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49 ren+®e enevold_non pollen palynomorphs
1. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Moesgaard Museum
&
Aarhus University, Denmark
Dept. of Geoscience
Dept. of Prehistoric Archaeology
NON POLLEN PALYNOMORPHS
Renée Enevold
- og hvad vi kan brug dem til
2. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Criteria:
All other micro-morphs than pollen in the samples
Morphological identifiable
Quantifiable
Size: 2-200 µm
NON POLLN PALYNOMORPHS
3. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Context for a NPP analysis NPP types pr. survey
Usually reported from pollen
preparations
Rough average: 10-30
NPPs
All sediment and soil types Increasingly more
Mostly Quaternary Fragments to whole
organisms
Archaeology, palaeolimnology,
palaeoecology, climate studies,
forensic surveys
New types are
continuously being
described
State of the art:
Most reported NPPs and their usual area of interpretation
Diatoms and dinocysts Climate studies
Fungal spores Fire regimes and anthropogenic
influence, archaeology
Testate amoeba Peat ecology, palaeoecology
Green algae and
cyanobacteria
Palaeolimnology and climate
studies
5. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
FUNGI
Mutualistic and parasitic fungi
Saprotrophic and coprophilic fungi
Suggested reading:
VAN GEEL, B., 2006. Fossil ascomycetes in Quaternary deposits. Nova Hedwigia, 82 (3), pp. 313; 313-329; 329.
8. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
ZOOLOGICAL PARTS
Photo : Renée Enevold
Photo : Renée Enevold
Suggested reading:
KVAVADZE, E. and KAKHIANI, K., 2010. Palynology of the Paravani burial
mound (Early Bronze Age, Georgia). Vegetation History and Archaeobotany,
19(5), pp. 469-478.
10. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Suggested reading:
Brinkkemper, O. & van Haaster, H. 2012, "Eggs of intestinal parasites whipworm (Trichuris) and mawworm (Ascaris): Non-pollen palynomorphs in archaeological
samples", Review of palaeobotany and palynology, vol. 186, no. 0, pp. 16-21.
PARASITE EGGS
Photo : Renée EnevoldPhoto : Renée Enevold
14. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
TESTING THE METHOD TO THE LIMIT
3. Criteria
All other micromorphs than
pollen in the samples -
cataloguing every potential
palynomorph
2. Preparation
Minimizing bias in the preparation
procedure – aiming at the highest
NPP diversity
1. Sampling
Approaching
representative
subsamples
4. Identifications
Contact with experts for all
groups
15. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
• 28 samples
• 6000 years of sediment
• Intense anthropogenic
disturbance approx. 800 years
• Sediment analysis
• Pollen analysis
• Macrofossil analysis
• NPP analysis
TÅRUP LUND
FOREST HOLLOW
Iron Age field system approx. 500 BC – AD 300
17. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Mild preparation procedure applied
Everything describable counted
Minimum count 300
Everything photographed
NPP ANALYSIS OF TÅRUP LUND SAMPLES
0
5
10
15
20
25
30
35
40
45
50
55
60
65
70
75
80
85
90
95
100
105
110
115
120
125
Depth
Sp
ore
1
20 40 6 0
Spor
e
19
S
pore
29
Sp
ore
30
Sp
ore
31
Spor
e
32(s
po
ro
rm
iella
)
S
pore
3
4
Sp
ore
36
Spor
e
43(s
ma
llpod
o
sp
ora)
S
pore
4
5
S
pore
5
3(+
sp
ore
28
2)
20
Sp
ore
63Sp
ore
64
Spor
e
66
S
pore
6
8
Sp
ore
69
Spor
e
71
Spor
e
73
S
pore
7
6
Sp
ore
83
Spor
e
85(p
od
osp
o
ra)
Spore
8
9
Spore
9
2
S
pore
95
Sp
ore
100
2 0
Spore
1
01S
pore
1
06
Sp
ore
108
Sp
ore
109
Spor
e
110
(+
sp
or
e
65)
Spore
1
12
S
pore
114
Sp
ore
116
Spore
1
25
S
pore
128
S
pore
129
Sp
ore
133
Spor
e
1
38
S
pore
140
Sp
ore
141
Spor
e
143
Spor
e
1
44
S
pore
1
49
Sp
ore
151
Spor
e
152
S
pore
1
56
S
pore
1
57
Sp
ore
158
(+
macr
o
9)
Spor
e
159
S
pore
1
64
Sp
ore
166
Sp
ore
167
20
Spor
e
170
("sigmo
po
lis
")
Spor
e
172
S
pore
1
75
(+
sp
ore
7
7)
S
pore
176
Sp
ore
181
Spor
e
182
S
pore
1
84
2 0
Sp
ore
197Sp
ore
200
Sp
ore
202
Spor
e
2
08
(+
m
acro
46
,50)
Hypo
podia
o
fGaeum
anom
yc
es
S
pore
2
10
20 4 0 6 0
Sp
ore
214
Spor
e
215
S
pore
2
17
S
pore
218
Sp
ore
219
Spor
e
220
(sm
allsp
o
rorm
iella
)
Spore
2
21
S
pore
222
20
Sp
ore
223
("gel
asino
spo
ra
")
Sp
ore
224
2 0
Spor
e
225
(+
sp
or
e
222
)
Spor
e
226
(kre
tzsch
m
aria
,spo
re
13
2)
Spore
2
30
S
pore
233
Sp
ore
234
Sp
ore
240
Spore
2
43
S
pore
245
Sp
ore
253
Spor
e
2
63
S
pore
265
S
pore
269
Sp
ore
280
Spor
e
281
S
pore
2
84
Sp
ore
290
Spor
e
291
Spor
e
292
S
pore
2
93
Sp
ore
294
20
Spor
e
295Spor
e
296
S
pore
2
97
Sp
ore
298
(+
sp
ore
180
)
Sp
ore
299
Spor
e
300
2 0
S
pore
3
01S
pore
3
02
Sp
ore
303
Spor
e
304
S
pore
3
06
S
pore
3
07
20 4 0
Sp
ore
308
2 0
S
pore
309Sp
ore
310
Spor
e
312
Spor
e
313
(+
sp
or
e
154
)
S
pore
3
14
Sp
ore
315
Spor
e
316
Spore
3
18
(+
sp
ore
2
78
)
S
pore
3
19
Sp
ore
320
Spor
e
321
Spore
3
22
S
pore
323
(+
oo
cy
te
24)
Sp
ore
324
Sp
ore
325
Spore
3
26
S
pore
327
Sp
ore
328
Spor
e
3
29
("
sigm
opolis
")
S
pore
330
S
pore
331
Sp
ore
332
Spor
e
333
S
pore
3
34
Sp
ore
335
Sp
ore
336
Spor
e
337
S
pore
3
38
Sp
ore
339
Spor
e
340
Spore
3
41
S
pore
3
42
Sp
ore
343
Spor
e
344
Spore
3
45
(+
sp
ore
3
05
)
S
pore
346
Sp
ore
347
Sp
ore
348
Spore
3
49
M
akro
1
(F
ungalfrui
tbod
y)
M
akro
2M
ak
ro
5
M
ak
ro
7M
akro
11
M
akro
16
M
ak
ro
17
M
akro
2
0
M
akro
21
M
akro
26
M
ak
ro
28(+
spo
re
13
9)
M
akro
3
0(s
mall-
pitte
txy
lem
)
M
akro
32
M
ak
ro
34
M
akro
3
6
M
akro
4
1
M
akro
42
M
ak
ro
47
2 0
M
akro
4
8(+
m
ac
ro
62,65
)
M
ak
ro
49(+
m
ac
ro
54)
M
akro
5
1(larg
e-p
itte
txyl
em)
M
akro
52
M
ak
ro
53(m
acro
86
)
M
ak
ro
56
M
akro
5
7
M
akro
60
M
ak
ro
61
M
akro
6
4
M
akro
66
M
akro
69
(+
m
ac
ro
7
7,90)
M
ak
ro
70
M
akro
7
1(H
dV-1
0
64
)
M
akro
72
M
akro
73(+
m
ac
ro
76
)
M
akro
74(+
m
ac
ro
79
,In
sec
to
rpla
nt20)
M
akro
7
5
M
akro
80
(m
a
cro
78
)
M
akro
81
M
ak
ro
8
2(m
acro
97
)
M
akro
83
M
akro
84
(c
onid
ie
sp
o
re)
M
akro
85
M
ak
ro
87
M
akro
8
8
M
akro
89
M
ak
ro
92
M
ak
ro
93
M
akro
9
4
M
akro
95
M
ak
ro
96
M
akro
9
8
M
akro
9
9
M
akro
10
0
20
Scla
riform
perf
ora
tio
n
pl
ate
Ph
ytolite
2
Bry
op
hyta
1
(sph
agn
um)
P
te
rid
oph
yta
1(p
o
lypodium
)
Pte
rid
o
ph
yt
a
4(d
ryop
teris
)
Pte
rid
oph
yta
5
(p
teridiu
m
)
T
rilete
s
po
re
un
iff.
A
lgae
1Algae
2Alga
e
3(m
acro
5
5)
A
lgae
4Algae
5(+
alg
ae1
1)
Algae
7(a
lga
e6,
meu
ge
o
tia
)
Alga
e
8(g
leotr
ichia
)
Algae
9A
lgae
10
Bo
tryo
co
cc
us1
Pedia
stru
m
1
Pedia
stru
m
9
P
ediast
ru
m
und
iff.
Sp
irog
yra
1
Spirogyr
a
2(o
rn
am
e
nted,o
oc
yte
17
)
D
ia
tom
3
Diato
m
6
Diatom
8
2 0
Diatom
undiff.
Anab
ena
1
R
ivo
lari
a
type
20
Sp
on
g
espi
cule
1
20
Spon
g
espi
cule
2
Spon
g
espi
cule
3(m
ac
ro
67)
F
oram
in
ife
ra
llin
ing
Oo
cyte
5
Oo
cyte
6
Ooc
yte7
Oocyt
e9
O
ocyte
10
Oo
cyte1
2
Oocyt
e13
Oocyt
e14
(S
po
re
18
3)
O
ocyte
15
Oo
cyte1
6
Ooc
yte19
O
ocyt
e20
(pa
ra
site
egg
5)
O
ocyte
21
(sp
ore
216,filin
ia
)
Oo
cyte2
2
Ooc
yte25
O
ocyt
e26
Oo
cyte
27
Ooc
yteun
diff.
Am
oe
ba5
(b
ottle
)
A
m
oe
ba5
Ha
ir1Hair2H
air3Ha
ir4Fir1
Fir2
In
se
ct
o
rp
lant1
In
se
ct
orplant7
In
se
ct
orplan
t9(+
macr
o
68)
In
se
ct
orplant
10
(+
fir3
)
In
se
ct
o
rp
lant12
In
se
ct
orplant13
In
se
ct
orplan
t15
(+
14
)
In
se
ct
o
rp
lant
16
In
se
ct
orplant17
In
se
ct
orplant18
In
se
ct
orplan
t21
In
se
ct
o
rp
lant
22
In
se
ct
orplant23
In
se
ct
orplan
t24
(+
11
)
In
se
ct
orp
lant
25
In
se
ct
antenna
1
In
se
ct
a
ntenna
5
In
se
ct
ant
enn
a
6
In
se
ct
ante
nna
7
In
se
ct
a
ntenna
8
In
se
ct
ant
enn
a
9
Claw
4
(h
air2)
In
se
ct
leg
1
In
se
ct
m
outh
parts
(m
ite
)
P
arasite
egg
3(S
po
re
146)
Fibre
3
Fibre
4Stjern
ehå
r1
T
rich
os
clerid
1
20 4 0 6 0
Fu
ng
a
ls
po
re
s
2 0 40 60 8 0
P
lant
pa
rts
2 0 40
A
lgae
2 0 40
Oth
eraquati
corg
anism
s
20 40
Art
hrop
oda
re
m
ainsO
th
er
18. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
Spores Plant
1, 5
3, 5
5, 56, 5
9, 5
12, 514, 5
16, 5
33, 5
51, 5
55, 5
58, 5
64, 5
70, 575, 5
79, 5
84, 5
87, 5
90, 5
94, 5
97, 5
101,5
104,5
109,5
119,5
124,5
CalAgeBP
20 40 60
Copro
philous
fu
ngal
sp
ores
Spore
3
2
S
pore
73
Spore
220
Sp
ore
43
Spore
8
5
Spor
e
290
S
pore
34
Spor
e
221
S
pore
294
Spor
e
3
12
S
pore
344
Spor
e
349
Sp
ore
138
Spore
2
18
S
pore
269
Spore
299
20
Sp
ore
309S
pore
328
Sp
ore
334
S
pore
348
Spor
e
8
9
S
pore
36
(C
h
aetom
iu
m)
2 0
Spore
2
23
(G
elas
inosp
ora
)
Sp
ore
292
Spore
2
93
20
Spor
e
295Spore
3
00
2 0
Spor
e
301S
pore
304
Spor
e
3
06
S
pore
307
Spore
3
15
Sp
ore
324
Spor
e
2
84
Sp
ore
337
Spore
92
Sp
ore
129
S
pore
202
Spor
e
226
(K
retz
sc
hm
aria
d
eu
sta)
S
pore
322
Spore
3
02
S
pore
303
Spor
e
2
22
S
pore
224
2 0
Spore
2
25
2 0
Sp
ore
101
2 0 4 0 60
Spore
214
2 0 4 0
Spore
3
08
20 40 60
Sp
ore
19
20
S
pore
63Spor
e
6
9
S
pore
166
Spor
e
110
Sp
ore
76
Spore
2
81
S
pore
318
Spor
e
3
19
Sp
ore
320
Spore
321
Sp
ore
116
S
pore
109
20
Spor
e
170
Sp
ore
291
Spor
e
1
76
S
pore
208
Spor
e
9
5
Sp
ore
141
Spore
1
43
Sp
ore
296
S
pore
83
Sp
ore
340
Spore
265
Spor
e
3
41
S
pore
336
Spor
e
297
Sp
ore
243
Spor
e
1
12
S
pore
140
Spore
68
Sp
ore
347
Spore
2
30
Spor
e
298
S
pore
345
Sp
ore
31
S
pore
100
Spor
e
3
0
S
pore
234
Spore
3
39
Sp
ore
133
Spore
2
00
S
pore
338
Spore
310
20
Sp
ore
197S
pore
325
Sp
ore
327
S
pore
323
Spor
e
1
59
S
pore
157
Spore
7
1
Sp
ore
215
Spor
e
2
17
Sp
ore
181
Spore
108
Sp
ore
53
S
pore
158
Spor
e
106
S
pore
164
Sp
ore
329
S
pore
233
Spor
e
3
30
S
pore
332
Spore
3
33
Sp
ore
313
Spor
e
1
52
Sp
ore
331
Spore
175
Sp
ore
314
S
pore
253
Spor
e
335
S
pore
167
Spore
2
10
S
pore
280
Spor
e
2
63
Sp
ore
45
Spore
1
14
2 0 4 0
Plantrem
a
ins
M
akro
1M
ak
ro
2
M
akro
5M
ak
ro
7M
akro
11
M
ak
ro
1
6
M
akro
17
M
akro
2
0
M
akro
21
M
akro
26
M
akro
28
M
akro
30
M
ak
ro
3
2
M
akro
34
M
ak
ro
36
M
akro
41
M
akro
4
2
M
akro
47
2 0
M
akro
48
M
akro
49
M
akro
51
M
ak
ro
52
M
akro
53
M
ak
ro
56
M
akro
57
M
ak
ro
6
0
M
akro
61
M
akro
64
M
akro
66
M
aro
69M
akro
70
M
akro
71
M
akro
72
M
akro
73
M
ak
ro
7
4
M
akro
75
M
ak
ro
80
M
akro
81
M
ak
ro
8
2
M
akro
83
M
akro
84
M
akro
85
M
akro
8
7
M
ak
ro
88
M
akro
89
M
akro
92
M
akro
93
M
ak
ro
9
4
M
akro
95
M
akro
9
6
M
akro
98
M
ak
ro
9
9
M
akro
100
2 0
Sclari
form
p
erfora
tio
n
plate
Ph
ytolith
2
B
ry
op
h
yta1
Pte
rid
oph
yta
1
P
te
rid
oph
yta
4
Pte
rid
oph
yta
5
T
rilete
spore
un
iff.
Stjer
nehå
r1
Tri
ch
os
cl
erid
1
Algae
1Algae
2Algae
3Alga
e
4Algae
5Algae
7A
lgae
8Alga
e
9
A
lgae
10Botryo
co
cc
us
1
P
ediast
ru
m
1
Pedi
astru
m
9
Pe
diastru
m
und
iff.
Spirogyra
1
Sp
iro
gyra
2
D
ia
tom
3
Diatom
6
Dia
tom
8
2 0
Diatom
undiff
.
20
S
pon
ges
picule
1
2 0
Spon
ge
spicul
e
2
S
pon
gespi
cule
3
Anab
ena
1
Rivo
laria
typ
e
Oocyt
e5
Oo
cyte6O
ocyte
20
Ooc
yteun
diff.
Oocyt
e22
Ooc
yte25
O
ocyte
26
Ooc
yte27
Am
oe
ba
5
Oocyt
e9
Oo
cyte7Oocyt
e10
Oo
cyte1
2
Oocyt
e13
Ooc
yte14
O
ocyte
15
Ooc
yte16
Oo
cyte1
9
Ooc
yte21
H
air1
Hair2Ha
ir3Hair4
Fir1
F
ir2
In
se
ct
orplant
7
In
se
ct
orplan
t9
In
se
ct
o
rp
lant10
In
se
ct
orplan
t12
In
se
ct
orp
lant
13
In
se
ct
orplan
t15
In
se
ct
o
rp
lant17
In
se
ct
orplan
t18
In
se
ct
orpla
nt22
In
se
ct
orplant
23
In
se
ct
o
rpla
nt24
In
se
ct
orp
lant25
In
se
ct
ante
nn
a
1
In
se
ct
ante
nn
a
5
In
se
ct
ante
nn
a
6
In
se
ct
a
nte
nn
a
7
In
se
ct
ante
nn
a
8
Claw
4(h
air2
)
In
se
ct
leg
1
In
se
ct
m
outhp
arts
1
Pa
rasite
egg
3
Fibre
3Fibre
4
20
A
lgae
2 0 4 0
Oth
eraqua
ti
corga
nisms
20
A
rthrop
odre
mains
Sporor miella Podospor a t ype Sor daria ty pe Cer cophor a t ype Sor dar iales Pyrenomycet es
Lept ospheria gr oup
Xylariaceae Plant rem ains Algae Zoological rem ainsO t her aquat ic or ganisms
Algal Zoological
• NPP types after reduction: 260
• Spores, mainly fungal: 149
• Plant remains: 53
• Algal remains: 25
• Zoological remains and resting eggs: 33
Reduced diagram
21. DEPARTMENT OF GEOSCIENCE
AARHUS
UNIVERSITETAU
Renée Enevold
ICP-IOCP 2016
Salvador
MODERN ANALOGUES
i. Forest hollow with closed canopy (Moesgaard Skov)
ii. Dried out forest hollow with closed canopy (Langland)
iii. Alder carr (Moesgaard Skov)
iv. Small lake surrounded by acidic common and pasture (Helgenæs)
v. Small lake in a mixed deciduous forest (Hennetved Haver)
vi. Small lake in a Hazel coppice (Næs Skov)
vii. Semi-dried out pond with willow thicket and surrounded by agricultural fields (Mols)
viii. Small dried out pond immediately surrounded by agricultural fields (Mols)
ix. Small lake immediately surrounded by agricultural land (Mols)
x. Small lake with mixed thicket and surrounded by agricultural fields (Mols)