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TDikow Afrotropical Mydidae ICD 2010
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TDikow Afrotropical Mydidae ICD 2010

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  • 1. 1 An overview of Afrotropical Mydidae – the most diverse mydid fauna world-wide Torsten Dikow Biodiversity Synthesis Center Field Museum of Natural History
  • 2. 2 Overview of presentation ◊ Mydidae in numbers ◊ Afrotropical Mydidae ◊ Syllegomydinae in detail ◊ Summary Eremomidas arabicus Bequaert, 1961 (photo Drew Gardner, UAE) Nomoneura caffra Hesse, 1969
  • 3. . mydidae in numbers 3 Mydidae – “mydas flies” ◊ Imagines flower-feeders, some do not feed at all ◊ Larvae possibly predatory ◊ 472* species and 66 genera world-wide ◊ 11 subfamily taxa ◊ Imagines 9 – 60 mm (wing length up to 52 mm) ◊ World-wide primarily in arid regions ◊ Oldest definitive fossil Cretomydas santanensis › Crato Formation, Brazil › ≈ 110 Million years * includes species currently being described Cretomydas santanensis Willkommen & Grimaldi, 2007
  • 4. .. afrotropical mydidae 4 Afrotropical Mydidae: geographic overview ◊ 206* species (= 44 % of world-wide species diversity) – 32 genera (= 48 %) – 5 subfamily taxa ◊ # species 4 Oman, Yemen, UAE (2 are endemic to Afrotropical Region) 5 Madagascar (all endemic) 4* Kenya (all previously unknown) 179* southern Africa (all endemic)
  • 5. .. afrotropical mydidae 5 Afrotropical Mydidae: taxonomic overview Author A. Gerstaecker M. Bezzi M. Bequaert A. Hesse Year 1868 1924 1938, 1940, 1969, 1972, 1951, 1959, 1974 1963 # species / # genera 7/2 16 / 4 26 / 4 126 / 14 Arenomydas namaquensis Hesse, 1969 taxon Afrotropical World # genera # species # genera # species Ectyphinae 2 7* (- 2, + 1) 4 10 Leptomydinae 3 5 6 51* (+ 3) Megascelinae 1 2 3 14* (+ 1) Rhopaliinae 2 5 4 21 Syllegomydinae 24 182* (+ 5) 25 199 total = 32 206* Afroleptomydas sp.
  • 6. .. afrotropical mydidae 6 Ectyphinae – 7* species Ectyphus Gerstaecker, 1868 – Parectyphus Hesse, 1972 › recorded from eastern Africa (Kenya) for the first time Ectyphus pinguis Gerstaecker, 1868 + Parectyphus namibiensis Hesse, 1972 (photos by Katie Lyons)
  • 7. .. afrotropical mydidae 7 Leptomydinae – 5 species Leptomydas Gerstaecker, 1868 – Eremomidas Semenov, 1896 – Hessemydas Kondratieff et al., 2005 › Afrotropical Eremomidas not properly placed in Leptomydinae Eremomidas arabicus Bequaert, 1961 (photo Drew Gardner, UAE) + Hessemydas seyrigi (Séguy, 1960)
  • 8. .. afrotropical mydidae 8 Megascelinae – 2 species Tongamya Stuckenberg, 1966 – the most enigmatic Afrotropical Mydidae › sister taxa in Australia (Neorhaphiomidas) and Chile (Megascelus) Tongamya miranda Stuckenberg, 1966 + Tongamya stuckenbergi Irwin & Wiegmann, 2001 (African Invertebrates 42: 234)
  • 9. .. afrotropical mydidae 9 Rhopaliinae – 5 species Rhopalia Macquart, 1838 – Perissocerus Gerstaecker, 1968 › small and very distinct Mydidae Rhopalia olivieri Macquart, 1838 + Perissocerus arabicus Bequaert, 1961 (photo Drew Gardner, UAE)
  • 10. .. afrotropical mydidae 10 Syllegomydinae – 187* species › 24 genera (23 are endemic to Afrotropical Region) › 9 genera monotypic › 5 genera with only 2 described species Cephalocera imitata Hesse, 1969
  • 11. ... afrotropical syllegomydinae 11 Syllegomydinae Afroleptomydas Bequaert, 1961 – predominant southern African genus › 88 species (many only known from type locality) › 5 species from Willowmore (Eastern Cape, South Africa) Afroleptomydas suffusipennis (Brunetti, 1929)
  • 12. ... afrotropical syllegomydinae 12 Syllegomydinae: Mydaselpidini Afromydas Bequaert, 1961 – Mydaselpis Bezzi, 1924 – Vespiodes Hesse, 1969 – 16* species › wasp-like ‘waist’ – largest Afrotropical Mydidae Mydaselpis peringueyi Bezzi, 1924 + Mydaselpis sp. nov. (Namibia)
  • 13. ... afrotropical syllegomydinae 13 Syllegomydinae: interesting taxa Eremohaplomydas Bequaert, 1959 – Haplomydas Bezzi, 1924 – Lachnocorynus Hesse, 1969 – 4 sp. › very distinct – not properly placed in Syllegomydinae Haplomydas crassipes Bezzi, 1924 + Eremohaplomydas desertorum Bequaert, 1959
  • 14. ... afrotropical syllegomydinae 14 Syllegomydinae: interesting taxa Namibimydas Hesse, 1972 – Nothomydas Hesse, 1969 – 4 species › very peculiar male terminalia Namibimydas sp. + Nothomydas gariepinus Hesse, 1969
  • 15. ... afrotropical syllegomydinae 15 Syllegomydinae: interesting taxa Agaperemius Hesse, 1969 – Halterorchis Bezzi, 1924 – Heteroleptomydas Bequaert, 1963 – Mahafalymydas Kondratieff et al., 2005 – Mimadelphus Hesse, 1972 – Neolaparopsis Hesse, 1969 – Notosyllegomydas Hesse, 1969 – Oreomydas Hesse, 1969 › only known from type localities or very few specimens Neolaparopsis puncturatus Hesse, 1969
  • 16. ... afrotropical syllegomydinae 16 Syllegomydinae: interesting taxa › several specimens of undescribed species cannot be placed in currently described genera ?genus sp. nov. Ugab (Namibia) + ?genus sp. nov. Gobabeb (Namibia)
  • 17. .... summary 17 Summary ◊ Afrotropical Region is most diverse ◊ Knowledge of western Africa very poor ◊ Still new species to be described ◊ Several species synonymies, particularly in southern Africa, need to be addressed ◊ Phylogenetic research will highlight ‘new genera’ for currently unplaced specimens ◊ I would be happy to study additional specimens from your collection! www.mydidae.tdvia.de Namadytes sp.
  • 18. 18 Acknowledgements ◊ Museum curators and collectors who made specimens available ◊ Ashley Kirk-Spriggs for organising this symposium ◊ Ashley & Mikhail Mostovski for spearheading the Afrotropical Manual ◊ U.S. National Science Foundation (REVSYS DEB-0919333)