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Morphine: opium poppy, pain relief        Make O2: photosynth.           Adaptations to living on land:          Origin of land plants (475-444 mya): first land   Paleozoic (500-250 mya): algae-          Endosymbiosis                               Secondary Endosymbiosis
Digitalin: foxglove; heart medication     Build soil : C buildup,        required multi cellular tissues for     plants, cuticle, spores, sporangia                >land plants: bryophytes (mosses)-       Multicellular green organisms               Second (or third, fourth)
Menthol: peppermint tree, cough           decomposition, rock            mechanical strength/support,            Silurian-Devonian explosion (444-359):            >early vascular plants: seedless         developed thru endosymbiosis: union         engulfing; Plastids evolve and
suppressant, relief of stuffy nose        weathering                     exposed light catching surfaces,        major morphological innovations: stomata,         plants (ferns, horsetails)->first seed   of a host and endosymbiont living           gene transfer occurs between
Taxol: pacific yew, ovarian cancer        Hold soil: root strength       anchoring system, water conduct         vasc. tissue, roots, leaves                       plants: gymnosperms (conifers)           within the host; division of chloroplasts   genomes and sometimes
Papain: papaya, reduce inflammation       and rmifcatIOn thru soil       system, obtaining nutrients, restrict   Carbinoferous: Lyoophytes and horsetails          Mesozoic (250-75): flowering plants:     closely resemble that of cyanobacteria      organisms
treat wounds                              Hold water                     water loss in desicating air, reprod    abundant (359-299): extensive coal-forming        angiosperms                              Process: anaerobic eukaryote engulfs        Mitochondria descended from
                                          Moderate climate:              and dispersing on land                  swamps                                            Cenozoic (75-0)                          aerobic bacerium; bac lives w/in euk        alpha-Proteobacteria
O2 & CO2                                  evapotranspiration,            Regulation of developmntal pthwys Gymnosperms abundant (299-145): both wet                                                         cell; euk supplies protection and C; bac    Chloroplasts  cyanobacteria
                                                                                                                                                                   (Shoot/Root) Apical meristem:
Conc                                      particle capture               -gene expression encoding transcr. and dry envir. blanketed with green plants for                                                  supplies ATP (used to produce pyruvate      (bluegreen algae)
                                                                                                                                                                   Cell divide, grow, differnte to form
                                          Food, fuel, fibre, drugs       Factors determine cell/tissue/organ first time                                                                                     & O2), handles toxic O2           Water Movement: along PE
                                                                                                                                                                   protodrm, grnd mristm,
                                                                         identity                                Angiosperms abundant (145-present):                                                                                          gradients (high ->low)
                                          Plasmodesmata: area                                                                                                      procambium
                                          betwn adj cell walls, filled   -cell fate det. By position (not clonal diversification of flowering plants                                                                                          Water PE: osmosis (tendency
                                          w/plasmalemma, comm            history)                                                                                                                                                             to move in response to diff.
                                          betwn cells, tubule of         -developmental pathways controlled                                                                                                                                   solute concentration), turgor
                                          smooth                         by networks of interacting genes                                                                                                                                     pressure (pressure of
                                          ER passes thru                 -development regulated by cell-to-                                                                                                                                   expanding cell volume against
                                                                         cell signaling:                                                                                                                                                      plasma membrane, cell wall
                                          Middle lamella: formed         •Ligand-induced signaling: cell wall                                                                                                                                 pushes back w/exqual opp.
                                          during cell div, outer cell    chemicals communicate local                                                                                                                                          force)
                                          wall, shared by cells          positional information                                                                                                                                               COHESION-TENSION THEORY
                                          Primary wall: forms after      •Hormonal signaling: auxin & others                                                                                                                                  (more accurate name:
                                          mid lamell, consists of        •Signaling via reg. proteins and/or                                                                                                                                  transpiration-cohesion-
                                          cellulose microfibrils and     mRNA through plasmodesmata                                                                                                                                           tension theory): water moves
                                          gel matrix of pectic                                                                                                                                                                                from soil/roots to leaves
        Carbon coal formation             compounds, hemicellulose,
*CO2 removed by plant synthesis and                                                                                                                                                                                                           along water pot. Grad. The
                                          glycoproteins                                                                                                                                                                                       grad. Exists bcuz water @ air-
limestone formation                       Secondary wall: formed                                                                                                                          WATER POTENTIAL: xylem sap rises against            water surf. In leaves is under
                                          after cell enlargement and                                                                                                                      gravity, driven by water pot grad.; 1 bar of press. neg. P (tens) great enough to
                                          provides compresn                                                                                                                               needed to push up a column of water 10m; 1          pull water up from roots thru
                                          Strength, made of cellulose,                                                                                                                    megapascal=10bars; pure water: 0 potential;         xylem)
                                          hemicellulose, lignin, often                                                                                                                    more P, more water potential; water under
                                          layered                                                                                                                                         tension decreases water potential; more solutes
                                          Roots meristem                                                                                                                                  decrease water pot.
                                                                                                                            Sclerenchyma: fibers, sclereids Collenchyma: cooking celery  Gradient water potential maintained by
                                          Roots force thru soil;
                                                                                                                            soft (break down collenchyma) Parenchyma: a) leaves:          creation and maintenance of gradient of
                                          Protection of apical
                                                                                                                            photosynthesis & gas exchange b) roots: carb storage          tension
                                          meristem; Delayed
                                          initiation of lateral                                                             Secondary Xylem: conducts/            H-BONDS AND
                                                                                                                            stores water & ions, provides         COHESION: O (weak       Water potential gradient between soil, plants,
                                          meristems; Diff reqs. for
                                                                                                                            support, made of parenchyma (rays) neg) + H (weak pos)       atmosphere, gain water from soil, lose to
                                          support and water
                                                                                                                            and sclerenchyma (fibers)             H-bond                  atmosphere: Water exits leaf thru stomata,
                                          collection/distribution
                                                                                                                            In Confiers/angiosperms: tracheids Cohesion: water            water replaced by evaporation from mesophyll
                                          Zea mays: junction                Secondary PHLOEM: sugar, amino acid,            (long, bordered pits that shut when molecule H-bonded to      cell lowering their water pot., causing them to
                                          between root apex and root        hormone transport up and down sieve tube,       tracheid collapses under low water four other water           extract water from neighbor cells, process
                                          cap                               comprised of vertical sieve tube members (no    pot.[torus moves and seals pit], can molecules, cohesion      connects back to tracheids/vessels causing
                                          Lateral Root development:         nucleus @ maturity and depend on bordering      be rehydrated, no cytoplasm), and     almost as stable as     water to be taken from xylem, water travels
                                          meristem develops from            companion cells to regulate physiological       ray cells (run horizontal thru xylem, covalent bond           from tracheids to air following water pot. grad. .
                                          parenchyma and lateral            processes) topped with sieve plates, support,   made of parenchyma and some           Adhesion: different     Cohesive and adhseive proerties of water and
Secondary Growth: Lateral Meristems       root grows out of root            made of parenchyma (sieve, comp. cells) and     tracheids)                            molecule types bond by small diameter of xylem aid in vertical
Stems and roots of woody plants           through cortex                    sclerenchyma (fibers/sclerids)                  Just angiosperms: vessels (short and similar process (e.g.    movement, pull decreases water pot. In xylem,
increase in diameter (apical mer. =                                                                                         wide, vessel elements stacked on      water-cellulose bond in roots take water from soil
length increase)                                                                                                            top of one another to form vessel,    xylem walls to          Corn, monocotyledon, C4                             Confier needle: broad, ONE
*vascular cambium: secondary vasc                                                                                           also contain pits, perforation walls, counteract gravity                                                          vascular bundle, mesophyll
tissues, xylem, phloem                                                                                                      moderate support but superior fluid enabling tension                                                              divided into palisade (top,
*cork cambium: bark tissues, periderm                                                                                       conduction)                           maintenance)                                                                spquished) and spongy
                                          Oldest xylem
(continually replaces epidermis)                                                                                                                                                                                                              (bottom, rounder) layers; slow
                                          cell on inside,
Periderm: cork (pretection),                                                                                                Diurnal pattern of shoot water pot.: During daylight, water                                                       water conduct, dry conditions;
                                          oldest phloem
phelloderm (synthesis & storage,                                                                                            loss exceeds water gain so shoot water potential decreases:                                                       stomata arranged in rows
                                          cells on outside
parenchyma), composed of AND                                                                                                                                                                                                                            STOMATA
derived from cork cambium (produces       Light->
cork and phelloderm, meristematic         Energy gradient ensures
cells)                                    down toward rxn center,
                                          and that transfer out of
Tomato, dicotyledon, C3                   peripheral antenna is
                                          unfavorable                                                                                                                                            Most veins run parallel to main vein but many
                                                                                                                                                                                                 cross veins so mesophyll cells close to a vein
                                                                                                                                                                                                                                                        Guard cells take up K causing
                                                                                                                                                                                                 Wind speed and increases transpiration (water          water into enter by osmosis,
                                                                                                                                                                                                 loss): wind reduces the boundary layer around          cells bow w/turgidity
                                                                                                                                                                                                 leaf which resists water from leaf, further wind       Isohydric: keeps leaf WP
                                                                                                                                                                                                 speed incr. reduces transp bcuz wind cools leaf        constant, maize, poplar
                                                                                                                                                                                                 directly                                               Anisohydric: leaf WP decr in
                                                                                                                                                                                                 Leaf plasticity w/light variation: sun leaves          day, sunflower, barley
                                                                                                                                                                                                 (than shade leaves)- have smaller area, 1.5-2.2        Carotenoids: tranfer photon
                                                                                                                                                                                                 mass/area, up to 1.5 density of stomata, more          energy to chlorophyll (light
                                          chloroplast                                                                                                                                            rubisco per chlorophyll, less chlorophyll per rxn      absorbed in photosynthesis),
                                          2 membranes:                                                                                                                                           center                                                 quench free rads by accepting
Veins arranged for support and so         endosymbiosis                              Excited e- in Photosynthsis                                                                                 Adaptation of xerophyte                                or stabilizing free electrons to
mesophyll cells close to a vein                                                      Fluorescence: e- drops back down to                                                                                                                                protect chlorophyll molecules;
                                                                                     lower E level, heat and fluorescence                                                                                                                               When a photo strikes, energy
Photosystem II (2H2OO2+4H+4e-): e- in rxn                                           emitted; Resonance: E in e- is moved to                                                                                                                            transferred to electron, e-
center excites chlorophyll, e- binds to                                              nearby pigment; Reduction/Oxidation:                                                                                                                               excited, raised to higher e-
pheophytin, chlorophyll oxidized, e- that reach                                      e- is transferred to a new compound                                                                                                                                shell w/greater PE
pheophytin are tranferred to plastoquinone           e- and H+ transf in thykaloid membr                                                                                                                                                                *too much light = too many
(lipid soluble), passed to an ETC (quinones and                                                                                    Dark Rxns occur in stroma                                                                                            free e- to dispense
cytochromes), passage of e- along ETC involves                           stroma                                                                                                                                                                         Chlorophyll absorb blue and
seriies of redox rxns that result im protons being
                                                                                                                                                                                                                                                        red light, transmit green,
pumped from stroma to thykaloid lumen, luman                                                                                                                                                     Incr. boundary layer, reduce water loss, reduce        carotenoids absorb blue and
pH reach 5 while strom pH is ~8 (H+ conc. 1000                                                                                                                                                   gas exchange (photosynth-->CO2)                        green, transmit yellow,
times higher in lumen than stroma), VERY fast,
                                                                                                                                                                                                                                                        orange, or red
rxn center than re-reduced by 2ndary donor                            STROMA                                                                                                                                                                            Pigments that absord blue/red
(usually water to produce O2) [END
                                                                                                                                                                                                                                                        best at photosynthesis, O2-
PHOTOSYSTEM II] e- then passed to protein
                                                                                                                                                                                                                                                        seeking bacteria congregate in
plastocyanin that can diffuse thru lumen of
                                                                                                                                                                                                                                                        these wavelengths bcuz alga is
thykaloid and donate e- to photosystem I
                                                                                                                                                                                                                                                        producing most O2
                                                                 Thykaloid
                                                     Protons diffuse to site of ATP synthase, black arrows rep. e-
                                                     transfer, blue lines rep proton movement                                      Thykaloid memb appear stacked but are folded and have                                                        C4 PHOTSYNTHESIS
                                                                                                                                   defined interior and exterior w/respect to stroma                                                            -the first product of CO2 fixation is
                                                        The Calvin Cycle:                                                                                                                                                                       malate (C40 in mesophyll (not PGA),
                                                        System stops is CO2,                                                                                                                                                                    this is transported to bundle sheath
                                                        ATP, or NADPH are                                                                                                                                                                       -CO2 released from malate in BS
                                                        not present                                                                                                                                                                             cells, fixed by rubsico and Calvin
                                                        -Rubisco-CO2 affinity                                                                                                                                                                   cycle proceeds, PEP back to
                                                        assures rapid carboxy-                                                                                                                                                                  mesophyll cells
                                                        lation at low C conc.                                                                                                                                                                   -decarboxylation of malate (CO2
                                                        -Rubisco will take O2                                                                                                                                                                   release) creates a higher conc of
                                                        rather than CO2 and                                                                                                                                                                     CO2 in BS cells than in
                                                        oxygenate RuBP                                                                                                                                                                          photosynthetic cells of C3 plants
                                                        (photorespiration)                                                                                                                                                                      This enables C4 plants to ssustain
                                                        -rubisco inefficient as                                                                                                                                                                 higher rates of photosynthesis, CO2
                                                        Catalyst for carboxyla-                                                                                                                                                                 conc higher than O2 in BS cells
                                                        Tion of RuBP                                                                                                                                                                            higher so photorespiration rates are
                                                        -CO2 competitive inhi-                                                                                                                                                                  lower
                                                        bition with O2

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Exam 3

  • 1. Morphine: opium poppy, pain relief Make O2: photosynth. Adaptations to living on land: Origin of land plants (475-444 mya): first land Paleozoic (500-250 mya): algae- Endosymbiosis Secondary Endosymbiosis Digitalin: foxglove; heart medication Build soil : C buildup, required multi cellular tissues for plants, cuticle, spores, sporangia >land plants: bryophytes (mosses)- Multicellular green organisms Second (or third, fourth) Menthol: peppermint tree, cough decomposition, rock mechanical strength/support, Silurian-Devonian explosion (444-359): >early vascular plants: seedless developed thru endosymbiosis: union engulfing; Plastids evolve and suppressant, relief of stuffy nose weathering exposed light catching surfaces, major morphological innovations: stomata, plants (ferns, horsetails)->first seed of a host and endosymbiont living gene transfer occurs between Taxol: pacific yew, ovarian cancer Hold soil: root strength anchoring system, water conduct vasc. tissue, roots, leaves plants: gymnosperms (conifers) within the host; division of chloroplasts genomes and sometimes Papain: papaya, reduce inflammation and rmifcatIOn thru soil system, obtaining nutrients, restrict Carbinoferous: Lyoophytes and horsetails Mesozoic (250-75): flowering plants: closely resemble that of cyanobacteria organisms treat wounds Hold water water loss in desicating air, reprod abundant (359-299): extensive coal-forming angiosperms Process: anaerobic eukaryote engulfs Mitochondria descended from Moderate climate: and dispersing on land swamps Cenozoic (75-0) aerobic bacerium; bac lives w/in euk alpha-Proteobacteria O2 & CO2 evapotranspiration, Regulation of developmntal pthwys Gymnosperms abundant (299-145): both wet cell; euk supplies protection and C; bac Chloroplasts  cyanobacteria (Shoot/Root) Apical meristem: Conc particle capture -gene expression encoding transcr. and dry envir. blanketed with green plants for supplies ATP (used to produce pyruvate (bluegreen algae) Cell divide, grow, differnte to form Food, fuel, fibre, drugs Factors determine cell/tissue/organ first time & O2), handles toxic O2 Water Movement: along PE protodrm, grnd mristm, identity Angiosperms abundant (145-present): gradients (high ->low) Plasmodesmata: area procambium betwn adj cell walls, filled -cell fate det. By position (not clonal diversification of flowering plants Water PE: osmosis (tendency w/plasmalemma, comm history) to move in response to diff. betwn cells, tubule of -developmental pathways controlled solute concentration), turgor smooth by networks of interacting genes pressure (pressure of ER passes thru -development regulated by cell-to- expanding cell volume against cell signaling: plasma membrane, cell wall Middle lamella: formed •Ligand-induced signaling: cell wall pushes back w/exqual opp. during cell div, outer cell chemicals communicate local force) wall, shared by cells positional information COHESION-TENSION THEORY Primary wall: forms after •Hormonal signaling: auxin & others (more accurate name: mid lamell, consists of •Signaling via reg. proteins and/or transpiration-cohesion- cellulose microfibrils and mRNA through plasmodesmata tension theory): water moves gel matrix of pectic from soil/roots to leaves Carbon coal formation compounds, hemicellulose, *CO2 removed by plant synthesis and along water pot. Grad. The glycoproteins grad. Exists bcuz water @ air- limestone formation Secondary wall: formed WATER POTENTIAL: xylem sap rises against water surf. In leaves is under after cell enlargement and gravity, driven by water pot grad.; 1 bar of press. neg. P (tens) great enough to provides compresn needed to push up a column of water 10m; 1 pull water up from roots thru Strength, made of cellulose, megapascal=10bars; pure water: 0 potential; xylem) hemicellulose, lignin, often more P, more water potential; water under layered tension decreases water potential; more solutes Roots meristem decrease water pot. Sclerenchyma: fibers, sclereids Collenchyma: cooking celery  Gradient water potential maintained by Roots force thru soil; soft (break down collenchyma) Parenchyma: a) leaves: creation and maintenance of gradient of Protection of apical photosynthesis & gas exchange b) roots: carb storage tension meristem; Delayed initiation of lateral Secondary Xylem: conducts/ H-BONDS AND stores water & ions, provides COHESION: O (weak Water potential gradient between soil, plants, meristems; Diff reqs. for support, made of parenchyma (rays) neg) + H (weak pos)  atmosphere, gain water from soil, lose to support and water and sclerenchyma (fibers) H-bond atmosphere: Water exits leaf thru stomata, collection/distribution In Confiers/angiosperms: tracheids Cohesion: water water replaced by evaporation from mesophyll Zea mays: junction Secondary PHLOEM: sugar, amino acid, (long, bordered pits that shut when molecule H-bonded to cell lowering their water pot., causing them to between root apex and root hormone transport up and down sieve tube, tracheid collapses under low water four other water extract water from neighbor cells, process cap comprised of vertical sieve tube members (no pot.[torus moves and seals pit], can molecules, cohesion connects back to tracheids/vessels causing Lateral Root development: nucleus @ maturity and depend on bordering be rehydrated, no cytoplasm), and almost as stable as water to be taken from xylem, water travels meristem develops from companion cells to regulate physiological ray cells (run horizontal thru xylem, covalent bond from tracheids to air following water pot. grad. . parenchyma and lateral processes) topped with sieve plates, support, made of parenchyma and some Adhesion: different Cohesive and adhseive proerties of water and Secondary Growth: Lateral Meristems root grows out of root made of parenchyma (sieve, comp. cells) and tracheids) molecule types bond by small diameter of xylem aid in vertical Stems and roots of woody plants through cortex sclerenchyma (fibers/sclerids) Just angiosperms: vessels (short and similar process (e.g. movement, pull decreases water pot. In xylem, increase in diameter (apical mer. = wide, vessel elements stacked on water-cellulose bond in roots take water from soil length increase) top of one another to form vessel, xylem walls to Corn, monocotyledon, C4 Confier needle: broad, ONE *vascular cambium: secondary vasc also contain pits, perforation walls, counteract gravity vascular bundle, mesophyll tissues, xylem, phloem moderate support but superior fluid enabling tension divided into palisade (top, *cork cambium: bark tissues, periderm conduction) maintenance) spquished) and spongy Oldest xylem (continually replaces epidermis) (bottom, rounder) layers; slow cell on inside, Periderm: cork (pretection), Diurnal pattern of shoot water pot.: During daylight, water water conduct, dry conditions; oldest phloem phelloderm (synthesis & storage, loss exceeds water gain so shoot water potential decreases: stomata arranged in rows cells on outside parenchyma), composed of AND STOMATA derived from cork cambium (produces Light-> cork and phelloderm, meristematic Energy gradient ensures cells) down toward rxn center, and that transfer out of Tomato, dicotyledon, C3 peripheral antenna is unfavorable Most veins run parallel to main vein but many cross veins so mesophyll cells close to a vein Guard cells take up K causing Wind speed and increases transpiration (water water into enter by osmosis, loss): wind reduces the boundary layer around cells bow w/turgidity leaf which resists water from leaf, further wind Isohydric: keeps leaf WP speed incr. reduces transp bcuz wind cools leaf constant, maize, poplar directly Anisohydric: leaf WP decr in Leaf plasticity w/light variation: sun leaves day, sunflower, barley (than shade leaves)- have smaller area, 1.5-2.2 Carotenoids: tranfer photon mass/area, up to 1.5 density of stomata, more energy to chlorophyll (light chloroplast rubisco per chlorophyll, less chlorophyll per rxn absorbed in photosynthesis), 2 membranes: center quench free rads by accepting Veins arranged for support and so endosymbiosis Excited e- in Photosynthsis Adaptation of xerophyte or stabilizing free electrons to mesophyll cells close to a vein Fluorescence: e- drops back down to protect chlorophyll molecules; lower E level, heat and fluorescence When a photo strikes, energy Photosystem II (2H2OO2+4H+4e-): e- in rxn emitted; Resonance: E in e- is moved to transferred to electron, e- center excites chlorophyll, e- binds to nearby pigment; Reduction/Oxidation: excited, raised to higher e- pheophytin, chlorophyll oxidized, e- that reach e- is transferred to a new compound shell w/greater PE pheophytin are tranferred to plastoquinone e- and H+ transf in thykaloid membr *too much light = too many (lipid soluble), passed to an ETC (quinones and Dark Rxns occur in stroma free e- to dispense cytochromes), passage of e- along ETC involves stroma Chlorophyll absorb blue and seriies of redox rxns that result im protons being red light, transmit green, pumped from stroma to thykaloid lumen, luman Incr. boundary layer, reduce water loss, reduce carotenoids absorb blue and pH reach 5 while strom pH is ~8 (H+ conc. 1000 gas exchange (photosynth-->CO2) green, transmit yellow, times higher in lumen than stroma), VERY fast, orange, or red rxn center than re-reduced by 2ndary donor STROMA Pigments that absord blue/red (usually water to produce O2) [END best at photosynthesis, O2- PHOTOSYSTEM II] e- then passed to protein seeking bacteria congregate in plastocyanin that can diffuse thru lumen of these wavelengths bcuz alga is thykaloid and donate e- to photosystem I producing most O2 Thykaloid Protons diffuse to site of ATP synthase, black arrows rep. e- transfer, blue lines rep proton movement Thykaloid memb appear stacked but are folded and have C4 PHOTSYNTHESIS defined interior and exterior w/respect to stroma -the first product of CO2 fixation is The Calvin Cycle: malate (C40 in mesophyll (not PGA), System stops is CO2, this is transported to bundle sheath ATP, or NADPH are -CO2 released from malate in BS not present cells, fixed by rubsico and Calvin -Rubisco-CO2 affinity cycle proceeds, PEP back to assures rapid carboxy- mesophyll cells lation at low C conc. -decarboxylation of malate (CO2 -Rubisco will take O2 release) creates a higher conc of rather than CO2 and CO2 in BS cells than in oxygenate RuBP photosynthetic cells of C3 plants (photorespiration) This enables C4 plants to ssustain -rubisco inefficient as higher rates of photosynthesis, CO2 Catalyst for carboxyla- conc higher than O2 in BS cells Tion of RuBP higher so photorespiration rates are -CO2 competitive inhi- lower bition with O2