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VEGETABLES AND FRUITS:
CLASSIFICATION, STRUCTURE,
COMPOSITION AND NUTRITIVE VALUE
Aktarafun Zannat
Assistant Professor (Food and Nutrition)
Govt. College of Applied Human Science, Dhaka
CLASSIFICATION
• Vegetables:
• Vegetables are classified according to the
parts of the plants consumed or colour of the
vegetable or according to the nutritive values.
Nutritionally they are classified into 3 groups-
1. Green leafy vegetables
2. Roots and Tubers
3. Other vegetables
Botanical Classification- B Srilakshmi book (page-
197)
Fruits: B Srilakshmi book (page- 227)
Structure
(a) Cell Wall System
1. Primary Wall
• The outermost living boundary of the cell. Composed mainly of cellulose,
hemicellulose, and pectic substances. It is thin, flexible, and permeable, allowing
exchange of water and solutes. Provides shape, rigidity, and protection to the cell.
2. Middle Lamella
• The intercellular layer that cements adjacent plant cells together. Composed largely of
pectin compounds. Responsible for firmness and cohesion of fruits and vegetables.
During ripening or cooking, pectin becomes soluble, causing the tissue to soften.
3. Intercellular Space
• Small air-filled gaps between cells. Influences texture, crispness, and light reflection,
affecting appearance. The amount of intercellular space differs among fruits and
vegetables.
(b) Plasmalemma (Cell Membrane)
• A semi-permeable membrane located inside the cell wall. Controls the movement of
water, ions, and solutes in and out of the cytoplasm. Maintains the integrity of the
protoplast.
(c) Protoplasmic Components
• The protoplasm includes all living parts of the cell, enclosed by the plasmalemma.
 Cytoplasm:
• Jelly-like substance containing organelles such as the nucleus and plastids. Site of
metabolic activities and enzyme reactions.
 Nucleus:
• The control center of the cell. Contains the nucleolus and chromatin material (DNA).
Directs cell metabolism, growth, and reproduction.
 Plastids:
• Specialized organelles responsible for color and storage. Three main types:
Chloroplasts (photosynthesis), Chromoplasts (pigments), Leucoplasts (storage).
 Vacuole:
• Large central cavity filled with cell sap (water, sugars, salts, acids, pigments).
Surrounded by the tonoplast. Maintains turgor pressure, keeping tissues firm and crisp.
 Tonoplast:
• Membrane surrounding the vacuole. Regulates the exchange of substances between
vacuole and cytoplasm.
 Starch Grains:
• Found mainly in leucoplasts (amyloplasts). Serve as storage form of carbohydrates.
Quantity and size vary with type of fruit or vegetable.
FUNCTIONAL IMPORTANCE OF CELL
STRUCTURE
a) Mechanical and Structural Functions
• The cell wall and middle lamella provide mechanical strength and rigidity. These
features maintain shape and resist deformation during handling and cooking. The
firmness of tissues depends on pectin structure and cell turgor.
(b) Physiological and Sensory Functions
• Vacuole and cytoplasm contain organic acids, sugars, and pigments responsible for
flavor and color. Air spaces between cells influence crispness and appearance. Loss of
turgor pressure results in softening or wilting. Texture changes during cooking occur due
to pectin solubilization and cell wall breakdown.
(c) Nutritional Importance
• Cell wall polysaccharides (cellulose, hemicellulose, and pectic substances) form
dietary fiber. Plastids store essential nutrients such as starch and pigments (carotenoids,
chlorophyll). Vacuole stores soluble nutrients like vitamin C and organic acids.
COMPARISON WITH ANIMAL CELLS
•Plant parenchyma cells differ from animal cells in several ways:
-Have rigid cell walls made of cellulose.
-Contain vacuoles and plastids.
-Possess a middle lamella for adhesion, absent in animal
cells.
•These structures provide firmness, storage capacity, and
turgidity, all of which influence the edible quality of fruits and
vegetables.
COMPOSITION AND NUTRITIVE
VALUES OF VEGETABLES
1. Nutritive Value of Green Leafy Vegetables
• Green leafy vegetables are the manufacturing organs of plants where
photosynthesis takes place.
• They are low in carbohydrates and energy but are good sources of β-
carotene, calcium, riboflavin, folic acid, and ascorbic acid (Vitamin C).
• The nutritive value depends on the variety and freshness of the leaves.
Vitamin and Mineral Content
• Rich sources of vitamins and minerals, especially B-vitamins (riboflavin, folic acid), β-
carotene, and Vitamin C.
• β-carotenes act as antioxidants and precursors of Vitamin A.
• The greener the leaves, the higher the carotene content due to more chlorophyll
pigments.
• Drying and withering reduce the content of Vitamin C and riboflavin.
Iron and Calcium Content
• Rich in iron — help in preventing anaemia.
• Amaranth, fenugreek, gogu are good iron sources.
• Agathi, colocasia, drumstick, and fenugreek leaves contribute calcium, though
absorption is limited by oxalic acid.
Other Nutrients and Fibre
• High moisture content; therefore, they wither easily and need proper
preservation.
• Not good sources of protein, fat, or carbohydrates.
• Excellent sources of fibre, aiding digestion and preventing degenerative
diseases.
• Discarded leaves like cauliflower leaves, beetroot leaves, and pumpkin
leaves also contain iron and should be used.
2. NUTRITIVE VALUE OF ROOTS AND TUBERS
Energy and Starch
• Provide more calories than leafy vegetables because they contain more starch.
• Important energy-giving foods.
Carotene and Vitamin C
• Carrots are rich in carotenes.
• Roots and tubers are fairly good sources of Vitamin C.
• Deficiency in Other Nutrients
• Poor sources of calcium, iron, and B-vitamins.
• Also low in protein.
3. NUTRITIVE VALUE OF OTHER VEGETABLES
General Composition
• Contain high moisture, making them highly perishable.
• Contribute fibre to the diet.
Vitamins and Pigments
• Tomatoes are rich in carotenoids (especially lycopene).
• Cluster beans, capsicum (green, red, yellow) — good sources of iron,
carotenoids, and Vitamin C.
• Capsicum contributes to both colour and Vitamin C content.
Antioxidant Activity of Vegetables
• Antioxidants protect cells against oxidative damage caused by free radicals.
• Measured by DPPH (2,2’-Diphenyl-1-picryl hydrazyl) radical scavenging activity.
• Okra and broad beans have the highest antioxidant activity among other vegetables.
Beetroot shows the highest antioxidant activity among roots and tubers.
Pigments in Vegetables
• Classification
• Pigments give colour and visual appeal to vegetables.
• Classified into:
• Water-insoluble pigments: Chlorophyll and Carotenoids
• Water-soluble pigments: Anthocyanins, Anthoxanthins, and Betalains
WATER-INSOLUBLE PIGMENTS
Chlorophyll
• Found in chloroplasts; responsible for green colour.
• Two forms:
• Chlorophyll-a: blue-green
• Chlorophyll-b: yellow-green
• Present in a 3 : 1 ratio in most plants.
• Masked by other pigments in some vegetables.
• Present in leafy vegetables, capsicum, beans, peas, and chillies.
Carotenoids
• Fat-soluble pigments: yellow, orange, red in colour.
• Include β-carotene, α-carotene, γ-carotene, lycopene, xanthophyll,
cryptoxanthin.
• Function as antioxidants and precursors of Vitamin A.
WATER-SOLUBLE PIGMENTS
Anthocyanins
• Water-soluble red to purple pigments; derivatives of cyanidin.
• Found in cherries, red apples, berries, grapes, pomegranates, and currants.
• Responsible for red colour in radish skin, sweet potato, red cabbage, and
purple brinjal.
• Sensitive to pH and heat; colour fades on cooking.
Anthocyanidins
• Anthocyanins without sugar.
• Examples:
• Pelargonidin – red
• Cyanidin – reddish blue
• Delphinidin – blue
Anthoxanthins
• Colourless to yellow pigments related to anthocyanins.
• Present in onions, spinach, and other pale-coloured vegetables.
• Sensitive to pH – become darker in acidic medium.
• Subgroups include flavones, flavonols, flavanones, catechins, etc.
Betalains
• Found in certain families instead of anthocyanins.
• Two pigments: Betacyanin (red-violet) and Betaxanthin (yellow).
• Present in beetroot.
ORGANIC ACIDS
•Vegetables contain formic, oxalic, malic, citric, tartaric, and
succinic acids.
•Give flavour and slightly acidic pH (5.0–6.3).
•Tomatoes and grapes have low pH due to high organic acid
content.
ENZYMES IN VEGETABLES
• Act as catalysts in biochemical reactions.
• Two major groups: Hydrolytic enzymes and Oxido-reductases.
 Functions
• Responsible for ripening (e.g., in tomatoes and bananas).
• Cause browning in potato, brinjal, plantain.
• Papain in papaya: used in tenderising meat and stabilising beer.
• Can cause loss of anthocyanin pigments (by anthocyanase).
• Heat treatment inactivates undesirable enzymes.
FLAVOUR COMPOUNDS
• Result from sugars, acids, salts, aldehydes, ketones, alcohols, tannins, and
sulphur compounds.
• Sweetness mainly from glucose, fructose, galactose.
• Astringency from tannins and phenolics.
Sulphur-Containing Compounds
• Found in onion, garlic, leek (Allium group) and cruciferous vegetables
(cabbage, turnip, cauliflower, kale, mustard).
• Responsible for characteristic odour and strong flavour.
• Overcooking or improper cooking enhances sulphur odour.
BITTER COMPOUNDS
• Cucurbitacins – found in cucumber and gourds, cause bitterness.
• Solanine and chaconine – steroidal glycoalkaloids in potatoes; toxic if
sprouted or damaged.
• Cooking destroys most toxic alkaloids.
• Proper storage reduces sprouting and bitterness.
Addition of Chemicals
• Preservative Treatment
• Studies at Agriculture College and Research Institute, Madurai:
• Onions treated with maleic hydrazide (2000 ppm) inhibited rooting and
sprouting.
• Prolonged storage life up to 150 days.
Factors That Extend Shelf Life
•Harvesting at optimum maturity with minimum injury.
•Maintaining proper sanitation during handling and storage.
•Optimum storage conditions: proper temperature, humidity, and sanitation.
•Controlled atmosphere during all marketing and transport steps.
COMPOSITION AND NUTRITIVE
VALUE OF FRUITS
• Fruits are generally poor sources of protein and fat.
• Avocado is an exception, containing 13.8 % fat.
• ω-3 fatty acids are present in plum, papaya, and guava — these fruits contribute to
dietary ω-3 fatty-acid intake (NIN 1999–2000).
• Contain high moisture, hence are highly perishable.
• Good sources of fibre aiding digestion.
Caloric Value
• Fruits are not high in calories.
• Bananas give a fairly good amount of energy compared to most fruits.
• Ripe fruits have a higher percentage of sugars than unripe ones.
• Major sugars: sucrose, fructose, and glucose.
Mineral Content
• Generally, fruits are poor in iron, but dry fruits are a fairly good source of iron.
VITAMINS AND CAROTENOIDS
β-Carotene and Carotenoids
• Mangoes are excellent sources of carotenes.
• Research at NIN, Hyderabad found Alphonso richest in β-carotene.
• Banganapally and Pedda Rasalu are fairly good sources.
• Indian dates, papaya, and oranges also contain β-carotene.
• Carotenoids provide yellow-orange colour and act as vitamin A precursors.
Vitamin C (Ascorbic Acid)
• Guavas and citrus fruits are excellent sources of vitamin C.
• Cashew apple is inexpensive yet rich in vitamin C (≈ 180 mg/100 g).
• Gooseberry (Amla) is the richest natural source of vitamin C (≈ 252 mg/100 g).
• Variation exists from fruit to fruit, but all fresh fruits contain some ascorbic acid.
• Loss of Vitamin C occurs when fruits are:
• Bruised or peeled,
• Cooked or exposed to air, alkali, or copper.
Antioxidant Value
•Antioxidant activity measured as DPPH (Trolox equivalent mg/100 g).
•Guava shows the highest antioxidant activity among fresh fruits.
Water Content
• Fruits contain 75–90 % water.
• Water dissolves sugars, salts, organic acids, and pigments within vacuoles.
• Insoluble substances remain colloidally dispersed.
• Pigments in Fruits
• Like vegetables, fruits owe their colour to several pigment groups.
Chlorophyll
• Found in guava, gooseberry, and country apple.
• Imparts green colour, though masked by carotenoids in ripe fruits.
Carotenoids
• Present in mango, papaya, orange, watermelon (lycopene), muskmelon (β-
carotene), jackfruit, peach (violaxanthin), tomato, grape pink (lycopene + β-
carotene), pineapple (violaxanthin + β-carotene).
• Contribute yellow, orange, and red hues.
Anthocyanins
• Found in grapes, blueberries, plums, and cherries.
• Provide red-blue-violet shades.
Anthoxanthins
• Present in guava, apple, gooseberry, pear, custard apple, banana.
• Yield pale yellow to creamy colours.
CELLULOSE AND PECTIC
SUBSTANCES
• Fruit framework built from cellulose forming the cell-wall matrix.
• Hemicellulose and pectic substances fill intercellular spaces and bind cells.
Pectic Compounds
• Include protopectin, pectinic acid, and pectin.
• Act as cementing substances giving firmness.
• Heat or ripening converts insoluble protopectin to soluble pectin, softening
fruit.
• Acids make tissue firm; alkalis disintegrate fibres.
• Apples and guavas are rich in pectin (a soluble dietary fibre).
FLAVOUR CONSTITUENTS
Volatile Compounds
• Include esters, aldehydes, acids, alcohols, ketones, ethers.
• Example: jackfruit flavour due to 13 esters (mostly 3-methyl butanoates), 9
alcohols, 5 aldehydes, 5 acids, and 2 acetyl-1-pyrroline compounds.
Other Influencing Substances
• Sugars, tannins, acids, and mineral salts also modify fruit flavour.
• Some fruits have essential oils contributing to characteristic aroma.
POLYPHENOLS
Nature and Occurrence
• Include tannins, catechins, leuco-anthocyanins, hydroxy acids.
• Present in apples, peaches, grapes, almonds, pears, absent in most herbs.
• Confer astringency and affect colour and flavour.
• Skins and seeds are high in tannin content.
Effects on Fruit Quality
• Cause a puckering sensation (astringency) in mouth.
• Lead to haze or precipitate formation in juice, beer, and wine due to protein-
phenolic interactions.
• Cause brown discolouration in cut fruits via oxidation (chlorogenic acid, catechins).
BITTERNESS IN FRUITS AND
VEGETABLES
Compounds Responsible
• Limonoids (triterpenes) and flavanone glycosides (naringin).
• Found especially in citrus fruits (grapefruit).
Mechanism
• Bitter flavour develops when precursor (limonoate A-ring lactone) reacts with acid
during juice extraction, forming limonin.
• Naringin, a major flavonoid in grapefruit, also contributes to bitterness.
Other Examples
• Cucumber contains cucurbitacins (oxygenated tetracyclic triterpenes) that impart
bitterness.
• Bitter compounds may exist both free and glycosidic forms.
IMPORTANCE OF VEGETABLES
• Vegetables are important for human health because oftheir vitamins,
minerals, phytochemical compounds, and dietary fiber content.
• Especially antioxidant vitamins (vitamin A, vitamin C, and vitamin E).
• Adequate vegetable consumption protects from some chronic diseases
such as diabetes, cancer, obesity, cardiovascular diseases.
ANTI-NUTRIENTS
• “Anti-nutrients” are substances that can block the absorption of
nutrients. Anti-nutrients are naturally found in animals and many
plant-based foods. In plants, they are compounds designed to
protect from bacterial infections and being eaten by insects.
• Glucosinolates in cruciferous vegetables (broccoli, cabbage) can
prevent the absorption of iodine, which may then interfere with
thyroid function and cause goiter. Those already with an iodine
deficiency or a condition called hypothyroidism are most susceptible.
• Lectins in legumes (beans, peanuts, soybeans), whole grains can
interfere with the absorption of calcium, iron, phosphorus, and zinc.
• Oxalates in green leafy vegetables, tea can bind to calcium and prevent it
from being absorbed.
• Phytates (phytic acid) in whole grains, seeds, legumes, some nuts can
decrease the absorption of iron, zinc, magnesium, and calcium.
• People who are at high risk for diseases related to mineral deficiencies, such
as osteoporosis with calcium deficiency or anemia with iron deficiency, may
wish to monitor their food choices for anti-nutrient content.
References
• Srilakshmi, B. (2019). Food Science (8th
Edition). New
Age International Publishers.
•Thank You