The Roman Empire fell in 476 AD, leading to a loss of knowledge and the rise of oral tradition in medieval music. Notation systems evolved from un-heightened neumes to square notation to capture melodies. Early medieval music was mostly monophonic and focused on beautiful, sacred melodies. Hildegrad von Bingen was a German nun who composed non-traditional melodies mimicking the inflected voice. Leonin and Perotin were French composers who invented the first polyphonic music through organum, writing multi-part chants in Ars Antiqua style in The Great Book of Organum.