,
1931; Mařan, 1939d; Schatzmayr, 1943b; Freude, 1989; Drovenik & Peks, 1994; 1999), are
assigned provisionally to the nominotypical subspecies rather than to the ssp. latifianus, since
the Zepë Mt. (an unmarked, but potential “loc. typ.” of the latter taxon) has an more southern
and well-isolated position from the other massifs in North Albania. Besides the material studied
above, in MNHUB the present author has seen other more 140 s. of Z. albanicus s.l. from
different localities, which remained unregistered.
[Zabrus albanicus jablanicensis Mařan, 1939]
References. Zabrus albanicus jablanicensis: Freude, 1989: 88 (“Mittel-Ost-Albanien an der jugo-
slavischen Grenze”).
Notes. Serrano & Andújar (2003: 570) indicated Bosnia Herzegovina as region of the
distribution of ssp. jablanicensis by mistake. In fact, this race was described from the Jablanitsa
Mt. shared between Albania and Macedonia and so far is cited only from the Macedonian part
of the mountain. Most likely the record of Freude (1989) is not based on material coming

184. COLEOPTERA: CARABIDAE: HARPALINAE 183
from collections, and may bases of supposition. The subspecies most probably inhabits the
Albanian part of the Jablanitsa Mt., too.
101.2.2. Zabrus albanicus latifianus Ganglbauer, 1915
References. Zabrus albanicus: Apfelbeck, 1907a: 503 (Zepë Mt.). Zabrus albanicus latifianus:
Ganglbauer, 1931: 39 (Zepë Mt.); Mařan, 1939d: 145-146 (Zepë Mt.); Schatzmayr, 1943b: 108
(Zepë Mt.); Freude, 1989: 83, 88, 97 (Zepë Mt.); Serrano & Andújar, 2003: 570.
Material studied. “Merdita Munela”, 1 s. (NMW); “Merdita Zebia”, 2 s. (NMW).
Notes. In my opinion, only the populations from the region of Merdita have to be considered
as belonging to spp. latifianus.
! Zabrus albanicus ssp.
Zabrus albanicus: Schatzmayr, 1943b: 108 (Tomor Mt.: Abbas Ali and Toik).
Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s. (NMW;
MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 8 s. (MNHUB);
“Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e Papingut”, 21 s. (MNHUB).
Notes. The specimens from the massifs of Tomor Mt., Maja e Dhembelit Mt. and Maja e
Papingut Mt. (all in South Albania) may concern another subspecies, since the latter are far
removed in the south from the massifs of the Zepë Mt. and Koritnik Mt.
# Zabrus apfelbecki Ganglbauer, 1915
References. Zabrus apfelbecki: Magistretti, 1965: 360.
Notes. Magistretti (1965) referred older records of Z. ignavus Csiki, 1907 to Z. apfelbecki,
but I failed to find any records in the literature. The latter species lives only in Bosnia
Herzegovina (Serrano & Andújar, 2003: 570), so that it is removed from the list of the
Albanian ground beetles.
101.3. Zabrus ganglbaueri Apfelbeck, 1906
101.3.1. Zabrus ganglbaueri ganglbaueri Apfelbeck, 1906
References. Zabrus ganglbaueri: Apfelbeck, 1906: 1667 (Merdita: Mal i Shenjt Mt. near Oroshi,
loc. typ.); Apfelbeck, 1907a: 499-500, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1936:
108 (Mal i Shenjt Mt. near Oroshi); Schatzmayr, 1943b: 108 (Mal i Shenjt Mt. near Oroshi);
Hieke, 1981: 75. Zabrus ganglbaueri ganglbaueri: Freude, 1989: 82, 110 (Mal i Shenjt Mt. near
Oroshi; Mal i That Mt. near Gjirokastër); Serrano & Andújar, 2003: 570.
Material studied. “Merdita M. Scheit”, 1 s. (NMW); “Merdita 18.19. n. 20/5 1934”, 2 s. (NMW).
Notes. Freude (1989) stated that the populations from the Mal i Shenjt Mt. (Merdita District)
and the Mal i That Mt. (Gjirokastër District) belong to the nominotypical race. In fact both
populations inhabit opposite outskirts of the country, so the specimens from the latter massif
may belong either to the following subspecies or to a separate geographical race.
101.3.2. Zabrus ganglbaueri lonae J. Müller, 1923
References. Zabrus lonae: Müller, 1923: 107-108 (Tomor Mt.: Toik, loc. typ.); Müller, 1935:
154 (Tomor Mt.: Toik); Mařan, 1940a: 30-31, 33, 35. Zabrus lonai: Schatzmayr, 1943b: 108
(Tomor Mt.: Toik). Zabrus ganglbaueri lonae: Freude, 1989: 83, 111 (Tomor Mt.: Toik); Serrano
& Andújar, 2003: 570.

185. 184 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Material studied. “Albania 1922 Ravasini, Lona” / “Tomor Monte Toik”, 1 s. (DEI); “Tomor:
Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Tojk” / “Albania
leg. Bischoff 1932”, 9 s. (MNHUB).
Species group “incrassatus”
101.4. Zabrus bischoffi J. Müller, 1936
References. Zabrus bischoffi: Müller, 1936: 107-108 (Mal i Dajtit Mt. near Tiranë, loc. typ.;
Llogora); Schatzmayr, 1943b: 106, 109 (Kruja; Mal i Dajtit Mt.; Llogora); Turin, 1981: 148.
Zabrus incrassatus bischoffi: Freude, 1989: 86, 120 (Mal i Dajtit Mt. near Tiranë; Llogara, south of
Vlorë); Serrano & Andújar, 2003: 571.
Material studied. 19 s. from “Mal i Dajtit” (MNHUB); 5 s. from “Mal i Krues” (MNHUB);
3 s. from “Mal i Shenjit” (MNHUB); 37 s. from “Llogora” (MNHUB); Llogora Pass, 1025 m,
6.6.1995, 1 m., G. Blagoev leg. (NMNHS).
Notes. This species has a quite different apex of the median lobe and sides of pronotum
narrower anteriorly, which is in contrast with the respective characteristics in Z. incrassatus s.l.
In my opinion, the former is separate species not subspecies of the latter: Zabrus (Pelor) bischoffi
J. Müller, 1936, stat. rest. of Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936.
101.5. Zabrus boldorii Schatzmayr, 1943
References. Zabrus boldori: Schatzmayr, 1943: 106, 108 (Dukati, south of Vlorë, loc. typ.);
Serrano & Andújar, 2003: 570. Zabrus baldori sic!: Turin, 1981: 148. Zabrus incrassatus boldorii:
Freude, 1989: 86, 119 (Dukati near Vlorë).
Material studied. “Südalbanien Kopstein” / “coll. Prof. F. Werner” / “Pašaliman”, 1 m.
(NMW); “Südalbanien Kopstein” / “Valona”, 2 f. (NMW).
101.6. Zabrus incrassatus (Ahrens, 1814)
101.6.1. Zabrus incrassatus incrassatus (Ahrens, 1814)
References. Zabrus incrassatus: Apfelbeck, 1904: 310 (Berat; Vlorë); Anonymous, 1907: 285;
Apfelbeck, 1907a: 495-496 (northeast of Shkodër; Maranajt Mt.: between Vorfai Postme, 300
m, and Vorfai Siperme, 700 m); Apfelbeck, 1909: 297 (Vlorë; Berat); Müller, 1936: 108 (Shëndelli
Mt.; Tepelenë; Maja e Tartarit Mt.; Tiranë; Gjirokastër; Berat; Vlorë); Schatzmayr, 1943b: 106,
108 (“Poggio Boschetto”; Lushnje; Berat; Vlorë; Shen Ili; Tepelenë; Maja e Tartarit Mt.; Tiranë;
Shkodër); Hieke & Wrase, 1988: 110.
Material studied. “Südalbanien Kopstein”, 2 s. (NMW); “Albanien Ochrida”, 1 s. (NMW);
“Hani Grabom Mai” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Rapsa 10-
18.V.” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Alban. Exp. 1918 Bicaj
14.15.VI.”, 3 s. (NMW); Ibë, 2-11.VII. 1962, 1 m. (MNST); Shkodër, 15.V.1963, 3 m., 1 f.
(MNST); 2 km N Durrës, 24.V.1993, 1 m., P. Stoev leg. (NMNHS); Vlorë, 1.V.1994, 2 m., 3 f.,
P. Stoev leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 f., P. Stoev leg. (NMNHS);
Saranda District, Borsh, 5.V.1994, 1 f., P. Stoev leg. (NMNHS); Saranda District, Ljukova, 3-
4.VI.1995, 1 f., G. Blagoev leg. (NMNHS); Vlorë District, Llogora Mt., 1027 m, 4.VI.1995, 1
f., G. Blagoev leg. (NMNHS).

186. COLEOPTERA: CARABIDAE: HARPALINAE 185
Notes. Serrano & Andújar (2003: 571) have omitted the species for Albania, so it is here
confirmed for the country. Besides the material studied above, in MNHUB I have seen other
more than 100 s. from different localities (“Shen Ili”, “Tirane”, “Murthi”, “Tepelene”, “Munela”,
“Burelle”, “Vlorë”, “Mal i Gjer”, “Myzege”, “Bose”, “Maja e Dhembellit”, “Tarabosh”, “Mal
i Tartarit”, etc.).
Species group “spinipes”
101.7. Zabrus spinipes (Fabricius, 1798)
101.7.1. Zabrus spinipes insignis J. Müller, 1932
References. Zabrus spinipes: Burmeister, 1939: 127; Hieke et al., 1986: 190 (Tomor Mt.: Abbas
Ali Monastery); Hieke & Wrase, 1988: 110; Freude, 1989: 147.
Material studied. “Shen-Ili Juni” / “Albania leg. Bischoff 1932”, 11 s. (MNHUB); “Mal i
That” / “Albania leg. Bischoff 1937”, 22 s. (MNHUB).
Notes. Serrano & Andújar (2003: 572) have omitted the species for Albania, so it is here
confirmed for the country.
Subgenus Zabrus s.str.
101.8. Zabrus tenebrioides (Goeze, 1777)
101.8.1. Zabrus tenebrioides tenebrioides (Goeze, 1777)
References. Zabrus tenebrioides: Schatzmayr, 1943b: 105-106 (Dukati; Lushnje). Zabrus tenebrioides
tenebrioides: Serrano & Andújar, 2003: 573.
Material studied. “Myzeqe” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); 11 km west of
Tiranë, 20.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 5 s., B.
Guéorguiev leg. (NMNHS).
Additional material included (studied A. Vigna Taglianti). Durrës, Currila, 8.V.2002, 1 f., A.
Paparistos leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of Vijosë River, 300-
360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).
Tribe ZUPHIINI Bonelli, 1810
Subtribe ZUPHIINA Bonelli, 1810
102. Genus Polystichus Bonelli, 1810
102.1. Polystichus connexus (Geoffroy, 1785)
References. Polystichus connexus: Hieke & Wrase, 1988: 163.
Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).
Notes. Baehr (2003e: 574) has omitted the species for Albania. Here it is confirmed for the
country as first detailed record is given.

187. 186 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
103. Genus Zuphium Latreille, 1806
Subgenus Zuphium s.str.
103.1. Zuphium olens (P. Rossi, 1790)
103.1.1. Zuphium olens olens (P. Rossi, 1790)
References. Zuphium olens: Schatzmayr, 1943b: 112 (Stan-Karbunarë); Hieke & Wrase, 1988:
162. Zuphium olens olens: Baehr, 2003e: 574.
Material studied. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

188. Results
When omitting doubtful species and taxa still not well defined, the Albanian Carabidae
listed in this catalogue represent 543 species. They are distributed among 11 subfamilies
(Nebriinae – 25, Cicindelinae – 10, Carabinae – 20, Elaphrinae – 1, Omophroninae –
1, Brachininae – 17, Siagoninae – 1, Scaritinae – 28, Apotominae – 2, Trechinae – 115,
and Harpalinae – 323), 31 tribes and 103 genera (Table 1).
One tribe, 10 genera, 11 subgenera and 55 species are new for the fauna of Albania,
and 1 subgenus and 3 species are new for the fauna of the Balkan Peninsula (Appendix
A.1-2). From these, 1 tribe, 1 subtribe, 7 genera, 20 subgenera and 72 species are
reported firstly with detailed record/s for the country (Appendix A.3). Altogether
119 species, omitted in the recent Palaearctic Catalogue (Löbl & Smetana, 2003), are
here confirmed for Albania (Appendix A.4).
Twelve taxa from the species group remain still not well defined, as likely most of
them represent new for the science species or subspecies (Appendix A.5). Two
subgenera and 22 species and subspecies are doubtful for the country (Appendix
A.6); generally the published records about them concern adjacent regions in
Montenegro, the Kosovo Region of Serbia, Macedonia or Northwestern Greece. Most
of the species and subspecies from the last category may inhabit Albania, but until
now we do not have reliable data to confirm this. Two subgenera and 29 species and
subspecies are deleted from the list (Appendix A.7). The systematic list includes also
some endemics described from massifs shared between Albania and neighbour
countries (but not recorded yet from the former), as well as several more widespread
species found from localities very close to Albania.
A moderate taxonomic contribution is also achieved (Appendix B). Principally, it is
based on study of types, topotypes or more extensive series of previously problematic
species and subspecies from Albania and neighbouring countries. In result of that 38
various taxonomic innovations (new synonyms, new status or revalidations) are
introduced. Further, nine taxa from the species group (5 valid species and 4 synonyms),
all described by V. Apfelbeck, have been found to have assigned an incorrect year of
description. Accordingly, the relevant earlier years of their descriptions are fixed. Finally,
eight nomina nuda were also traced out, as one of them may be a result of mislabeling,
and one is supplied with year of description in the literature.
RESULTS 187

189. 188 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Conclusions
Undoubtedly the Balkan Peninsula contains the richest ground beetle diversity
compared to other diverse European regions like the Iberian Peninsula, Italy or the
Caucasus Major (Turin, 1981; Kryzhanovskij et al., 1995; Serrano, 2003; Vigna Taglianti,
1993; 2004; 2005a; 2005b). Currently an impressive total of at least 1487 species of
Carabidae, following the classification used in Löbl & Smetana, 2003, is reported
from this territory (unpublished data). This figure is provisional while the systematics,
diversity and distribution of the Carabidae in the Balkan Peninsula are better known.
Until now, the carabid fauna of Bulgaria is probably the best investigated one in the
region (Hieke & Wrase, 1988; Guéorguiev & Guéorguiev, 1995). The species
composition of no other Balkan regions has been examined in details; only that of
ex-Yugoslavia was subject of uncritical registration (Drovenik & Peks, 1994; Drovenik
& Peks, 1999). Therefore, the Carabidae of Slovenia, Croatia, Bosnia Herzegovina,
Montenegro, Serbia, Macedonia, Romania, Greece and European Turkey need scrutiny
and writing of up-to-date checklists.
Bearing in mind the surface of Albania, the number of the carabid species and
subspecies there seems relatively high (543), if compared it with that of other Balkan
or Mediterranean regions. In Bulgaria there are known more than 720 species
(Guéorguiev & Guéorguiev, 1995), from Macedonia there are reported about 420
species (Guéorguiev, 1998; Hristovski, personal communication), and for ex-Yugoslavia
more than 1500 species and subspecies have been cited (Drovenik & Peks, 1999).
Both the Iberian Peninsula and Italy, having territories from 10 to 20 times bigger
than Albania, have a number of species only 2.13-2.42 times higher than is that in the
last country (Serrano, 2003; Vigna Taglianti, 2005a; 2005b).

190. Subfamily Tribe Subtribe Genus Nr. Subgenus Nr.
species species
Nebriinae Nebriini Eurynebria 1
Leistus 7 Leistus 2
Pogonophorus 5
Nebria 9 Alpaeus 5
Boreonebria 1
Eunebria 1
Nebria 2
Notiophilini Notiophilus 8
Cicindelinae Cicindelini Calomera 2
Cicindela 4 Cicindela 4
Cylindera 3 Cylindera 1
Eugrapha 2
Myriochila 1 Myriochila 1
Carabinae Carabini Calosoma 4 Calosoma 2
Microcallisthenes 2
Carabus 15 Archicarabus 1
Carabus 1
Chaetocarabus 1
Megodontus 3
Oreocarabus 3
Pachystus 2
Procerus 1
Procrustes 1
Tachypus 1
Tomocarabus 1
Cychrini Cychrus 1
Elaphrinae Elaphrini Elaphrus 1 Neoelaphrus 1
Omophroninae Omophronini Omophron 1 Omophron 1
Brachininae Brachinini Aptinus 3 Aptinus 3
Brachinus 14 Brachinus 5
Brachynidius 4
Brachynolomus 1
Cnecostolus 3
insertae sedis 1
Siagoninae Siagonini Siagona 1
Scaritinae Clivinini Clivinina Clivina 3
Reicheiina Reicheadella 1 Reicheadella 1
Dyschiriini Dyschirius 21 Chiridysus 1
Dyschiriodes 14
Dyschirius 1
Eudyschirius 3
Paradyschirius 2
Scaritini Scaritina Scarites 3 Parallelomorphus 2
Scallophorites 1
Apotominae Apotomini Apotomus 2
Trechinae Bembidiini Bembidiina Asaphidion 4
Bembidion 63 Bembidion 1
Taxonomic structure of Albanian Carabidae (Table 1)
TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE 189

191. 190 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Bembidionetolitzkya 9
Chlorodium 1
Diplocampa 2
Emphanes 5
Euperyphus 3
Metallina 2
Microserrullula 1
Nepha 3
Notaphemphanes 1
Notaphus 1
Ocydromus 3
Ocyturanes 6
Odontium 2
Omoperyphus 1
Peryphanes 4
Peryphus 4
Philochthus 6
Phyla 1
Princidium 1
Talanes 1
Terminophanes 1
Testedium 1
Trepanes 3
Ocys 1
Sinechostictus 3 Pseudolimnaeum 1
Sinechostictus 2
Tachyina Lymnastis 1
Paratachys 3
Polyderis 1
Sphaerotachys 1
Tachys 1
Tachyta 1 Tachyta 1
Tachyura 2 Tachyura 2
Patrobini Deltomerus 2 Deltomerus 1
Paradeltomerus 1
Pogonini Pogonistes 2
Pogonus 2 Pogonus 2
Trechini Perileptina Perileptus 1 Perileptus 1
Trechina Albanotrechus 1
Duvalius 10 Duvalius 9
Euduvalius 1
Neotrechus 3
Trechus 12 Trechus 12
Trechodina Thalassophilus 1
Harpalinae Chlaeniini Callistina Callistus 1
Chlaeniina Chlaenius 13 Chlaeniellus 6
Chlaenites 1
Chlaenius 1
Dinodes 2
Epomis 2
Trichochlaenius 1
Cyclosomini Masoreina Masoreus 1 Masoreus 1
Dryptini Drypta 1 Drypta 1
Subfamily Tribe Subtribe Genus Nr. Subgenus Nr.
species species

192. Harpalini Amblystomina Amblystomus 3
Anisodactylina Anisodactylus 4 Anisodactylus 3
Pseudodichirius 1
Diachromus 1
Gynandromorphus 1
Ditomina Carterus 3 Carterus 3
Ditomus 2
Dixus 3
Graniger 1
Odontocarus 1
Harpalina Acinopus 4 Acinopus 2
Oedematicus 1
Osimus 1
Harpalus 32 Cryptophonus 1
Harpalus 29
Pseudophonus 2
Ophonus 20 Brachyophonus 1
Hesperophonus 4
Incisophonus 1
Metophonus 10
Ophonus 4
Parophonus 6 Ophonomimus 1
Parophonus 5
Trichotichnus 1 Trichotichnus 1
Stenolophina Acupalpus 10 Acupalpus 10
Anthracus 2
Bradycellus 4 Bradycellus 4
Dicheirotrichus 1 Dicheirotrichus 1
Loxoncus 1
Stenolophus 7 Egadroma 1
Stenolophus 6
Lebiini Cymindidina Cymindis 9 Cymindis 7
Menas 1
Tarulus 1
Dromiina Dromius 1 Dromius 1
Microlestes 7
Paradromius 1 Manodromius 1
Philorhizus 2 Philorhizus 2
Lebiina Lebia 5 Lamprias 1
Lebia 4
Lionychina Apristus 2
Lionychus 2 Lionychus 2
Syntomus 4
Licinini Licinina Badister 4 Badister 2
Baudia 2
Licinus 4 Licinus 3
Neorescius 1
Odacanthini Odacantha 1 Odacantha 1
Oodini Oodes 2 Oodes 2
Omphreini Omphreus 3 Omphreus 3
Panagaeini Panagaeus 1 Panadaeus 1
Platynini Agonum 15 Agonum 13
Europhilus 2
Subfamily Tribe Subtribe Genus Nr. Subgenus Nr.
species species
TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE 191

193. 192 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Anchomenus 1 Anchomenus 1
Olisthopus 2
Paranchus 1
Platynus 2 Batenus 1
Platynus 1
Pterostichini Abax 2 Abacopercus 1
Abax 1
Molops 15 Molops 14
Stenochoromus 1
Myas 1 Myas 1
Pedius 2
Poecilus 8 Angoleus 1
Poecilus 7
Pterostichus 24 Argutor 1
Bothriopterus 1
Cheporus 1
Feronidius 1
Melanius 2
Morphnosoma 1
Parahaptoderus 1
Phonias 3
Platypterinus 1
Platysma 1
Pseudomaseus 4
Pterostichus 7
Speluncarius 2 Hypogium 1
Speluncarius 1
Stomis 2 Stomis 2
Tapinopterus 6 Tapinopterus 6
Sphodrini Atranopsina Platyderus 4 Platyderus 4
Calathina Calathus 14 Bedelinus 1
Calathus 6
Neocalathus 7
Synuchidius 1
Sphodrina Laemostenus 7 Antisphodrus 2
Laemostenus 3
Pristonychus 2
Synuchina Synuchus 1 Synuchus 1
Zabrini Amarina Amara 32 Amara 13
Amarocelia 1
Bradytus 2
Celia 3
Curtonotus 1
Paracelia 2
Parapercosia 1
Percosia 1
Xenocelia 4
Zezea 4
Zabrina Zabrus 8 Pelor 7
Zabrus 1
Zuphiini Polystichus 1
Zuphium 1 Zuphium 1
11 subfamilies 31 tribes 103 genera 543 species
Subfamily Tribe Subtribe Genus Nr. Subgenus Nr.
species species

194. APPENDIX 193
Appendix
A. Faunistic contributions
A.1. New taxa for the fauna of the Balkan Peninsula
Subgenera
1. Terminophanes Müller-Motzfeld, 1998
Species
1. Bembidion (Terminophanes) terminale Heer, 1841
2. Cymindis (Cymindis) angularis Gyllenhal, 1810
3. Lionychus (Lionychus) maritimus Fairmaire, 1862
A.2. Taxa new for the fauna of Albania
Tribes
1. ODACANTHINI Laporte, 1834
Genera
1. Eurynebria Ganglbauer, 1891
2. Myriochila Motschulsky, 1858
3. Lymnastis Motschulsky, 1862
4. Polyderis Motschulsky, 1862
5. Pogonistes Chaudoir, 1871
6. Odontocarus Solier, 1835
7. Dromius Bonelli, 1810
8. Lionychus Wissmann, 1846
9. Odacantha Paykull, 1798
10. Paranchus Lindroth, 1974
Subgenera
1. Myriochila Motschulsky, 1858 s.str.
2. Dyschirius Bonelli, 1810 s.str.
3. Talanes Motschulsky, 1864
4. Terminophanes Müller-Motzfld, 1998
5. Pseudolimnaeum Kraatz, 1888
6. Menas Motschulsky, 1864
7. Dromius Bonelli, 1810 s.str.
8. Lamprias Bonelli, 1810
9. Lionychus Wissmann, 1846 s.str.
10. Odacanta Paykull, 1798 s.str.
11. Parahaptoderus Jeanne, 1969
Species (subspecies not considered)
1. Eurynebria complanata (Linnaeus, 1767)
2. Leistus (Pogonophorus) apfelbecki Ganglbauer, 1891

195. 194 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
3. Nebria (Alpaeus) germarii Heer, 1837
4. Myriochila (Myriochila) melancholica (Fabricius, 1798)
5. Aptinus (Aptinus) acutangulus Chaudoir, 1876
6. Dyschirius (Dyschiriodes) luticola Chaudoir, 1850
7. Dyschirius (Dyschirius) latipennis Seidlitz, 1867
8. Scarites (Parallelomorphus) terricola Bonelli, 1813
9. Asaphidion stierlini (Heyden, 1880)
10. Bembidion (Bembidion) quadripustulatum Audinet-Serville, 1821
11. Bembidion (Euperyphus) testaceum (Duftschmid, 1812)
12. Bembidion (Peryphanes) deletum Audinet-Serville, 1821
13. Bembidion (Peryphus) femoratum Sturm, 1825
14. Bembidion (Philochthus) mannerhaimii C.R. Sahlberg, 1827
15. Bembidion (Talanes) subfasciatum Chaudoir, 1850
16. Bembidion (Terminophanes) terminale Heer, 1841
17. Bembidion (Trepanes) maculatum Dejean, 1831
18. Sinechostictus (Pseudolimnaeum) doderoi (Ganglbauer, 1891)
19. Sinechostictus (Sinechostictus) millerianus (Heyden, 1883)
20. Lymnastis galilaeus Piochard de la Brûlerie, 1876
21. Polyderis cardioderus (Chaudoir, 1850)
22. Pogonistes convexicollis Chaudoir, 1871
23. Pogonistes gracilis (Dejean, 1828)
24. Carterus (Carterus) rotundicollis Rambur, 1837
25. Odontocarus robustus (Dejean, 1831)
26. Harpalus (Harpalus) euchlorus Ménétriés, 1839
27. Harpalus (Harpalus) flavescens (Piller et Mitterpacher, 1783)
28. Harpalus (Harpalus) hospes Sturm, 1818
29. Ophonus (Hesperophonus) similis (Dejean, 1829)
30. Ophonus (Metophonus) melletii (Heer, 1837)
31. Ophonus (Ophonus) stictus Stephens, 1828
32. Acupalpus (Acupalpus) brunnipes (Sturm, 1825)
33. Stenolophus (Stenolophus) abdominalis Gené, 1836
34. Cymindis (Cymindis) angularis Gyllenhal, 1810
35. Cymindis (Cymindis) coadunata Dejean, 1825
36. Cymindis (Menas) miliaris (Fabricius, 1801)
37. Dromius (Dromius) quadrimaculatus (Linnaeus, 1758)
38. Lebia (Lamprias) cyanocephala (Linnaeus, 1758)
39. Lebia (Lebia) trimaculata (Villers, 1789)
40. Apristus reticulatus Schaum, 1857
41. Lionychus (Lionychus) maritimus Fairmaire, 1862
42. Lionychus (Lionychus) quadrillum (Duftschmid, 1812)
43. Badister (Badister) bullatus (Schrank, 1798)
44. Odacantha (Odacantha) melanura (Linnaeus, 1767)
45. Agonum (Agonum) afrum (Duftschmid, 1812)
46. Agonum (Agonum) muelleri (Herbst, 1784)
47. Paranchus albipes (Fabricius, 1796)
48. Poecilus (Poecilus) versicolor (Sturm, 1824)

196. APPENDIX 195
49. Pterostichus (Parahaptoderus) brevis (Duftschmid, 1812)
50. Pterostichus (Pseudomaseus) nigrita (Paykull, 1790)
51. Calathus (Calathus) ellipticus Reitter, 1889
52. Calathus (Neocalathus) ambiguus (Paykull, 1790)
53. Laemostenus (Laemostenus) dalmatinus (Dejean, 1828)
54. Amara (Amara) similata (Gyllenhal, 1810)
55. Amara (Bradytus) consularis (Duftschmid, 1812)
A.3. Taxa firstly reported with detailed record/s for Albania
Tribes
1. CYCLOSOMINI Laporte, 1834
Subtribes
1. MASOREINA Chaudoir, 1870
Genera
1. Tachys Dejean, 1821
2. Masoreus Dejean, 1821
3. Diachromus Erichson, 1837
4. Paradromius Fowler, 1887
5. Myas Sturm, 1826
6. Synuchus Gyllenhal, 1810
7. Polystichus Bonelli, 1810
Subgenera
1. Boreonebria Jeannel, 1937
2. Eunebria Jeannel, 1937
3. Procerus Dejean, 1821
4. Notaphemphanes Netolitzky, 1920
5. Omoperyphus Netolitzky, 1931
6. Phyla Motschulsky, 1844
7. Masoreus Dejean, 1821 s.str.
8. Pseudodichirius Lutshnik, 1921
9. Brachyophonus Sciaky, 1987
10. Incisophonus Sciaky, 1987
11. Tarulus Bedel, 1906
12. Manodromius Reitter, 1905
13. Batenus Motschulsky, 1865
14. Myas Sturm, 1826 s.str.
15. Bothriopterus Chaudoir, 1835
16. Morphnosoma Lutshnik, 1915
17. Platypterinus Reitter, 1908
18. Bedelinus Ragusa, 1885
19. Synuchus Gyllenhal, 1810 s.str.
20. Curtonotus Stephens, 1827
Species (subspecies not considered)
1. Leistus (Leistus) nitidus (Duftschmid, 1812)
2. Leistus (Pogonophorus) parvicollis Chaudoir, 1869

197. 196 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
3. Nebria (Boreonebria) rufescens (Strøm, 1768)
4. Nebria (Eunebria) jockischi Sturm, 1815
5. Notiophilus aquaticus (Linnaeus, 1758)
6. Notiophilus germinyi Fauvel, 1863
7. Calosoma (Calosoma) inquisitor (Linnaeus, 1758)
8. Carabus (Procerus) gigas Creutzer, 1799
9. Clivina collaris (Herbst, 1784)
10. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
11. Bembidion (Emphanes) axillare (Motschulsky, 1844)
12. Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
13. Bembidion (Odontium) foraminosum Sturm, 1825
14. Bembidion (Omoperyphus) hypocrita Dejean, 1831
15. Bembidion (Philochthus) guttula (Fabricius, 1792)
16. Bembidion (Phyla) tethys Netolitzky, 1926
17. Paratachys micros (Fischer von Waldheim, 1828)
18. Tachys scutellaris Stephens, 1828
19. Tachyura (Tachyura) parvula (Dejean, 1831)
20. Chlaenius (Chlaeniellus) nitidulus (Schrank, 1781)
21. Chlaenius (Chlaeniellus) tristis (Schaller, 1783)
22. Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829
23. Chlaenius (Epomis) circumscriptus (Duftschmid, 1812)
24. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813)
25. Anisodactylus (Pseudodichirius) intermedius Dejean, 1829
26. Diachromus germanus (Linnaeus, 1758)
27. Harpalus (Harpalus) laevipes Zetterstedt, 1828
28. Harpalus (Harplaus) pumilus Sturm, 1818
29. Harpalus (Harpalus) subcylindricus Dejean, 1829
30. Harpalus (Harpalus) latus (Linnaeus, 1758)
31. Harpalus (Harpalus) solitaris Dejean, 1829
32. Harpalus (Harpalus) autumnalis (Duftschmid, 1812)
33. Ophonus (Brachyophonus) krueperi Apfelbeck, 1902
34. Ophonus (Incisophonus) incisus (Dejean, 1829)
35. Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
36. Ophonus (Metophonus) schaubergerianus (Puel, 1937)
37. Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)
38. Acupalpus (Acupalpus) elegans (Dejean, 1829)
39. Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)
40. Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
41. Acupalpus (Acupalpus) suturalis Dejean, 1829
42. Bradycellus (Bradycellus) distinctus (Dejean, 1829)
43. Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
44. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758)
45. Paradromius (Manodromius) linearis (Olivier, 1795)
46. Philorhizus (Philorhizus) notatus (Stephens, 1827)
47. Lebia (Lebia) humeralis Dejean, 1825
48. Lebia (Lebia) scapularis (Geoffroy, 1785)

198. APPENDIX 197
49. Licinus (Licinus) cassideus (Fabricius, 1792)
50. Agonum (Agonum) marginatum (Linnaeus, 1758)
51. Platynus (Batenus) scrobiculatus (Fabricius, 1801)
52. Abax (Abax) ovalis (Duftschmid, 1812)
53. Myas (Myas) chalybaeus (Palliardi, 1825)
54. Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
55. Pterostichus (Morphnosoma) melanarius (Illiger, 1798)
56. Pterostichus (Phonias) diligens (Sturm, 1824)
57. Pterostichus (Platypterinus) ottomanus Apfelbeck, 1908
58. Calathus (Bedelinus) circumseptus Germar, 1824
59. Calathus (Calathus) bosnicus Ganglbauer, 1891
60. Calathus (Neocalathus) erratus (C.R. Sahlberg, 1827)
61. Calathus (Neocalathus) metallicus Dejean, 1828
62. Synuchus (Synuchus) vivalis (Illiger, 1798)
63. Amara (Amara) convexior Stephens, 1828
64. Amara (Amara) ovata (Fabricius, 1792)
65. Amara (Amara) proxima Putzeys, 1866
66. Amara (Amara) saphyrea Dejean, 1828
67. Amara (Celia) montana Dejean, 1828
68. Amara (Curtonotus) aulica (Panzer, 1796)
69. Amara (Paracelia) dalmatina Dejean, 1828
70. Amara (Xenocelia) messae Baliani, 1924
71. Amara (Zezea) tricuspidata Dejean, 1831
72. Polystichus connexus (Geoffroy, 1785)
A.4. Species omitted for Albania in Löbl & Smetana (2003); here their presence in the
country is confirmed
1. Leistus (Leistus) fulvibarbis Dejean, 1826
2. Leistus (Pogonophorus) magnicollis Motschulsky, 1866
3. Nebria (Eunebria) jockischi Sturm, 1815
4. Notiophilus aestuans Dejean, 1826
5. Notiophilus aquaticus (Linnaeus, 1758)
6. Notiophilus biguttatus (Fabricius, 1779)
7. Notiophilus danieli Reitter, 1897
8. Notiophilus germinyi Fauvel, 1863
9. Notiophilus palustris (Duftschmid, 1812)
10. Notiophilus rufipes Curtis, 1829
11. Notiophilus substriatus C.R. Waterhouse, 1833
12. Calomera fischeri (M.F. Adams, 1817)
13. Carabus (Oreocarabus) preslii Dejean, 1830
14. Elaphrus (Neoelaphrus) uliginosus Fabricius, 1792
15. Brachinus (Brachinus) elegans Chaudoir, 1842
16. Brachinus (Brachinus) plagiatus Reiche, 1868
17. Brachinus (Cnecostolus) bayardi Dejean, 1831
18. Dyschirius (Dyschiriodes) nitidus (Dejean, 1825)

199. 198 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
19. Dyschirius (Dyschiriodes) cylindricus (Dejean, 1825)
20. Dyschirius (Dyschiriodes) laeviusculus Putzeys, 1846
21. Dyschirius (Eudyschirius) gracilis (Heer, 1837)
22. Scarites (Scallophorites) buparius (Forster, 1771)
23. Asaphidion flavipes (Linnaeus, 1761)
24. Bembidion (Bembidionetolitzkya) conforme (Dejean, 1831)
25. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
26. Bembidion (Bembidionetolitzkya) rhodopense Apfelbeck, 1902
27. Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
28. Bembidion (Bembidionetolitzkya) varicolor (Fabricius, 1803)
29. Bembidion (Chlorodium) splendidum Sturm, 1825
30. Bembidion (Diplocampa) assimile Gyllenhal, 1810
31. Bembidion (Emphanes) axillare (Motschulsky, 1844)
32. Bembidion (Emphanes) latiplaga Chaudoir, 1850
33. Bembidion (Emphanes) tenellum Erichson, 1837
34. Bembidion (Euperyphus) scapulare Dejean, 1831
35. Bembidion (Metallina) properans (Stephens, 1828)
36. Bembidion (Microserrullula) quadricolle (Motschulsky, 1844)
37. Bembidion (Nepha) callosum Küster, 1847
38. Bembidion (Nepha) caucasicum (Motschulsky, 1844)
39. Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
40. Bembidion (Ocyturanes) pindicum Apfelbeck, 1901
41. Bembidion (Odontium) foraminosum Sturm, 1825
42. Bembidion (Odontium) striatum (Fabricius, 1792)
43. Bembidion (Omoperyphus) hypocrita Dejean, 1831
44. Bembidion (Peryphanes) brunnicorne Dejean, 1831
45. Bembidion (Peryphanes) grandipenne Schaum, 1862
46. Bembidion (Peryphanes) tetracolum Say, 1823
47. Bembidion (Philochthus) guttula (Fabricius, 1792)
48. Bembidion (Philochthus) inoptatum Schaum, 1857
49. Bembidion (Philochthus) lunulatum (Geoffroy, 1785)
50. Bembidion (Phyla) tethys Netolitzky, 1926
51. Bembidion (Princidium) punctulatum Drapiez, 1820
52. Bembidion (Testedium) bipunctatum (Linnaeus, 1761)
53. Bembidion (Trepanes) octomaculatum (Goeze, 1777)
54. Ocys harpaloides (Audinet-Serville, 1821)
55. Sinechostictus (Sinechostictus) tarsicus (Peyron, 1858)
56. Paratachys bistriatus (Duftschmid, 1812)
57. Paratachys fulvicollis (Dejean, 1831)
58. Paratachys micros (Fischer von Waldheim, 1828)
59. Sphaerotachys haemorrhoidalis (Ponza, 1805)
60. Tachys scutellaris Stephens, 1828
61. Tachyta (Tachyta) nana (Gyllenhal, 1810)
62. Tachyura (Tachyura) diabrachys (Kolenati, 1845)
63. Tachyura (Tachyura) parvula (Dejean, 1831)
64. Pogonus (Pogonus) littoralis (Duftschmid, 1812)

200. APPENDIX 199
65. Chlaenius (Chlaeniellus) terminatus Dejean, 1826
66. Chlaenius (Epomis) dejeani (Dejean, 1831)
67. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813)
68. Harpalus (Harpalus) cupreus Dejean, 1829
69. Harpalus (Pseudophonus) griseus (Panzer, 1796)
70. Ophonus (Hesperophonus) subquadratus (Dejean, 1829)
71. Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
72. Acupalpus (Acupalpus) parvulus (Sturm, 1825)
73. Bradycellus (Bradycellus) caucasicus (Chaudoir, 1846)
74. Cymindis (Cymindis) axillaris (Fabricius, 1794)
75. Cymindis (Cymindis) humeralis (Geoffroy, 1785)
76. Cymindis (Cymindis) lineola L. Dufour, 1820
77. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758)
78. Microlestes luctuosus Holdhaus, 1904
79. Microlestes minutulus (Goeze, 1777)
80. Microlestes negrita (Wollaston, 1854)
81. Paradromius (Manodromius) linearis (Olivier, 1795)
82. Philorhizus (Philorhizus) notatus (Stephens, 1827)
83. Lebia (Lebia) cruxminor (Linnaeus, 1758)
84. Lebia (Lebia) humeralis Dejean, 1825
85. Lebia (Lebia) scapularis (Geoffroy, 1785)
86. Apristus subaeneus Chaudoir, 1846
87. Syntomus impressus (Dejean, 1825)
88. Syntomus pallipes (Dejean, 1825)
89. Syntomus truncatellus (Linnaeus, 1761)
90. Omphreus (Omphreus) morio Dejean, 1828
91. Agonum (Agonum) extensum Ménétriés, 1849
92. Agonum (Agonum) hypocrita (Apfelbeck, 1904)
93. Agonum (Agonum) marginatum (Linnaeus, 1758)
94. Agonum (Agonum) sexpunctatum (Linnaeus, 1758)
95. Agonum (Agonum) sordidum Dejean, 1828
96. Agonum (Agonum) viridicupreum (Goeze, 1777)
97. Agonum (Europhilus) antennarium (Duftschmid, 1812)
98. Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
99. Olisthopus fuscatus Dejean, 1828
100. Olisthopus glabricollis (Germar, 1817)
101. Platynus (Batenus) scrobiculatus (Fabricius, 1801)
102. Platynus (Platynus) assimilis (Paykull, 1790)
103. Molops (Molops) alpestris (Dejean, 1828)
104. Molops (Molops) apfelbecki Ganglbauer, 1891 (considering former Molops reseri Apfelbeck, 1904)
105. Molops (Molops) curtulus Ganglbauer, 1891
106. Molops (Molops) holdhausi J. Müller, 1917
107. Molops (Molops) plurisetosus J. Müller, 1918
108. Molops (Molops) piceus (Panzer, 1793)
109. Pterostichus (Cheporus) burmeisteri Heer, 1838
110. Pterostichus (Platysma) niger (Schaller, 1783)

201. 200 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
111. Pterostichus (Pterostichus) reiseri Ganglbauer, 1889
112. Platyderus (Platyderus) dalmatinus L. Miller, 1881
113. Calathus (Neocalathus) cinctus Motschulsky, 1850
114. Calathus (Neocalathus) mollis (Marsham, 1802)
115. Laemostenus (Antisphodrus) elongatus (Dejean, 1828)
116. Laemiostenus (Laemostenus) janthinus (Duftschmid, 1812)
117. Zabrus (Pelor) incrassatus (Ahrens, 1814)
118. Zabrus (Pelor) spinipes (Fabricius, 1798)
119. Polystichus connexus (Geoffroy, 1785)
A.5. Taxa still not well defined
1. Nebria (Alpaeus) germarii ssp.
2. Dalmatoreicheia sp.
3. Reicheadella (Reicheadella) sp.
4. Bembidion (Bembidionetolitzkya) sp. (aff. complanatum Heer, 1937)
5. Bembidion (Bembidionetolitzkya) sp. (ex. gruppo “fasciolatum”)
6. Bembidion (Terminophanes) terminale ssp.
7. Adriaphaenops sp.
8. Trechini gen. & sp. indet. (? aff. Pheggomisetes Knirsch, 1923)
9. Omphreus (Omphreus) ravasinii ssp.
10. Tapinopterus (Tapinopterus) protensus ssp.1
11. Tapinopterus (Tapinopterus) protensus ssp.2
12. Zabrus (Pelor) albanicus ssp.
A.6. Taxa (cited, mapped or found in collections) with doubtful presence in Albania
Subgenera
1. Platycarabus A. Morawitz, 1886
2. Trachycarabus Géhin, 1876
Species and subspecies
1. Nebria (Alpaeus) bosnica tresnjevikensis Horvatovich, 1974
2. Nebria (Alpaeus) dahli (Duftschmid, 1812) (considering ssp. montenegrina Apfelbeck, 1904)
3. Carabus (Platycarabus) irregularis Fabricius, 1792 (considering ssp. ramanus Sokolář, 1909)
4. Carabus (Trachycarabus) scabriusculus Olivier, 1795 (considering ssp. bulgarus Lapouge, 1908)
5. Siagona europaea oberleitneri Dejean, 1830
6. Dyschirius (Eudyschirius) gracilis (Heer, 1837) (considering nominotypical subspecies)
7. Asaphidion cyanicorne (Pandellé, 1867) (considering nominotypical subspecies)
8. Bembidion (Bembidion) humerale Sturm, 1825
9. Bembidion (Bembidion) quadrimaculatum (Linnaeus, 1761) (considering nominotypical subspecies)
10. Bembidion (Bembidionetolitzkya) relictum Apfelbeck, 1904
11. Deltomerus (Deltomerus) malissorum Apfelbeck, 1918
12. Deltomerus (Paradeltomerus) paradoxus paradoxus Apfelbeck, 1908
13. Pogonus (Pogonus) luridipennis (Germar, 1823)
14. Trechus (Trechus) irenis Csiki, 1912
15. Cymindis (Cymindis) pindicola Apfelbeck, 1901

202. APPENDIX 201
16. Olisthopus rotundatus (Paykull, 1790) (considering nominotypical subspecies)
17. Molops (Molops) spartanus parnassicola Kraatz, 1875
18. Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908
19. Pterostichus (?) epiroticus Csiki, 1930
20. Calathus (Calathus) glabricollis Dejean, 1828
21. Amara (Amara) famelica C. Zimmermann, 1832
22. Amara (Celia) praetermissa (C.R. Sahlberg, 1827)
A.7. Taxa deleted from the list of Albanian Carabidae
Subgenera
1. Eucarabus Géhin, 1876
2. Oreoplatysma Jakobson, 1907
Species and subspecies
1. Nebria (Nebria) salina Fairmaire & Laboulbène, 1854
2. Cicindela (Cicindela) hybrida Linnaeus, 1758
3. Carabus (Eucarabus) catenulatus Scopoli, 1763
4. Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
5. Cychrus semigranosus semigranosus Palliardi, 1825
6. Cychrus semigranosus balcanicus Hopffgarten, 1881
7. Cychrus semigranosus montanus Winkler, 1939
8. Brachinus (Brachinus) alexandri F. Battoni, 1984
9. Dyschirius (Dyschiriodes) punctatus (Dejean, 1825)
10. Bembidion (Bembidionetolitzkya) ascendens K. Daniel, 1902
11. Bembidion (Bembidionetolitzkya) fasciolatum (Duftschmid, 1812)
12. Bembidion (Peryphanes) milleri Jacquelin du Val, 1852
13. Tachyura (Tachyura) sexstriata (Duftschmid, 1812)
14. Duvalius (Bicharotrechus) maglicensis Winkler, 1933
15. Trechus (Trechus) obtusiusculus Ganglbauer, 1889
16. Acinopus (Acinopus) subquadratus Brullé, 1832
17. Harpalus (Harpalus) anatolicus Tschitschérine, 1898
18. Harpalus (Harpalus) danieli Reitter, 1900
19. Molops (Molops) weiratheri J. Müller, 1930
20. Molops (Molops) simplex simplex Chaudoir, 1868
21. Molops (Molops) rufipes klisuranus Apfelbeck, 1902
22. Pterostichus (Oreoplatysma) lodosi Heinz, 1977
23. Tapinopterus (Tapinopterus) comita Jedlička, 1935
24. Tapinopterus (Tapinopterus) extensus (Dejean, 1831)
25. Tapinopterus (Tapinopterus) miridita jacupicensis Jedlička, 1935
26. Laemostenus (Antisphodrus) cavicola modestus (Schaufuss, 1862)
27. Amara (Amara) spreta Dejean, 1831
28. Amara (Zezea) erythrocnema Dejean, 1828
29. Zabrus (Pelor) apfelbecki Ganglbauer, 1915

203. 202 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
B. Taxonomic contributions
B.1. Nomenclatorial acts
Nebriini
Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. for Nebria (Alpaeus) aetolica macedonica Mařan, 1938
Nebria (Alpaeus) macedonica galicica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)
aetolica galicica Horvatovich, 1974a
Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)
aetolica papingutica Horvatovich, 1974a
Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assignment of Nebria (Alpaeus) aetolica
rambouseki Mařan, 1938
Carabini
Carabus (Oreocarabus) neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri
Schaum, 1856
Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii Dejean, 1830
Cychrini
Cychrus semigranosus albanicus Hopp, 1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck,
1904
Cychrus semigranosus dinaricus Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck,
1904
Licinini
Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn. nov. of Licinus (Licinus) graecus Apfelbeck, 1901
Platynini
Platynus (Batenus) scrobiculatus serbicus Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904
Platynus (Batenus) scrobiculatus turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus
Apfelbeck, 1904
Pterostichini
Molops (Molops) maderi J. Müller, 1933, syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917
Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri
reiseri Apfelbeck, 1904
Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri
corallipes J. Müller, 1936
Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri
pseudoalpestris Mlynář, 1977
Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri
victoris J. Müller, 1918
Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Müller, 1917
Aphaon Heyden, 1906, syn. nov. of Pterostichus s. str. Bonelli, 1810
Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment of Pterostichus (Aphaon)
bischoffianus Jedlička, 1936

204. APPENDIX 203
Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment of Pterostichus (Aphaon) brucki
Schaum, 1859
Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment of Pterostichus
(Aphaon) latifianus latifianus Apfelbeck, 1906
Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment of Pterostichus (Aphaon)
latifianus strupii J. Müller, 1935
Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment of Pterostichus
(Aphaon) lumensis lumensis Apfelbeck, 1905
Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment of Pterostichus (Aphaon) lumensis
litae Csiki, 1940
Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment of Pterostichus
(Aphaon) lumensis ljubetensis Apfelbeck, 1908
Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment of Pterostichus (Aphaon)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. for Pterostichus (Pterostichus)
malissorum Apfelbeck, 1905
Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment of Pterostichus (Aphaon) meisteri
Reitter, 1885
Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment of Pterostichus (Aphaon)
merklii (J. Frivaldszky, 1879)
Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment of Pterostichus (Aphaon) pentheri
Apfelbeck, 1918
Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment of Pterostichus (Aphaon) reiseri
Ganglbauer, 1889
Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment of Pterostichus
(Aphaon) rhilensis rhilensis Rottenberg, 1874
Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment of Pterostichus (Aphaon)
rhilensis kourili Mařan, 1933
Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment of Pterostichus (Aphaon)
rhilensis vitosensis Mařan, 1933
Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment of Pterostichus (Aphaon) walteri
Reitter, 1883
Tapinopterus (Tapinopterus) extensoides (Straneo, 1936), stat. rest. for Tapinopterus (Tapinopterus)
extensus extensoides (Straneo, 1936).
Sphodrini
Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat. nov. for Calathus (Calathus) macedonicus
Mařan, 1935
Zabrini
Zabrus (Pelor) bischoffi J. Müller, 1936, stat. rest. for Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936.
B.2. Year of description
The following taxa (valid or synonyms) have long time been accredited to have a correct year
of description. However, it was found out that this is incorrect. Below, the relevant earlier

205. 204 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
years of publication dates of the original descriptions are given. All the descriptions considered
fulfil the conditions of the ICZN (1999).
Carabus croaticus ljubetensis Apfelbeck, 1917 (synonym of Carabus croaticus durmitorensis Apfelbeck,
1904) instead Carabus croaticus ljubetensis Apfelbeck, 1918a
Carabus violaceus merditanus Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)
instead Carabus violaceus merditanus Apfelbeck, 1918a
Carabus violaceus shardagensis Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)
instead Carabus violaceus shardagensis Apfelbeck, 1918a
Aptinus merditanus Apfelbeck, 1917 (valid species) instead Aptinus merditanus Apfelbeck, 1918b
Aptinus merditanus var. granulipennis Apfelbeck, 1917 (synonym of Aptinus merditanus Apfelbeck,
1917) instead Aptinus merditanus var. granulipennis Apfelbeck, 1918b
Trechus albanicus Apfelbeck, 1905 (valid species) instead Trechus albanicus Apfelbeck, 1907b
Ophonus krueperi Apfelbeck, 1902 (valid species) instead Ophonus krueperi (Apfelbeck, 1904)
Licinus graecus Apfelbeck, 1901 (valid species) instead Licinus cassideus graecus Apfelbeck, 1904
Molops malshentianus Apfelbeck, 1917 (valid species) instead Molops malshentianus Apfelbeck, 1918b
B.3. Nomina nuda found in both collections and literature
Eurynebria complanata orientalis Apfelbeck
Eurynebria complanata albanica Apfelbeck
Eurynebria complanata var. albanica Apfelbeck
? Nebria ganglbaueri ditana Apfelbeck (or incorrect spelling of Nebria ganglbaueri merditana
Apfelbeck, 1906)
Aptinus lugubris rufipes Apfelbeck (according to Apfelbeck, 1907a: 499, 501)
Bembidion merditanum Apfelbeck
Harpalus bischoffi Schauberger
Molops transfuga Apfelbeck (according to Müller, 1930: 124; Mlynář, 1977: 120)
Molops apfelbecki bischoffi J. Müller
Pterostichus djumanus Apfelbeck

206. REFERENCES 205
References
General work on the names, the systematics and the distribution of the Carabidae
from the Balkan Peninsula (the separate parts of the first work are cited in the following
section)
1. Löbl I., A. Smetana. 2003. Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata -
Myxophaga - Adephaga. Stenstrup: Apollo Books, 819 pp.
2. Lorenz W. 1998. Nomina Carabidarum - a directory of the scientific names of ground
beetles (Insecta, Coleoptera “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae,
Cicindelinae, Rhysodinae). Tutzing: W. Lorenz, iv + 937 pp.
Faunistic and taxonomic references dealing with Albanian Carabidae (sources containing
data but not found are in brackets)
1. Absolon K., J. Mařan. 1947. Příspěvek k poznání geografického rozšíření, systematiku a
vzniku druhů rodu Neotrechus J. Müll. – Příroda, 39: 1-6.
2. Anonymous. 1907. Collections du Musée d’Histoire Naturelle de son altesse Royale
Ferdinand I Prince de Bulgarie. Sofia, xiv + 484 p.
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u godini 1900. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 13: 405-438.
4. Apfelbeck V. 1901b. Zwei neue Rassen der Gattung Carabus aus Albanien. – Societas
Entomologica, 16: 121-122.
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zoogeographische Beiträge. – Münchener Koleopterologische Zeitschrift, 1 [1902-1903]: 95-101.
6. Apfelbeck V. 1903. Ophonus suturifer Reitt. Ist = O. fallax Peyr. = suturalis Chaud. – Wiener
Entomologische Zeitung, 22: 53-54.
7. Apfelbeck V. 1904. Die Käferfauna der Balkanhalbinsel, mit Berücksichtigung Klein-
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9. Apfelbeck V.1906.NeueKoleopteren, gesammeltwährendeinerinJahre1905mitSubvention
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in Wien, Mathematisch-naturwissenschaftliche Klasse (Abt.I), 116: 493-506.

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11. Apfelbeck V. 1907b. Neue Koleopteren von der Balkanhalbinsel. – Wissenschaftliche
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1900. prigodom arheoloskog putovanja. – Glasnik Zemaljskog Museja u Bosni i
Hercegovini, 21: 297-301.
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Nomenclature, London, online version: http://www.iczn.org/iczn.
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Carabidi per la valutazione ambientale e la conservazione della biodiversità. Manuale
operativo. APAT, Manuali e Linee Guida, 34: 240 pp.

225. 224 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
Index
(valid taxa are given in bold)
Abacopercus 138
Abax 138
Abax s.str. 139
abdominalis (Stenolophus) 119
abdominalis persicus (Stenolophus) 119
Acinopus 97
Acinopus s.str. 97
Acupalpus 12, 115
Acupalpus s.str. 115
acutangulus (Aptinus) 39
Adriaphaenops 79
aenea (Amara) 174
aeneocephalus (Chlaenius) 92
aeneocephalus aeneocephalus (Chlaenius) 92
aeneus (Dyschirius) 46
aeneus (Harpalus) 107
aeneus aeneus (Dyschirius) 46
aestuans (Notiophilus) 23
aetolica (Nebria) 18
aetolica galicica (Nebria) 18, 21
aetolica korabica (Nebria) 21
aetolica macedonica (Nebria) 18
aetolica papingutica (Nebria) 18, 21, 22
aetolica rambouseki (Nebria) 18
aetolica tomoricensis (Nebria) 21, 22
aetolicum (Bembidion) 63
aetolicum aetolicum (Bembidion) 63
aetolicus (Carabus) 30
aetolicus (Tapinopterus) 163
aetolicus peristericus (Tapinopterus) 162
aetolicus tomoricensis (Tapinopterus) 163
aetolus (Zabrus) 182
aetolus kodymi (Zabrus) 182
aetolus winkleri (Zabrus) 182
affinis (Harpalus) 107
affinis ab. limbopunctatus (Harpalus) 107
affinis ab. viridulus (Harpalus) 107
afrum (Agonum) 133
agnatus (Dyschirius) 46
Agonum 133
Agonum s.str. 133
albanensis (Carabus) 31
albanensis (Harpalus) 111
albanensis (Ophonus) 111
albanicola (Carabus) 34
albanicum (Bembidion) 68
albanicum (Hypogeobium) 160
albanicum (Platysma) 160
albanicus (Calathus) 169, 171
albanicus (Carabus) 35
albanicus (Cychrus) 37
albanicus (Harpalus) 103
albanicus (Molops) 142
albanicus (Omphreus) 132
albanicus (Ophonus) 111
albanicus (Pterostichus) 160
albanicus (Speluncarius) 160
albanicus (Tapinopterus) 160
albanicus (Trechus) 87
albanicus (Zabrus) 182, 183
albanicus albanicus (Zabrus) 182
albanicus latifianus (Zabrus) 182, 183
albanicus jablanicensis (Zabrus) 182
Albanotrechus 79
albipes (Paranchus) 137
alexandri (Brachinus) 40
alinae (Molops) 140
Alpaeus 18
alpestre (Agonum) 133
alpestris (Molops) 141
alpestris imitator (Molops) 141
Amara 12, 174
Amara s.str. 174
AMARINA 174
Amarocelia 177
ambiguus (Calathus) 169
ambiguus (Chlaenius) 91
ambiguus ambiguus (Calathus) 169

226. INDEX 225
AMBLYSTOMINA 93
Amblystomus 93
ammophilus (Acinopus) 98
anatolicus (Harpalus) 104
anatolicus anatolicus (Harpalus) 104
Ancholeus 149
Anchomenus 136
Anchomenus s.str. 136
andreae (Bembidion) 68
andreae albanicum (Bembidion) 68
andreae bualei (Bembidion) 68
angularis (Cymindis) 121
angustatum (Agonum) 134
ANISODACTYLINA 94
Anisodactylus 12, 94
Anisodactylus s.str. 94
antennarium (Agonum) 136
antennarius (Platynus) 136
anthobia (Amara) 174
anthracinus (Pterostichus) 154
anthracinus anthracinus (Pterostichus) 154
anthracinus biimpressus (Pterostichus) 155
Anthracus 117
Antisphodrus 171
anxius (Harpalus) 102
apfelbecki (Leistus) 17
apfelbecki (Molops) 143
apfelbecki (Nebria) 19
apfelbecki (Pterostichus) 153
apfelbecki (Zabrus) 183
apfelbecki apfelbecki (Molops) 143
apfelbecki bischoffi (Molops) 143
apfelbecki corallipes (Molops) 142
apfelbecki kulti (Nebria) 19
apfelbecki plurisetosus (Molops) 145
apfelbecki pseudoalpestris (Molops) 142
apfelbecki reiseri (Molops) 142, 143
apfelbecki victoris (Molops) 142, 143
Aphaenopsis 79
Aphaon 155
apicalis (Dyschirius) 47
APOTOMINAE 51, 187
APOTOMINI 51
Apotomus 51
apricaria (Amara) 177
Apristus 127
apterus (Microlestes) 124
Aptinus 39
Aptinus s.str. 39
aquaticus (Notiophilus) 24
Archicarabus 30
ardosiacus (Ophonus) 112
arenaria (Cicindela) 28
arenaria (Cylindera) 28
arenaria viennensis (Cylindera) 28
areolatus (Perileptus) 79
areolatus areolatus (Perileptus) 79
Argutor 151
articulatum (Bembidion) 72
Asaphidion 52
ascendens (Bembidion) 56, 57
assimile (Bembidion) 58
assimilis (Platynus) 138
ATRANOPSINA 165
atratus (Harpalus) 100
atratus var. subsinuatus (Harpalus) 100
atrocaeruleum (Bembidion) 56
attemsi (Nebria) 19
attenuatus (Harpalus) 99
atterimus (Pterostichus) 152
atterimus atterimus (Pterostichus) 152
aulica (Amara) 178
auriculatus (Dyschirius) 47
austriacus (Molops) 147
autumnalis (Harpalus) 104
axillare (Bembidion) 58
axillare occiduum (Bembidion) 58
axillaris (Cymindis) 121
axillaris axillaris (Cymindis) 121
axillaris adriatica (Cymindis) 121
axillaris lineola (Cymindis) 122
azurescens (Bembidion) 59
azurescens azurescens (Bembidion) 59
azureus (Chlaenius) 91
azureus (Harpalus) 109
azureus (Ophonus) 109, 110
azureus oberthuri (Harpalus) 109
babinjensis (Carabus) 32
babinjensis (Trechus) 85
bacillus (Dyschirius) 47
bacillus arbensis (Dyschiriodes) 47
bacillus arbensis (Dyschirius) 47

227. 226 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
bacillus bacillus (Dyschirius) 47
bacillus lucanus (Dyschirius) 47
Badister 128
Badister s.str. 128
balcanicum (Bembidion) 64
balcanicum balcanicum (Bembidion) 64, 65, 66
baldori sic! (Zabrus) 184
Batenus 138
Baudia 129
baudii (Acinopus) 97
baudii hellenicus (Acinopus) 97, 98
bayardi (Brachinus) 43
Bedelinus 166
BEMBIDIINA 52
BEMBIDIINI 12, 52
Bembidion 12, 53
Bembidion s.str. 53
Bembidionetolitzkya 54
beroni (Albanotrechus) 12, 79
Bicharotrechus 79
bifrons (Amara) 178
biguttatus (Notiophilus) 24
binaghii (Agonum) 133
binotatus (Anisodactylus) 94
binotatus ab. spurcaticornis (Anisodactylus) 94
bipunctatum (Bembidion) 71
bipunctatum nivale (Bembidion) 71
bipunctatum var. pyritosum (Bembidion) 71
bipustulatus (Brachinus) 43
bisbimaculatus (Tachys) 76
bischoffi (Amara) 180
bischoffi (Duvalius) 80
bischoffi (Harpalus) 105
bischoffi (Neotrechus) 83
bischoffi (Reicheadella) 45
bischoffi (Reicheia) 45
bischoffi (Tapinopterus) 163, 164
bischoffi (Zabrus) 184
bischoffianus (Pterostichus) 155, 156
bistriatus (Paratachys) 74
bistriatus (Tachys) 74
bodemeyeri (Brachinus) 41
boldorii (Zabrus) 184
bombarda (Aptinus) 39
Boreonebria 22
bosnica (Nebria) 19
bosnica apfelbecki (Nebria) 19
bosnica jezercensis (Nebria) 19
bosnica kulti (Nebria) 19
bosnica prokletiensis (Nebria) 19
bosnica sturanyi (Nebria) 19
bosnica tresnjevikensis (Nebria) 20
bosnicus (Calathus) 166
Bothriopterus 151
Brachynidius 41
BRACHININAE 7, 39, 187
BRACHININI 39
Brachynolomus 42
Brachinus 40
Brachinus s.str. 40
Brachyophonus 109
Bracteon 57
Bradycellus 12, 118
Bradycellus s.str. 118
Bradytus 177
brevicollis (Brachinus) 42
brevicollis (Nebria) 22, 23
brevicollis (Ophonus) 111
brevicollis var. salina (Nebria) 22, 23
brevis (Pterostichus) 153
brevis brevis (Pterostichus) 153
brucki (Pterostichus) 155, 156
brunnicorne (Bembidion) 66
brunnicorne brunnicorne (Bembidion) 66
brunnipes (Acupalpus) 115
brunoi (Bembidion) 56, 57
bualei (Bembidion) 68
bualei bualei (Bembidion) 68
bucephalus (Carabus) 35
bullatus (Badister) 128
buparius (Scarites) 51
burmeisteri (Pterostichus) 151
burmeisteri burmeisteri (Pterostichus) 151
caelatus (Carabus) 31
caelatus caelatus (Carabus) 31
caelatus caelatus m. malissorum (Carabus) 31
caelatus caelatus n. metalkanus (Carabus) 31
caelatus dalmatinus (Carabus) 32
caelatus dalmatinus n. procerus (Carabus) 32
caelatus malissorum (Carabus) 31
caelatus metalkanus (Carabus) 31
caelatus metalkanus v. malissorum (Carabus) 31

228. INDEX 227
caelatus sarajevensis sic! (Carabus) 31
caelatus sarajevoensis (Carabus) 31
caelatus var. procerus (Carabus) 32
CALATHINA 166
Calathus 166
Calathus s.str. 166
CALLISTINA 89
Callistus 89
callosum (Bembidion) 61
callosum callosum (Bembidion) 61
Calomera 25
Calosoma 29
Calosoma s.str. 29
calydonius (Ditomus) 95
calydonius calydonius (Ditomus) 95
campestris (Cicindela) 26, 27
campestris campestris (Cicindela) 26
campestris olivieria (Cicindela) 26, 27
campestris pontica (Cicindela) 26
cancellatus (Carabus) 36
cancellatus alessiensis (Carabus) 36
cancellatus emarginatus (Carabus) 36
cancellatus subgraniger var. alessiensis (Carabus) 36
cancellatus var. alessiensis (Carabus) 36
CARABINAE 7, 29, 187
CARABINI 29
caraboides (Asaphidion) 52, 53
caraboides balcanicus (Asaphidion) 52
caraboides var. balcanicus (Asaphidion) 52
Carabus 29
Carabus s.str. 30
carbonarium (Agonum) 133
carbonarium carbonarium (Agonum) 133
cardioderus (Polyderis) 74
cardioderus balcanicus (Trechus) 86
carinatus (Abax) 138
carinatus carinatus (Abax) 138
carinatus var. latus (Abax) 138
Carterus 95
Carterus s.str. 95
cassideus (Licinus) 129
cassideus cassideus (Licinus) 129
cassideus dohrni (Licinus) 129
cassideus graecus (Licinus) 129, 130
cassideus latus (Licinus) 129
catenatus (Carabus) 31
catenatus var. albanensis (Carabus) 31
catenulatus (Carabus) 31
catenulatus catenulatus (Carabus) 31
caucasicum (Bembidion) 62
caucasicus (Bradycellus) 118
cavernosus (Carabus) 34
cavernosus cavernosus (Carabus) 34
cavernosus n. cavernosus (Carabus) 34
cavernosus var. sterbai (Carabus) 34
cavicola (Laemostenus) 171
cavicola albanicus (Laemostenus) 171
cavicola modestus (Laemostenus) 171, 172
cavicola var. modestus (Laemostenus) 171, 172
Celia 178
centralis (Harpalus) 98
Cephalota 26
Chaetocarabus 30
chalybaeus (Myas) 148
chalybaeus (Pterostichus) 158
chalybeus (Dyschirius) 49
chalybeus gibbifrons (Dyschirius) 49
chaudoiri (Amara) 181
chaudoiri incognita (Amara) 181
Cheporus 151
chionobatus (Carabus) 30
chionophilus (Carabus) 36
Chiridysus 46
Chlaeniellus 89
CHLAENIINA 89
CHLAENIINI 89
Chlaenites 90
Chlaenius 89
Chlaenius s.str. 91
Chlorodium 58
Cicindela 26
Cicindela s.str. 26
CICINDELINAE 7, 12, 25, 187
CICINDELINI 25
cimmerius (Laemostenus) 173
cimmerius (Pristonychus) 173
cimmerius (Sphodrus) 173
cimmerius cimmerius (Laemostenus) 173
cimmerius weiratheri (Laemostenus) 173
cinctus (Calathus) 169, 170, 171
circumdata (Cephalota) 26
circumdata circumdata (Cephalota) 26

229. 228 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
circumscriptus (Chlaenius) 92
circumseptus (Calathus) 166
clathratus (Carabus) 31
clathratus auraniensis (Carabus) 31
Clivina 44
CLIVININA 44
CLIVININI 44
clypeatus (Aristus) 96
clypeatus (Ditomus) 96
clypeatus (Dixus) 96
Cnecostolus 43
coadunata (Cymindis) 121
coadunata coadunata (Cymindis) 121
coerulescens (Poecilus) 151
coerulescens (Pterostichus) 151
coeruleum (Bembidion) 57
coeruleum concoeruleum (Bembidion) 57
collaris (Bradycellus) 118
collaris (Clivina) 44
columbinus (Ophonus) 113
combustum (Bembidion) 60, 78
combustum combustum (Bembidion) 60
comita (Tapinopterus) 164
complanata (Amara) 180
complanata (Eurynebria) 16
complanata albanica (Eurynebria) 16
complanata orientalis (Eurynebria) 16
complanata var. albanica (Eurynebria) 16
complanatum (Bembidion) 55
concoeruleum (Bembidion) 57
concoeruleus (Ocydromus) 57
conforme (Bembidion) 55
connexus (Polystichus) 185
consularis (Amara) 178
convexicollis (Pogonistes) 78
convexior (Amara) 175
convexiusculus (Argutor) 153
convexiusculus (Pterostichus) 153
convexiusculus (Tapinopterus) 161
convexus (Carabus) 36
convexus dilatatus (Carabus) 36
convexus dilatatus n. miliusi (Carabus) 36
convexus dilatatus var. miliusi (Carabus) 36
convexus miliusi (Carabus) 36
convexus var. chionophilus (Carabus) 36
corax (Pterostichus) 160
corax sic! (Calathus) 168
cordatus (Ophonus) 111
coriaceus (Carabus) 35
coriaceus cerisyi (Carabus) 35
coriaceus cerisyi m. weiratheriana (Carabus) 35
coriaceus excavatus (Carabus) 35
coriaceus excavatus dalmaticus (Carabus) 36
coriaceus excavatus m. dalmatica (Carabus) 35
coriaceus excavatus n. joniensis (Carabus) 35
coriaceus joniensis (Carabus) 36
coriaceus joniensis var. albanicus (Carabus) 36
coriaceus mediterraneus (Carabus) 36
coriaceus rugosus (Carabus) 35
coriaceus rugosus n. joniensis m. albanicus (Carabus) 35
coriaceus subrugosus (Carabus) 35
coriaceus subrugosus v. weiratherianus (Carabus) 35
coriaceus var. hopffgarteni (Carabus) 35
corticalis (Microlestes) 123
corticalis escorialensis (Microlestes) 123
crepitans (Brachinus) 40
crepitans ab. fallax (Brachinus) 40
cribricollis (Harpalus) 110
cribricollis (Ophonus) 110
croaticus (Carabus) 32
croaticus babinjensis (Carabus) 32
croaticus durmitorensis (Carabus) 32
croaticus durmitorensis var. babinjensis (Carabus) 32
croaticus durmitorensis var. ljubetensis (Carabus) 32
croaticus ljubetensis (Carabus) 32
croaticus ljubetensis m. durmitorensis (Carabus) 32
croaticus ljubetensis n. ljubetensis (Carabus) 32
cruciatum (Bembidion) 68
cruciatum albanicum (Bembidion) 68
cruralis (Chlaenius) 91
cruxmajor (Panagaeus) 133
cruxmajor ab. schaumi (Panagaeus) 133
cruxminor (Lebia) 126
cruxminor cruxminor (Lebia) 126
Cryptophonus 98
cupreus (Harpalus) 105
cupreus (Poecilus) 149
cupreus (Pterostichus) 149
cupreus cupreus (Poecilus) 149
cupreus fastuosus (Harpalus) 105
cursitans (Amara) 180
cursor (Pterostichus) 151

230. INDEX 229
cursorius (Poecilus) 150
cursorius (Pterostichus) 150
cursorius cursorius (Poecilus) 150
curta (Amara) 175
Curtonotus 178
curtulus (Molops) 144
curtulus cakorensis (Molops) 144
curtulus winklerianus (Molops) 144
cyaneomicans (Bembidion) 61
cyanicorne (Asaphidion) 52
cyanicorne cyanicorne (Asaphidion) 52
cyanocephala (Lebia) 125, 126
cyanocephala cyanocephala (Lebia) 125
CYCHRINI 37
Cychrus 37
CYCLOSOMINI 92
Cylindera 28
Cylindera s.str. 28
cylindricus (Dyschirius) 47
cylindricus hauseri (Dyschirius) 47
CYMINDIDINA 121
Cymindis 121
Cymindis s.str. 121
dahli (Nebria) 20
dahli montenegrina (Nebria) 20
dahli velebitica (Nebria) 20
dalmaticum sic! (Bembidion) 67
dalmaticus (Carabus) 35
dalmatina (Amara) 179
dalmatinum (Bembidion) 67, 90
dalmatinum dalmatinum (Bembidion) 67
dalmatinus (Laemostenus) 172
dalmatinus (Platyderus) 165
dalmatinus dalmatinus (Platyderus) 165
dalmatinus malissorum (Neotrechus) 84
dalmatinus pentheri (Neotrechus) 84
dalmatinus procerus (Carabus) 32
dalmatinus suturalis var. pentheri (Neotrechus) 84
Dalmatoreicheia 45
dama (Carterus) 95
dama (Ditomus) 95
danieli (Harpalus) 104
danieli (Notiophilus) 24
decipiens (Chlaenius) 91
decipiens laticollis (Chlaenius) 91
decipiens var. ambiguus (Chlaenius) 91
decolor (Bembidion) 69
decorum (Bembidion) 63
decorum decorum (Bembidion) 63
dejeani (Chlaenius) 92
dejeani (Epomis) 92
dejeani (Parophonus) 114
deletum (Bembidion) 67
deletum deletum (Bembidion) 67
Deltomerus 76
Deltomerus s.str. 76
dentata (Drypta) 93
depressa (Siagona) 44
detonii (Tapinopterus) 164
diabrachys (Tachys) 76
diabrachys (Tachyura) 76
Diachromus 94
Dicheirotrichus 118
Dicheirotrichus s.str. 118
didonnai (Duvalius) 80
diffinis (Harpalus) 113
diffinis (Ophonus) 113
dilatatus (Badister) 129
diligens (Pterostichus) 153
dimidiatus (Harpalus) 105
dinaricus (Cychrus) 37
Dinodes 91
Diplocampa 58
discophorus (Acupalpus) 119
discophorus (Stenolophus) 119
distinctissimus (Trechus) 87
distinctus (Bradycellus) 118
distinguendus (Calathus) 167
distinguendus (Harpalus) 107
distinguendus distinguendus (Harpalus) 107
DITOMINA 95
Ditomus 95
Dixus 96
djumanus (Pterostichus) 158
dochii (Pterostichus) 164
dochii (Tapinopterus) 164
doderoi (Sinechostictus) 73
dorsale (Platynus) 136
dorsalis (Acupalpus) 117
dorsalis (Anchomenus) 136
dorsalis (Platynus) 136

231. 230 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
DROMIINA 123
Dromius 123
Dromius s.str. 123
droveniki (Duvalius) 79
Drypta 93
Drypta s.str. 93
DRYPTINI 93
duftschmidi (Agonum) 133
Duvalius 79
Duvalius s.str. 80
dyrrhachii (Bembidion) 63
DYSCHIRIINI 46
Dyschiriodes 46
Dyschirius 12, 46
Dyschirius s.str. 49
effluviorus (Sinechostictus) 73
Egadroma 119
egregius (Trechus) 87
ejaculans (Brachinus) 41
ELAPHRINAE 38, 187
ELAPHRINI 38
Elaphrus 38
elegans (Acupalpus) 115
elegans (Bembidion) 52, 53, 57, 58
elegans (Brachinus) 41
ellipticus (Calathus) 167
ellipticus ellipticus (Calathus) 167
elongatum (Bembidion) 73
elongatum tarsicum (Bembidion) 73
elongatum var. effluviorum (Bembidion) 73
elongatus (Laemostenus) 172
elongatus (Pterostichus) 152
elongatus (Sinechostictus) 73
elongatus pentheri (Laemostenus) 172
emarginatus (Acinopus) 97
Emphanes 58
eomita sic! (Tapinopterus) 164
ephippium (Bembidion) 62
epirensis (Carabus) 31
epiroticus (Pterostichus) 160
Epomis 92
equestris (Amara) 179, 180
equestris equestris (Amara) 179, 180
eremita (Ditomus) 96
eremita (Dixus) 96
erratica (Amara) 177
erratus (Calathus) 170
erratus erratus (Calathus) 170
erythrocnema (Amara) 181
erythroderus (Calathus) 169
escorialensis (Microlestes) 123
etruscus (Gynandromorphus) 95
etruscus etruscus (Gynandromorphus) 95
Eucarabus 31
euchlorus (Harpalus) 105
Euduvalius 82
Eudyschirius 49
Eugrapha 28
Eunebria 22
Euperyphus 60
europaea (Siagona) 44
europaea europaea (Siagona) 44
europaea oberleitneri (Siagona) 44
europaea var. oberleitneri (Siagona) 44
Europhilus 136
Eurynebria 16
eurynota (Amara) 175
exhalans (Brachinus) 43
exilis (Microlestes) 124
exilis luctuosus (Microlestes) 124
explodens (Brachinus) 42
explodens ab. glabratus (Brachinus) 42
explodens ab. strepens (Brachinus) 42
explodens sichemita (Brachinus) 42
extensoides Jedlička, 1936 (Tapinopterus) 161, 162
extensoidesStraneo,1936(Tapinopterus)161,162
extensum (Agonum) 133
extensus (Tapinopterus) 161, 162
extensus abreshensis (Tapinopterus) 161
extensus convexiusculus (Pterostichus) 161
extensus extensoides (Tapinopterus) 161, 162
fallaciosa (Amara) 179
fallaciosus (Zabrus) 179
fallax (Brachinus) 40
fallax (Parophonus) 114
fallax (Tachyophonus) 114
famelica (Amara) 175
familiaris (Amara) 175
fasciatopunctatus (Pterostichus) 155
fasciolatum (Bembidion) 56, 57
femoralis (Carterophonus) 96
femoralis (Graniger) 96

232. INDEX 231
femoratum (Bembidion) 68
femoratum femoratum (Bembidion) 68
Feronidius 152
festivus (Chlaenius) 91
festivus festivus (Chlaenius) 91
fischeri (Calomera) 25
fischeri (Cicindela) 25
fischeri fischeri (Calomera) 25
fissuralis (Microlestes) 123
flavescens (Harpalus) 103
flavicornis (Harpalus) 101
flavicornis flavicornis (Harpalus) 101
flavipes (Asaphidion) 52, 53
flavipes (Chlaenius) 89
fodori (Duvalius) 82
foraminosum (Bembidion) 66
fossor (Clivina) 45
fossor fossor (Clivina) 45
franziniorum (Ophonus) 113
fuchsi (Duvalius) 81
fugax (Harpalus) 108
fulvibarbis (Leistus) 16
fulvibarbis fulvibarbis (Leistus) 16
fulvicollis (Paratachys) 74
fulvicollis (Tachys) 74
fulvipes (Amara) 181
fumigatum (Bembidion) 58
fusca (Amara) 180
fuscatus (Olisthopus) 137
fuscipes (Calathus) 167
fuscipes fuscipes (Calathus) 167
fuscipes graecus (Calathus) 167
fuscipes var. graecus (Calathus) 167
fuscipes var. latus (Calathus) 167
galilaeus (Lymnastis) 73
ganglbaueri (Brachinus) 41
ganglbaueri (Bradycellus) 118
ganglbaueri (Nebria) 20
ganglbaueri (Synuchidius) 171
ganglbaueri (Zabrus) 183
ganglbaueri ditana (Nebria) 21
ganglbaueri ganglbaueri (Nebria) 20
ganglbaueri ganglbaueri (Zabrus) 183
ganglbaueri gyalicaensis (Nebria) 20
ganglbaueri korabensis (Nebria) 20
ganglbaueri lonae (Zabrus) 183
ganglbaueri merditana (Nebria) 20, 21
ganglbaueri zebiaensis (Nebria) 21
genei illigeri (Bembidion) 62
geniculatum (Bembidion) 55
geniculatum geniculatum (Bembidion) 55
georgii (Duvalius) 80
germanica (Cicindela) 28
germanica (Cylindera) 28
germanica germanica (Cylindera) 28
germanica muelleri (Cylindera) 28
germanus (Diachromus) 94
germarii (Nebria) 21
germarii durmitorensis (Nebria) 21
germinyi (Notiophilus) 24
gestroi (Tapinopterus) 165
gibbifrons (Dyschirius) 49
giganteus (Calathus) 167, 168
gigas (Carabus) 35
gigas gigas (Carabus) 35
gigas parnassicus (Carabus) 35
gilvipes (Carterus) 95
gisellae (Agonum) 134
glabratus (Brachinus) 42
glabricollis (Calathus) 168
glabricollis (Olisthopus) 137
globosus (Dyschirius) 49
gracilis (Dyschirius) 49, 50
gracilis (Oodes) 131
gracilis (Pogonistes) 78
gracilis (Pterostichus) 155
gracilis gracilis (Dyschirius) 49
gracilis gracilis (Pterostichus) 155
graecus (Carabus) 35
graecus (Licinus) 129, 130
graecus (Platyderus) 165
graecus graecus (Carabus) 35
graecus morio (Carabus) 35
graecus morio var. neowinkleri (Carabus) 35
graecus morio var. winkleri (Carabus) 35
graecus n. winkleri (Carabus) 35
graecus winkleri (Carabus) 35
grandipenne (Bembidion) 67
grandipenne grandipenne (Bembidion) 67
Graniger 96
granulatus (Carabus) 30
granulatus aetolicus (Carabus) 30

233. 232 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
granulatus granulatus m. merdita sic! (Carabus) 30
granulatus granulatus n. interstitialis (Carabus) 30
granulatus interstitialis (Carabus) 30
granulatus interstitialis v. miridita (Carabus) 30
granulatus miridita (Carabus) 30
granulatus m. miridita (Carabus) 30
granulatus var. miridita (Carabus) 30
granulipennis (Aptinus) 39
griseus (Harpalus) 108
gudenzii (Bembidion) 64
guttula (Bembidion) 69
guttula guttula (Bembidion) 69
gyllenhali (Nebria) 22
Gynandromorphus 95
haemorrhoidalis (Sphaerotachys) 75
haemorrhoidalis (Tachys) 75
hannemanni (Amara) 178
HARPALINA 97
HARPALINAE 7, 89, 187
HARPALINI 12, 93
harpaloides (Ocys) 72
Harpalus 12, 98
Harpalus s.str. 99
helopioides (Oodes) 131
helopioides helopioides (Oodes) 131
herzegowinensis (Carabus) 34
Hesperophonus 109
hilleri (Carabus) 34
hirsutulus (Harpalus) 113
hirsutulus (Ophonomimus) 113
hirsutulus (Ophonus) 113
hirsutulus (Parophonus) 113
holdhausi (Agonum) 134
holdhausi (Molops) 142, 143, 144, 145
holdhausi (Platynus) 134
honestus (Harpalus) 99
honestus honestus (Harpalus) 99
hopffgarteni (Carabus) 35
hortensis (Carabus) 33, 34
hortensis hortensis (Carabus) 34
hortensis neumeyeri (Carabus) 33, 34
hortensis neumeyeri n. herzegowinensis (Carabus) 34
hortensis neumeyeri n. neumeyeri (Carabus) 34
hortensis neumeyeri var. albanicola (Carabus) 34
hortensis neumeyeri var. hilleri (Carabus) 34
hortensis neumeyeri var. pindicus (Carabus) 34
hortensis preslii (Carabus) 33
hortensis preslii m. jonica (Carabus) 34
hortensis rhodopensis (Carabus) 34
hospes (Harpalus) 106
hospes hospes (Harpalus) 106
humerale (Bembidion) 53
humeralis (Cymindis) 121
humeralis (Lebia) 126
hybrida (Cicindela) 27
hybrida albanica (Cicindela) 27
hybrida riparia (Cicindela) 27
Hygrocarabus 31
hypocrita (Agonum) 134
hypocrita (Bembidion) 66
hypocrita (Notiophilus) 24
hypocrita illyricum (Bembidion) 66
Hypogeobium 160
Hypogium 160
hypsobius (Carabus) 36
ignavus (Zabrus) 183
illigeri (Bembidion) 62
immaculicornis (Brachinus) 42
immaculicornisimmaculicornis(Brachinus)42
immundus (Acupalpus) 116
importunus (Dyschirius) 50
importunus importunus (Dyschirius) 50
impressum sic! (Syntomus) 127
impressus (Metabletus) 127
impressus (Syntomus) 127
impressus impressus (Syntomus) 127
inaequalis (Tachys) 76
incertus (Brachinus) 43
Incisophonus 111
incisus (Ophonus) 111
incrassatus (Zabrus) 184
incrassatus bischoffi (Zabrus) 184
incrassatus boldorii (Zabrus) 184
incrassatus incrassatus (Zabrus) 184
inoptatum (Bembidion) 70
inquinatus (Pedius) 149
inquinatus (Pterostichus) 149
inquisitor (Calosoma) 29
inquisitor inquisitor (Calosoma) 29
inserticeps (Bembidion) 61
intermedius (Anisodactylus) 94
intermedius (Dyschirius) 46

234. INDEX 233
intricatus (Carabus) 30
intricatus chinobatus (Carabus) 30
intricatus intricatus (Carabus) 30
intricatus intricatus m. montenegrinus (Carabus) 30
intricatus intricatus n. weiratheri (Carabus) 30
intricatus intricatus var. weiratheri (Carabus) 30
intricatus montenegrinus (Carabus) 30
intricatus var. montenegrinus (Carabus) 30
irenis (Trechus) 86
iricolor (Bembidion) 70
irregularis (Carabus) 35
irregularis bucephalus (Carabus) 35
irregularis ramanus (Carabus) 35
janthinus (Laemostenus) 172
janthinus janthinus (Laemostenus) 172
javurkovae (Bembidion) 68
jockischi (Nebria) 22
jockischi jockischi (Nebria) 22
jonicus (Carabus) 34
joniensis (Carabus) 35
justinae (Bembidion) 55
kaufmanni (Tapinopterus) 161
klisuranus (Molops) 144, 146
kobingeri (Trechus) 87
kobingeri bulgaricus (Trechus) 88
kobingeri kobingeri (Trechus) 87, 88
kobingeri pawlowskianus (Trechus) 88
korabensis (Carabus) 33
korabica (Nebria) 21
korax (Calathus) 167, 168
koyi (Poecilus) 150
koyi (Pterostichus) 150
koyi dinaricus (Pterostichus) 150
koyi koyi (Poecilus) 150
kratteri (Nebria) 23
kratteri kratteri (Nebria) 23
kratteri valonensis (Nebria) 23
krueperi (Harpalus) 109
krueperi (Ophonus) 109
Laemostenus 171
Laemostenus s.str. 172
laeviceps (Parophonus) 114, 115
laevicollis (Trichotichnus) 115
laevicollis laevicollis (Trichotichnus) 115
laevifrons (Clivina) 45
laevigatus (Scarites) 50
laevipes (Harpalus) 101
laeviusculus (Dyschirius) 47
lafertei (Dyschirius) 49
Lamprias 125
lampros (Bembidion) 60
lampros var. properans (Bembidion) 61
laterale (Bembidion) 61
laterale pseudolaterale (Bembidion) 61
laticollis (Chlaenius) 91
laticollis (Ophonus) 111
latifianus (Pterostichus) 157
latifianus ab. nigripes (Pterostichus) 158, 159
latifianus latifianus (Pterostichus) 156, 157
latifianus strupii (Pterostichus) 156, 157,
158, 159
latipennis (Dyschirius) 49
latiplaga (Bembidion) 59
latiplaga latiplaga (Bembidion) 59
latus (Abax) 138
latus (Calathus) 167
latus (Harpalus) 103
latus (Licinus) 129
Lebia 125
Lebia s.str. 126
LEBIINA 125
LEBIINI 121
Leistus 16
Leistus s.str. 16
leonhardi stenocephalus (Duvalius) 81
leonhardi sturanyi (Duvalius) 81, 82
lepidus (Poecilus) 150
lepidus (Pterostichus) 150
lepidus lepidus (Poecilus) 150
levantinus (Amblystomus) 93
levantinus levantinus (Amblystomus) 93
LICININA 128
LICININI 128
Licinus 129
Licinus s.str. 129
limbatum (Omophron) 38
limbopunctatus (Harpalus) 107
Limnocarabus 31
linearis (Dromius) 125
linearis (Paradromius) 125
linearis linearis (Paradromius) 125
lineata (Cymindis) 122

235. 234 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
lineola (Cymindis) 122
LIONYCHINA 127
Lionychus 127
Lionychus s.str. 127
literata sic! (Cicindela) 28
litorale (Bembidion) 57, 58
litoralis sic! (Pogonus) 78
litterata (Cicindela) 28
littoralis (Calomera) 26
littoralis (Pogonus) 78
littoralis nemoralis (Calomera) 26
ljubetensis (Carabus) 32
lodosi (Pterostichus) 153
lonae (Neotrechus) 82
lonae (Trechus) 82
lonae atavus (Neotrechus) 83
lonae lonae (Neotrechus) 82, 83
lonae shkodrensis (Neotrechus) 83
lonai (Omphreus) 131
lonai sic! (Neotrechus) 82
lonai sic! atavus (Neotrechus) 83
lonai sic! lonai sic! (Neotrechus) 82
lonai sic! shkodrensis (Neotrechus) 83
longicollis (Pedius) 149
longicorne (Agonum) 134
longicornis (Anthracus) 117
longicornis (Thalassophilus) 88
Loxoncus 119
lucida (Amara) 176
lucidus (Dyschirius) 46
luctuosus (Microlestes) 124
luctuosus luctuosus (Microlestes) 124
lugubris rufipes (Aptinus) 39, 40
lumensis (Pterostichus) 157, 158
lumensis litae (Pterostichus) 156, 158
lumensis ljubetensis (Pterostichus) 156, 158
lumensis lumensis (Pterostichus) 156, 157
lunatus (Callistus) 89
lunatus lunatus (Callistus) 89
lunulata (Cicindela) 26
lunulata var. nemoralis (Cicindela) 26
lunulatum (Bembidion) 70
luridipennis (Pogonus) 78
luteatus (Acupalpus) 115
luticola (Dyschirius) 47
luticola luticola (Dyschirius) 47
Lymnastis 73
macedonica (Nebria) 18, 21
macedonica galicica (Nebria) 18, 21
macedonica papingutica (Nebria) 18, 21
macedonica rambouseki (Nebria) 18, 22
macedonicus (Calathus) 168
macedonicus (Pachycarus) 97
macroderus (Dyschirius) 48
macroderus macroderus (Dyschirius) 48
maculatum (Bembidion) 72
maculatum maculatum (Bembidion) 72
maculatus (Acupalpus) 116, 117
maculicornis (Harpalus) 114
maculicornis (Ophonus) 114
maculicornis (Parophonus) 114
maculicornis (Trichotichnus) 114
maderi (Molops) 139
maglicensis (Duvalius) 79
magnicollis (Leistus) 17
malissorum (Carabus) 31, 32
malissorum (Deltomerus) 76, 77
malissorum (Neotrechus) 83, 84
malissorum (Molops) 145
malissorum (Pterostichus) 156, 158, 159
malissorum (Trechus) 84
malissorum ab. munella (Pterostichus) 157
malshentianus (Molops) 139
mannerheimii (Bembidion) 70
Manodromius 125
marginata (Egadroma) 119
marginatum (Agonum) 134
marginatus (Stenolophus) 119
maritimus (Lionychus) 127
MASOREINA 92
Masoreus 92
Masoreus s.str. 92
maurus (Microlestes) 124
maurus maurus (Microlestes) 124
megacephalus (Acinopus) 98
Megodontus 29, 31
meisteri (Pterostichus) 156
melanarius (Pterostichus) 153
melanarius melanarius (Pterostichus) 153
melancholica (Myriochila) 28
melancholica melancholica (Myriochila) 28
Melanius 152

236. INDEX 235
melanocephalus (Calathus) 170
melanocephalusmelanocephalus(Calathus)170
melanura (Odacantha) 131
melas (Pterostichus) 152
melas depressus (Pterostichus) 152
melletii (Ophonus) 111
Menas 122
mendax (Harpalus) 114
mendax (Ophonus) 114
mendax (Parophonus) 114
mendax (Tachyophonus) 114
merditana (Nebria) 20, 21
merditanum (Bembidion) 57
merditanus (Aptinus) 39
merditanus (Carabus) 33
merditanus (Molops) 141, 142
merditanus (Trechus) 86
merditanus intermedius (Molops) 142
merditanus merditanus (Aptinus) 39, 40
merditanus merditanus (Molops) 141
merditanus var. granulipennis (Aptinus) 40
meridianus (Acupalpus) 116
meridionale (Bembidion) 59
meridionalis (Ophonus) 110
merklii (Pterostichus) 156
meschniggi (Bembidion) 56
messae (Amara) 180
metalkanus (Carabus) 31
metallescens (Amblystomus) 93
metallicus (Calathus) 169, 170, 171
metallicus (Pterostichus) 151
metallicus metallicus (Calathus) 170
metallicus prokletijensis (Calathus) 170
Metallina 60
Metophonus 111
Microcallisthenes 29
Microlestes 12, 123
micros (Paratachys) 74
micros (Tachys) 74
Microserrullula 61
miliaris (Cymindis) 123
miliusi (Carabus) 36
milleri (Bembidion) 67, 68
milleri milleri (Bembidion) 68
millerianus (Sinechostictus) 73
minor (Pterostichus) 155
minor minor (Pterostichus) 155
minutulus (Blechrus) 124
minutulus (Microlestes) 124
minutulus (Speluncarius) 160
minutulus (Tapinopterus) 160
minutulus apfelbecki (Microlestes) 124
minutus (Dyschirius) 48
minutus (Platyderus) 166
minutus albanicus (Dyschirius) 48
minutus obtusangulus (Platyderus) 166
miridita (Tapinopterus) 164
miridita jablanicensis (Tapinopterus) 164, 165
miridita jacupicensis (Tapinopterus) 165
miridita miridita (Tapinopterus) 164
miriditus (Carabus) 30
mixtus (Stenolophus) 120
moestum (Agonum) 133
moestus (Platynus) 133
mollis (Calathus) 171
mollis f. erythroderus (Calathus) 169
mollis mollis (Calathus) 171
Molops 139
Molops s.str. 139
monachum var. binaghii (Agonum) 133
monastirensis (Tapinopterus) 165
monastirensis gestroi (Tapinopterus) 165
montana (Amara) 178
montenegrinus (Carabus) 30
montenegrinus (Molops) 147, 148
montenegrinus cavernicola (Molops) 147
monticola (Cicindela) 27
monticola albanica (Cicindela) 27
montivagus (Carabus) 30
montivagus m. sutomorensis (Carabus) 30
montivagus vellepiticus (Carabus) 30
morio (Amara) 176
morio (Dyschirius) 48
morio (Omphreus) 132
morio albanicus (Omphreus) 132
morio malissorum (Omphreus) 132
morio nivium (Amara) 176
Morphnosoma 153
muelleri (Agonum) 134, 135
muelleri (Duvalius) 80
muelleri (Platynus) 134
munella (Pterostichus) 157

237. 236 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
municipalis (Amara) 180
municipalis bischoffi (Amara) 180
Myas 148
Myas s.str. 148
Myriochila 28
Myriochila s.str. 28
Mystropterus 97
nana (Tachyta) 75
Nebria 12, 18
Nebria s.str. 22
NEBRIINAE 7, 16, 187
NEBRIINI 16
nebulosum (Asaphidion) 52, 53
nebulosum balcanicum (Asaphidion) 52, 58
negrita (Microlestes) 124
negrita apfelbecki (Microlestes) 124
nemoralis nemoralis (Lophyridia) 26
nemorivagus (Anisodactylus) 94
Neocalathus 169
Neoelaphrus 38
Neorescius 130
Neotrechus 82
neowinkleri (Carabus) 35
Nepha 61
neresheimeri (Bembidion) 70
neumeyeri (Carabus) 33, 34
niger (Amblystomus) 93
niger (Pterostichus) 154
niger niger (Pterostichus) 154
nigricornis (Brachinus) 43
nigricornis (Chlaenius) 67, 89, 90
nigrinus (Trechus) 87
nigripes (Pterostichus) 158
nigrita (Pterostichus) 155
nitidulus (Chlaenius) 90
nitidus (Dyschirius) 46
nitidus (Leistus) 17
nitidus nitidus (Dyschirius) 46
nivalis (Synuchus) 173
nodulosus (Carabus) 31
normannum (Bembidion) 59
normannum apfelbecki (Bembidion) 59
normannum mediterraneum (Bembidion) 59
normannum meridionale (Bembidion) 59
Notaphemphanes 62
Notaphus 63
notatus (Acupalpus) 116
notatus (Dromius) 125
notatus (Philorhizus) 125
NOTIOPHILINI 23
Notiophilus 12, 23
oberthuri (Ophonus) 109
oblitus (Harpalus) 108
oblitus oblitus (Harpalus) 108
oblongopunctatus (Pterostichus) 151
oblongopunctatus oblongopunctatus
(Pterostichus) 151
oblongum (Bembidion) 60
obscuroguttatus (Metabletus) 128
obscuroguttatus (Syntomus) 128
obscurus (Aristus) 96
obscurus (Ditomus) 96
obscurus (Dixus) 96
obsoletus (Dicheirotrichus) 118
obtusangulus intermedius (Molops) 142
obtusiusculus (Trechus) 88
obtusus (Trechus) 85
obtusus obtusus (Trechus) 85
octomaculatum (Bembidion) 72
Ocydromus 63
Ocys 72
Ocyturanes 64
Odacantha 130
Odacanta s.str. 131
ODACANTHINI 130
Odontium 66
Odontocarus 97
Oedematicus 98
oertzeni (Licinus) 130
oertzeni albanicus (Licinus) 130
oertzeni oertzeni (Licinus) 130
olens (Zuphium) 186
olens olens (Zuphium) 186
Olisthopus 12, 137
Omoperyphus 66
Omophron 38
Omophron s.str. 38
OMOPHRONINAE 7, 38, 187
OMOPHRONINI 38
OMPHREINI 131
Omphreus 131
Omphreus s.str. 131

238. INDEX 237
Oodes 131
OODINI 131
Ophonomimus 113
Ophonus 12, 109
Ophonus s.str. 112
Oreocarabus 33
Oreoplatysma 153
Osimus 98
osmanilis (Molops) 145
osmanilis osmanilis (Molops) 145, 146
osmanilis tomoricensis (Molops) 145, 146
ottomanus (Pterostichus) 154
ottomanus ottomanus (Pterostichus) 154
ovalis (Abax) 139
ovata (Amara) 176
Pachycarus 97
Pachystus 34
pallipes (Metabletus) 128
pallipes (Syntomus) 128
paludicola (Acupalpus) 116, 117
palustris (Notiophilus) 24
PANAGAEINI 133
Panagaeus 133
Panagaeus s.str. 133
Paracelia 179
Paradeltomerus 77
paradoxus (Deltomerus) 77
paradoxus korabensis (Deltomerus) 77
paradoxus paradoxus (Deltomerus) 77
Paradromius 125
Paradyschirius 50
Parahaptoderus 153
Parallelomorphus 50
parallelus (Dyschirius) 50
parallelus (Ophonus) 111
parallelus albanensis (Harpalus) 111
parallelus albanicus (Ophonus) 111
parallelus ruficornis (Dyschiriodes) 50
parallelus ruficornis (Dyschirius) 50
Paranchus 137
Parapercosia 179
Paratachys 74
parnassicola (Molops) 140
parnassicola alinae (Molops) 140
parnassicola valonensis (Molops) 140
parnassicus (Carabus) 35
Parophonus 12, 113
Parophonus s.str. 114
parreyssi (Molops) 141
parreyssi lonae (Molops) 141
parumpunctatus (Deltomerus) 77
parvicollis (Leistus) 17
parvula (Tachyura) 76
parvulus (Acupalpus) 117
parvulus (Tachys) 76
PATROBINI 76
Pedius 149
Pelor 181
peltatus (Badister) 129
peltatus peltatus (Badister) 129
pentheri (Calosoma) 29
pentheri (Pterostichus) 156, 159
pentheroides (Calosoma) 29
Percosia 179
peregrinus (Brachinus) 42
PERILEPTINA 79
Perileptus 79
Perileptus s.str. 79
peristericus (Tapinopterus) 162
permoestum (Agonum) 135
Peryphanes 66
Peryphus 68
Phaon 155
Pheggomisetes 88
Philochthus 69
Philorhizus 125
Philorhizus s.str. 125
Phonias 153
Phyla 70
picenus peristericus (Carabus) 33
piceus (Molops) 147
piceus balcanicus (Molops) 147
picipennis (Harpalus) 102
picipes (Acinopus) 97
pindicola (Cymindis) 122
pindicum (Bembidion) 64, 65
pindicus (Carabus) 34
plagiatus (Brachinus) 40, 41
plagiatus (Microlestes) 124
plagiatus escorialensis (Microlestes) 123
planicollis (Acupalpus) 117
planicollis Apfelbeck (Harpalus) 114

239. 238 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
planicollis Apfelbeck (Ophonus) 114
planicollis Apfelbeck (Parophonus) 114, 115
planicollis Dejean (Harpalus) 114
planicollis Dejean (Ophonus) 114
planicollis Dejean (Parophonus) 114
Platycarabus 35
Platyderus 165
Platyderus s.str. 165
PLATYNINI 133
Platynus 138
Platynus s.str. 138
Platypterinus 154
Platysma 154
plurisetosus (Molops) 145
Poecilus 149
Poecilus s.str. 149
POGONINI 78
Pogonistes 78
Pogonophorus 17
Pogonus 78
Pogonus s.str. 78
Polyderis 74
Polystichus 185
pominii (Duvalius) 81
praetermissa (Amara) 178
praeustum (Bembidion) 65, 74
praeustum praeustum (Bembidion) 65
praeustum viridifluum (Bembidion) 65
preslii (Carabus) 33, 34
preslii neumeyeri (Carabus) 34
priapus (Trechus) 86
priapus priapus (Trechus) 86
Princidium 71
Pristonychus 173
Procerus 29, 35
procerus (Anoplogenius) 119
procerus (Carabus) 32
procerus (Loxoncus) 119
Procrustes 29, 35
properans (Amara) 180
properans (Bembidion) 61
protensus (Tapinopterus) 162
protensus aetolicus (Tapinopterus) 162, 163
protensus peristericus (Tapinopterus) 162
protensus tomoricensis (Tapinopterus) 163
proxima (Amara) 176
proximus (Stenolophus) 120
Pseudodichirius 94
pseudolaterale (Bembidion) 61
Pseudolimnaeum 73
Pseudomaseus 154
Pseudophonus 108
psophia (Brachinus) 41
PTEROSTICHINI 138
Pterostichus 151
Pterostichus s.str. 155
pubescens (Harpalus) 108
pubescens (Ophonus) 108
pueli (Anisodactylus) 94
pueli pueli (Anisodactylus) 94
pumicatus (Stomis) 160
pumicatus pumicatus (Stomis) 160
pumilus (Harpalus) 102
punctatostriatus (Harpalus) 106
punctatulus (Harpalus) 111
punctatulus (Ophonus) 111
punctatus (Dyschirius) 48
punctatus (Laemostenus) 173
punctatus (Leistus) 18
punctatus albanicus (Dyschirius) 48
puncticeps (Harpalus) 112
puncticeps (Ophonus) 112
puncticollis Coquerel (Acupalpus) 117
puncticollis Auct. nec Coquerel (Acupalpus) 116
puncticollis (Harpalus) 112
puncticollis (Ophonus) 112
puncticollis (Poecilus) 149
puncticollis (Pterostichus) 149
punctulatum (Bembidion) 71
punctulatum punctulatum (Bembidion) 71
punctulatus sic! (Ophonus) 111
pusillus (Dyschirius) 48
pusillus (Notiophilus) 23
pygmaeus (Harpalus) 106
quadricolle (Bembidion) 61
quadriguttatum (Fabricius) (Bembidion) 53
quadriguttatum (Olivier) (Bembidion) 54
quadrillum (Lionychus) 127
quadrimaculatum (Bembidion) 53, 54
quadrimaculatum quadrimaculatum
(Bembidion) 53
quadrimaculatus (Dromius) 123

240. INDEX 239
quadripunctatus (Harpalus) 101
quadripustulatum (Bembidion) 54
quadripustulatum quadripustulatum
(Bembidion) 54
quadristriatus (Trechus) 85
quarnerensis (Anthracus) 117
quenseli (Amara) 179
quenseli quenseli (Amara) 179
ravasinianus (Trechus) 88
ravasinii (Calathus) 168
ravasinii (Omphreus) 132
ravasinii (Trechus) 88
ravasinii macedonicus (Calathus) 168
ravasinii ravasinii (Calathus) 168
rebeli (Poecilus) 150
rebeli (Pterostichus) 150
reflexicollis (Amara) 181
reflexicollis reflexicollis (Amara) 181
regiszogui (Duvalius) 81
Reicheadella 45
Reicheadella s.str. 45
REICHEINA 45
reiseri (Bembidion) 66
reiseri (Molops) 143, 144
reiseri (Pterostichus) 156, 159
reiseri corallipes (Molops) 142
reiseri holdhausi (Molops) 142, 143, 144, 145
reiseri pseudoalpestris (Molops) 142
reiseri reiseri (Bembidion) 65
reiseri reiseri (Molops) 142, 143
reiseri victoris (Molops) 142, 143, 144
reiseri vranense (Bembidion) 65
relictum (Bembidion) 55
relictum (Calosoma) 29
relictum var. pentheroides (Calosoma) 29
reticulatus (Apristus) 127
rhilensis (Pterostichus) 155
rhilensis kourili (Pterostichus) 156
rhilensis rhilensis (Pterostichus) 156
rhilensis vitosensis (Pterostichus) 156
rhodopense (Bembidion) 54
rhodopensis (Carabus) 34
rilvensis (Carabus) 32
riparia (Cicindela) 27
riparius (Pogonus) 78
rivulare (Bembidion) 58
robustus (Odontocarus) 97
rossii (Asaphidion) 53
rostratus (Stomis) 161
rostratus rostratus (Stomis) 161
rotundatus (Olisthopus) 137
rotundatus rotundatus (Olisthopus) 137
rotundicollis (Carterus) 95
rubripes (Harpalus) 99
rufescens (Nebria) 22
rufescens rufescens (Nebria) 22
rufibarbis (Harpalus) 112
rufibarbis (Ophonus) 112
ruficornis (Dyschirius) 50
rufipalpis (Harpalus) 100
rufipalpis rufipalpis (Harpalus) 100
rufipes (Harpalus) 108
rufipes (Molops) 146
rufipes (Notiophilus) 25
rufipes (Pseudophonus) 108
rufipes klisuranus (Molops) 146
rufipes munelensis (Molops) 146
rufipes pseudoosmanilis (Molops) 146
rufipes rufipes (Molops) 146
rufipes steindachneri (Molops) 147
rufipes sturanyi (Molops) 147
rufitarsis (Harpalus) 100
rufithorax (Apotomus) 51
rufomarginatus (Leistus) 18
rufus (Apotomus) 51
rufus (Platyderus) 166
rufus rufus (Platyderus) 166
rugosicollis (Myas) 148
rugosus (Carabus) 35
rumelicum (Bembidion) 63
rupicola (Harpalus) 112
rupicola (Ophonus) 112
sabulicola (Harpalus) 113
sabulicola (Ophonus) 113
sabulicola columbinus (Harpalus) 113
sabulicola columbinus (Ophonus) 113
sahlbergi albanica (Cicindela) 27
salina (Nebria) 23
salinus (Dyschirius) 49
salinus striatopunctatus (Dyschirius) 49
saphyrea (Amara) 177
sarajevoensis (Carabus) 31

241. 240 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
saxicola (Harpalus) 108
scabriusculus (Carabus) 37
scabriusculus bulgarus (Carabus) 37
Scallophorites 51
scapulare (Bembidion) 60
scapulare oblongum (Bembidion) 60
scapularis (Cymindis) 122
scapularis (Lebia) 126
scapularis scapularis (Cymindis) 122
scapularis scapularis (Lebia) 126
Scarites 50
SCARITINA 50
SCARITINAE 7, 12, 44, 187
SCARITINI 50
schaubergerianus (Ophonus) 112
schaumi (Panagaeus) 133
scrobiculatus (Platynus) 138
scrobiculatus serbicus (Platynus) 138
scrobiculatus turcicus (Platynus) 138
scutellaris (Tachys) 75
semigranosus (Cychrus) 37
semigranosus albanicus (Cychrus) 37, 38
semigranosus balcanicus (Cychrus) 37
semigranosus dinaricus (Cychrus) 37, 38
semigranosus jakupicensis (Cychrus) 38
semigranosus montanus (Cychrus) 37
semigranosus montenegrinus (Cychrus) 37, 38
semigranosus n. balcanicus (Cychrus) 37
semigranosus n. montenegrinus (Cychrus) 37
semigranosus semigranosus (Cychrus) 37
semigranosus travnikanus (Cychrus) 38
serbicus (Platynus) 138
serripes (Harpalus) 101
serripes ab. subchalybaeus (Harpalus) 101
serripes serripes (Harpalus) 101
sexpunctatum (Agonum) 135
sexstriata (Tachyura) 76
sexstriatus (Tachys) 76
sexstriatus var. bisbimaculatus (Tachys) 76
shardagensis (Carabus) 33
shljebensis (Carabus) 32
Siagona 44
SIAGONINAE 7, 44, 187
SIAGONINI 44
sichemita (Brachinus) 42
siculum (Bembidion) 64
siculum smyrnense (Bembidion) 64
sigma (Dromius) 125
sigma (Philorhizus) 125
signatipenne (Bembidion) 65
silphoides (Licinus) 130
similata (Amara) 177
similis (Ophonus) 110
simplex (Molops) 140, 141
simplex meridionalis (Molops) 140, 141
simplex simplex (Molops) 140
simulator (Dyschirius) 48
Sinechostictus 10, 73
Sinechostictus s.str. 73
skrimshiranus (Stenolophus) 120
skutariensis (Harpalus) 104
smaragdinus (Harpalus) 104
solitaris (Harpalus) 103
sordidum (Agonum) 135
sordidum sordidum (Agonum) 135
sordidus (Platynus) 135
spartanus (Molops) 140
spartanus parnassicola (Molops) 140
spartanus valonensis (Molops) 140
Speluncarius 12, 160
Speluncarius s.str. 160
Sphaerotachys 75
SPHODRINA 171
SPHODRINI 165
spinibarbis (Leistus) 17, 18
spinibarbis punctatus (Leistus) 18
spinibarbis rufipes (Leistus) 18
spinipes (Zabrus) 185
spinipes insignis (Zabrus) 185
splendidum (Bembidion) 58
splendidum pincum (Bembidion) 58
spoliatus (Chlaenius) 90
spoliatus spoliatus (Chlaenius) 90
spreta (Amara) 177
spurcaticornis (Anisodactylus) 94
steindachneri (Molops) 147
stenocephalus (Duvalius) 82
stenocephalus (Trechus) 81
Stenochoromus 147
STENOLOPHINA 12, 115
Stenolophus 119
Stenolophus s.str. 119

242. INDEX 241
sterbai (Carabus) 34
stictus (Ophonus) 113
stierlini (Asaphidion) 53
stolfai (Bembidion) 65, 66
Stomis 160
Stomis s.str. 160
strenuus (Argutor) 154
strenuus (Pterostichus) 154
strepens (Brachinus) 42
striatopunctatus (Poecilus) 150
striatul sic! (Bembidion) 66
striatum (Bembidion) 66
strumosus (Dyschirius) 46
strupii (Duvalius) 82
sturanyi (Duvalius) 81, 82
sturanyi (Molops) 147
sturanyi (Nebria) 19
sturanyi matejkai (Duvalius) 82
sturanyi munelensis (Molops) 146
sturanyi stenocephalus (Duvalius) 81, 82
sturanyi sturanyi (Duvalius) 82
subaeneus (Apristus) 127
subchalybaeus (Harpalus) 101
subcostatum (Bembidion) 68
subcostatum javurkovae (Bembidion) 68, 69
subcostatum spartanum (Bembidion) 69
subcostatum vau (Bembidion) 68, 69
subcylindricus (Harpalus) 102
subfasciatum (Bembidion) 71
submarginatus (Licinus) 129, 130
subnotatus (Trechus) 85
subnotatus ljubetensis (Trechus) 86
subnotatus subnotatus (Trechus) 85
subquadratus (Acinopus) 97, 98
subquadratus (Ophonus) 110
subrugosus (Carabus) 35
subsinuatus (Harpalus) 100
substriatum (Bembidion) 62
substriatus (Dyschiriodes) 50
substriatus (Dyschirius) 50
substriatus (Notiophilus) 25
substriatus priscus (Dyschirius) 50
sulphuripes (Harpalus) 100
sulphuripes sulphuripes (Harpalus) 100
sutomorensis (Carabus) 30
suturalis (Acupalpus) 117
suturalis (Neotrechus) 84
suturalis (Trechus) 84
suturalis pentheri (Neotrechus) 84
suturalis pretneri (Neotrechus) 83
suturalis suturalis (Acupalpus) 117
sycophanta (Calosoma) 29
sylvatica (Cicindela) 27
sylvatica sylvatica (Cicindela) 27
Syntomus 12, 127
Synuchidius 171
SYNUCHINA 173
Synuchus 173
Synuchus s.str. 173
TACHYINA 73
TACHYINI 10
Tachypus 36
Tachys 75
Tachyta 75
Tachyta s.str. 75
Tachyura 76
Tachyura s.str. 76
Taenidia 26
Talanes 71
Tapinopterus 12, 161
Tapinopterus s.str. 161
tardus (Harpalus) 103
tarsicus (Sinechostictus) 73
Tarulus 123
taurica (Amara) 179
tenebrioides (Zabrus) 185
tenebrioides tenebrioides (Zabrus) 185
tenebrosus (Harpalus) 98
tenebrosus var. centralis (Harpalus) 98
tenellum (Bembidion) 59
tenellum tenellum (Bembidion) 59
terminale (Bembidion) 71
terminatus (Chlaenius) 90
Terminophanes 71
terricola (Laemostenus) 173
terricola (Pristonychus) 173
terricola (Scarites) 51
terricola punctatus (Laemostenus) 173
terricola terricola (Scarites) 51
testaceum (Bembidion) 60
testaceum testaceum (Bembidion) 60
Testedium 71

243. 242 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV
tethys (Bembidion) 70
tetracolum (Bembidion) 69
tetracolum tetracolum (Bembidion) 69
teutonus (Acupalpus) 120
teutonus (Stenolophus) 120
Thalassophilus 88
thessalicus tomoricensis (Tapinopterus) 163
thoreyi (Agonum) 136
thoreyi (Platynus) 136
thoreyi thoreyi (Agonum) 136
tibiale (Bembidion) 54
tibialis (Amara) 177
Tomocarabus 36
tomoricensis (Amara) 179, 180
tomoricensis (Molops) 145
tomoricensis (Nebria) 21
Trachycarabus 37
transfuga (Molops) 143
transversalis (Cicindela) 27
transversalis transversalis (Cicindela) 27
TRECHINA 79
TRECHINAE 7, 52, 187
TRECHINI 79
TRECHINITAE 10
TRECHODINA 88
Trechus 85
Trechus s.str. 85
Trepanes 72
Trichochlaenius 92
Trichotichnus 12, 115
Trichotichnus s.str. 115
tricolor (Bembidion) 56
tricuspidata (Amara) 181
tricuspidatus (Carterus) 95
tricuspidatus (Ditomus) 95
trimaculata (Lebia) 126
triseriatus (Harpalus) 105
trisignata (Cylindera) 28
trisignata trisignata (Cylindera) 28
tristis (Chlaenius) 90
tristis (Harpalus) 104
tristis (Trechus) 87
tristis tristis (Chlaenius) 90
truncatellus (Metabletus) 128
truncatellus (Syntomus) 128
turcicus (Platynus) 138
uliginosus (Elaphrus) 38
unipustulatus (Badister) 128
ustulatum (Bembidion) 69
valbonensis (Trechus) 85
valonensis (Nebria) 23
vaporariorum (Cymindis) 123
varicolor (Bembidion) 56
varicolor varicolor (Bembidion) 56
variolosus (Carabus) 31
variolosus nodulosus (Carabus) 31
varium (Bembidion) 63
varium dyrrhachii (Bembidion) 63
velutinus (Chlaenius) 91
venustus (Laemostenus) 172
verbasci (Bradycellus) 118
vernalis (Harpalus) 102
vernalis cursor (Pterostichus) 151
versicolor (Poecilus) 151
vestitus (Chlaenius) 90
victoris (Molops) 143, 144
viduus moestus (Platynus) 133
viduus var. moestus (Platynus) 133
vignai (Duvalius) 82
violaceus (Carabus) 32
violaceus azurescens (Carabus) 32
violaceus azurescens m. merditana (Carabus) 33
violaceus azurescens m. merditanus (Carabus) 33
violaceus azurescens m. zabljakensis (Carabus) 32
violaceus azurescens n. zabljakensis (Carabus) 32
violaceus azurescens var. shljebensis (Carabus) 32
violaceus dryas (Carabus) 33
violaceus dryas var. korabensis (Carabus) 33
violaceus dryas var. merditanus (Carabus) 33
violaceus exasperatus (Carabus) 33
violaceus merditanus (Carabus) 33
violaceus shardagensis (Carabus) 33
violaceus var. korabensis (Carabus) 33
violaceus var. shardagensis (Carabus) 33
violaceus vlasuljensis (Carabus) 33
violaceus zabljakensis v. merditanus (Carabus) 33
viridicupreum (Agonum) 135
viridicupreum viridicupreum (Agonum) 135
viridicupreus (Platynus) 135
viridulus (Harpalus) 107
vivalis (Synuchus) 173
vivalis vivalis (Synuchus) 173

244. INDEX 243
vlasuljensis (Carabus) 33
vseteckai (Bembidion) 62
vseteckai dissimile (Bembidion) 62
walteri (Pterostichus) 156
weiratheri (Carabus) 30
weiratheri (Molops) 140
weiratherianus (Carabus) 35
wetterhallii (Masoreus) 92
wetterhallii wetterhallii (Masoreus) 92
winkleri (Carabus) 35
winkleri (Zabrus) 182
winklerianus (Molops) 144
Xenocelia 180
zabljakensis (Carabus) 32
ZABRINA 181
ZABRINI 174
Zabrus 181
Zabrus s.str. 185
Zezea 181
zhalovi (Duvalius) 82
ZUPHIINA 185
ZUPHIINI 185
Zuphium 186
Zuphium s.str. 186

245. 244 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV