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ANOMALOUS
SECONDARY GROWTH
Saloni Rani Swain
Botany Student
President science college, Ahmedabad, Gujarat.
TOPIC
Introduction
Dracaena stem(Monocot)
Bougainvillea and Boerhavia stem(Dicot)
Raphanus and Beet (Root)
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Introduction
Anomalous or Abnormal secondary
growth doesn’t follow normal secondary
growth in which vascular cambium produces
xylem on inside and phloem on outside.
ABNORMAL GROWTH
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STEM ROOT
Monocot Dicot
Dracaena
Boerhaavia Bougainvillea
Raphanus Beet
DRACAENA
(MONOCOT)
PRIMARY
VASCULAR BUNDLE
❑ It is a monocot plant but due to abnormal growth of
cambium it behave like a dicot.
❑ Roughly circular
❑ Epidermis is made up of single layered.
❑ Ground tissue – Parenchyamatous cell.
❑ Primary vascular bundle are collateral, conjoint, closed and
endarch.
❑ They are scattered in parenchyma ground tissue. Each
primary bundle is enclosed with thick-walled
sclerenchymatous sheath.
SECONDARY
GROWTH
❑ In old stem, epidermis replaced by formation of periderm
(cork cambium – phellogen).
❑ Multilayered cortex.
❑ Old stem cambium develop outside the primary vascular
bundle.
❑ Cambium ring form and it divide into two side outer and
inner:
if talk about outer side of cambium then found little
parenchyma and conjunctive tissue is present inner side.
❑ Special type of stele present in dracaena such as
Concentric Amphivasal vascular bundle( means secondary
phloem inner side and secondary phloem outside )
MICROSCOPIC VIEW
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BOERHAAVIA
(DICOT)
PRIMARY GROWTH
o Border- Nearly circular
o Epidermis- Uniseriate, covered with cuticle, a few multicellular hair present.
o Cortex- Outer 3-5 layer collenchyma(hypodermis) and inner layer is made up
chlorenchyma cell.
o Endodermis and Pericycle are not distinct.
o Primary vascular bundle – Conjoint, collateral, Open, Endarch.
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Three Vascular rings are form
Outside from
cambium there 15 to
20 small vascular
bundles are present
Middle ring – 6 to 14 ,
oval in shape. It help
in secondary growth
Inner ring – 2 to 3
larger vascular
bundle (medullary
ray)
The protoxylem of
these bundle face
each other (help in
secondary growth)
SECONDARY GROWTH
Cambium
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Secondary phloem – It present outer side ,thus it cut
parenchymatous ground tissue.
Secondary xylem- inner side and parenchymatous tissue
(lignified)
The activity of this cambium may stop and new cambium ring
formed
❖ SUCCESSIVE
CAMBIUM RING
ARISE.
PITH LARGER IN
SIZE AND MADE UP
OF PARENCHYMA
CELL
MICROSCOPIC
VIEW
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BOUGAINVILLEA
(DICOT)
PRIMARY GROWTH
• It is similar to Boerhavia ,but minor difference can be seen like, inside the stele
only two vascular bundles are form instead of three.
• Epidermis- single layered (cuticle, thick, made up of parenchyma).
• Hypodermis- Few layer thick (collenchyma).
• Endodermis- Not distinct yet present.
• Pericycle- Heterogenous arrangement of tissues (Parenchyma and sclerenchyma
cell alter to each other).
• Vascular bundle - Irregularly arrange.
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SECONDARY GROWTH
• Successive rings of Accessort cambia form.
• Secondary growth - Formation of first cambial ring which is extra stelar in origin.
• Cambium arise from pericycle.
• First cambium life is too short
• Secondary parenchyma- fusiform.
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when new
cambium form it
break from interally
and externally
Internally-
secondary xylem
alternate to
parenchyma
Externally-
secondary phloem
above secondary
xylem and
parenchyma
• Thus successive concentric rings of
vascular bundles are formed thick walled
conjuctive tissue.
• New cambium never make complete ring
surrounding the pericycle it attach to
above cambium
• In some species phloem can not be
distinguished.
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MICROSCOPIC VIEW
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RAPHANUS
(ROOT)
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RAPHANUS
• It is a dicot root.
• Primary xylem and phloem tissue arranged alternative.
• Arrangement of protoxylem on outer side pr towards the cortex. It is called exarch.
• During secondary growth the cells of pericycle convert in cambium.
• The cells of upper portion of the xylem and cells of lower protion of phloem convert in cambium ring.
• The first ring of cambium behave normal it produce less amount of secondary phloem on outer side and
higher amount of secondary xylem inner side.
• The parenchymatous cells of the secondary phloem produced remaining all cambium ring it is known as a
accessory cambium.
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➢ The all accessory cambium behave
anomalous or abnormal function it
produce less amount of secondary
phloem on outer side and higher
amount of secondary in inner side.
➢ Im secondary xylem the component of
xylem parenchyma is higher it a
anomalous of the root.
➢ The function of secondary xylem
parenchyma is storage the food it is
also a anomaly.
➢ Due to higher amount of secondary
xylem parenchyma the Raphanus root
look thick and fleshy.
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RAPHANUS IN
MICROSCOPE
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BEET
(ROOT)
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BEET
• It is a dicot root, thus storage zone is present.
• Secondary growth in these storage root is anamalous.
• Primary structure- Cotyledon type root with radical diarch.
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Secondary
Abnormal activity
of cambium
Abnormal
position of
cambium
• Secondary growth is initiated by the formation of a cambial ring.
• Cambial ring formed pericycle outside protoxylem and parenchyma
between the xylem and phloem.
• Primary cambium--- Abnormal
• Collateral, Conjoint.
• Cambium after sometime ceases its activity.
• Then second arise just outside the first parenchyma or outer ring of
pericycle.
• Second-- Abnormally behave like first.
• Grow continiuosly parenchyma cell.
• Dark red colour just because of Anthocyanin.
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Inner side- sec. xylem and
parenchyma alternative.
Outer side- sec. xylem opposite to
sec. xylem and parenchyma.
BEET ROOT
MICROSCOPIC VIEW
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REFERENCE
• By S.N.PANDEY
• https://images.app.goo.gl/T8DqP8i4n1hhoMBEA
• https://images.app.goo.gl/ffE49jVmjeLj3uDLA
• https://images.app.goo.gl/vw4X7z8zjnB6X2Bh7
• https://images.app.goo.gl/YwTJzYb2qvon5BbQ9
• https://images.app.goo.gl/ys7QYm1YYNG91ut2A
• https://images.app.goo.gl/JKfyKaXRNUMLDMEv9
• https://images.app.goo.gl/sD1H3V4YQnLVVnK59
• https://images.app.goo.gl/XsiysexqJt4HLbbA6
• https://images.app.goo.gl/9GHC9EQNXMEKXMLW7
• https://images.app.goo.gl/3oeEwJr9QzCrz1ho6
• https://images.app.goo.gl/YWhKPxxd5K7jFZvn6
• https://images.app.goo.gl/BMfC5FmZXhh8bGvJA
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THANK
YOU
Saloni Rani Swain
saloniswain225@gmail.com