Pteridophytes were the first vascular plants to evolve, originating around 400 million years ago. They are characterized by having well-developed vascular tissue for transporting water and nutrients. Pteridophytes reproduce via spores and have alternation of generations, with both gametophyte and sporophyte generations. They differ from other plants in being non-flowering and non-seed producing. Common examples include ferns, clubmosses, and horsetails. Pteridophytes first dominated terrestrial ecosystems but now many species thrive in moist, shaded environments.