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The Knowledge Network (or Community of Practice) is a powerful organizational model for organizations to innovate, coordinate, build capacity, and translate ideas rapidly into action. On November 30, 2011 Kate Pugh of AlignConsulting and Columbia University led a Nonprofit Webinar on "knowledge networks." We introduced recent research funded by the Bill and Melinda Gates Foundation, and discussed the six behaviors of an enduring network: common objectives, collaborative behavior, working platform, cohesion, generous connectivity, and trust. More than half of all webinar participants were concerned about the levels of trust in their networks, and the resulting compromised collaboration. What builds trust in a network? What erodes it? What can we design into the network -- into its structures, relationships, measures -- to build trust and trustworthiness? We'll look at recent research on the mechanics and science of trust, and look at several case studies of networks where trust translates into volunteerism, innovation, and collaboration. We'll also revisit each of the 8 network design dimensions with a trust lens.