10/14/13
	


Diversity And Classification of
Flowering Plants:
Eudicots

Michael G. Simpson

Eudicots
•  Palynological apo...
10/14/13
	


Eudicots
UNPLACED**
Apodanthaceae, Balanophoraceae,
Cynomoriaceae, Cytinaceae,

	


Hoplestigmataceae, Medusa...
10/14/13
	


Diversity And Classification of
Flowering Plants:
Eudicots: Rosids

ROSIDS
	

•  Very large, monophyletic gro...
10/14/13
	


Aphloiaceae
MYRTALES
Geissolomataceae
Alzateaceae
Ixerbaceae
Combretaceae
Picramniaceae
Cryteroniaceae
Strasb...
10/14/13
	


Myrtaceae - Myrtle family
(myrtus, Gr. name for myrtle). 120 genera / 3850 species

•  trees and shrubs	

•  ...
10/14/13
	


Leptospermum laevigatum
	


FABALES
Fabaceae (Leguminosae)
- Bean/Pea family
(after faba, Latin name for broa...
10/14/13
	


Fabaceae: 3 subfamilies
	


Caesalpinioideae	

	

Flowers zygomorphic; petals distinct; posterior petal inner...
10/14/13
	


style	

ovary	


stipe	

Bauhinia variegata Orchid Tree
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
Eudicots rosids fs2013.ppt
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Eudicots rosids fs2013.ppt

  1. 1. 10/14/13 Diversity And Classification of Flowering Plants: Eudicots Michael G. Simpson Eudicots •  Palynological apomorphy: tricolpate or tricolpate-derived pollen grain 1
  2. 2. 10/14/13 Eudicots UNPLACED** Apodanthaceae, Balanophoraceae, Cynomoriaceae, Cytinaceae, Hoplestigmataceae, Medusandraceae, Metteniusaceae, Pottingeriaceae BUXALES** Buxaceae Didymelaceae** Sabiaceae CORE EUDICOTS BERBERIDOPSIDALES** Aextoxicaceae Berberidopsidaceae DILLINEALES** Dilleniaceae GUNNERALES Gunneraceae Myrothamnaceae** TROCHODENDRALES** Trochodendraceae RANUNCULALES Berberidaceae Circaeasteraceae Eupteleaceae CARYOPHYLLALES Achatocarpaceae Aizoaceae Amaranthaceae Ancistrocladaceae Asteropeiaceae Lardizabalaceae Menispermaceae Papaveraceae Ranunculaceae PROTEALES Nelumbonaceae Platanaceae* [Proteaceae] Barbeuiaceae Basellaceae Cactaceae Caryophyllaceae Didiereaceae Dioncophyllaceae Eudicots (continued) Droseraceae Drosophyllaceae Frankeniaceae Gisekiaceae Halophytaceae** Hectorellaceae** Limeaceae** Molluginaceae Nepenthaceae Nyctaginaceae Physenaceae Phytolaccaceae Plumbaginaceae Polygonaceae Portulacaceae Rhabdodendraceae Sarcobataceae Simmondsiaceae Stegnospermataceae Tamaricaceae SANTALALES Erythropalaceae** Olacaceae Opiliaceae Loranthaceae Misodendraceae Santalaceae Schoepfiaceae** SAXIFRAGALES Altingiaceae Aphanopetalaceae** Cercidiphyllaceae Crassulaceae Daphniphyllaceae Grossulariaceae Haloragaceae Hamamelidaceae Iteaceae Paeoniaceae Penthoraceae** Peridiscaceae** Pterostemonaceae** Saxifragaceae ROSIDS (see Table 8.2) ASTERIDS (see Table 8.3) Tetracarpaeaceae** Proteaceae 2
  3. 3. 10/14/13 Diversity And Classification of Flowering Plants: Eudicots: Rosids ROSIDS •  Very large, monophyletic group of Eudicots •  Linked by no clear non-molecular apomorphies •  Ovules bitegmic (2 integuments) & crassinucellate [contrast with Asterids] •  13 orders Michael G. Simpson 3
  4. 4. 10/14/13 Aphloiaceae MYRTALES Geissolomataceae Alzateaceae Ixerbaceae Combretaceae Picramniaceae Cryteroniaceae Strasburgeriaceae Heteropyxidaceae Vitaceae Lythraceae CROSSOSOMATALES Melastomataceae Crossosomataceae (incl. Memecylaceae) Stachyuraceae Myrtaceae Staphyleaceae Oliniaceae GERANIALES Onagraceae Geraniaceae Penaeaceae (incl. Hypseocharitaceae) Psiloxylaceae Ledocarpaceae Rhynchocalycaceae Melianthaceae Vochysiaceae (incl. Francoaceae) Vivianiaceae EUROSIDS I (continued) OXALIDALES Brunelliaceae Cephalotaceae Connaraceae Cunoniaceae Elaeocarpaceae Oxalidaceae ROSALES Barbeyaceae Cannabaceae Dirachmaceae Elaeagnaceae Moraceae Rhamnaceae Rosaceae Ulmaceae Urticaceae EUROSIDS I Zygophyllaceae Krameriaceae* Huaceae CELASTRALES Celastraceae (incl. Lepidobotryaceae) Huaceae Parnassiaceae (incl. Lepuropetalaceae) CUCURBITALES Anisophylleaceae Begoniaceae Coriariaceae Corynocarpaceae Cucurbitaceae Datiscaceae Tetramelaceae EUROSIDS I (continued) FABALES Fabaceae (Leguminosae) Polygalaceae Quillajaceae Surianaceae FAGALES Betulaceae Casuarinaceae Fagaceae Juglandaceae (incl. Rhoipteleaceae) Myricaceae Nothofagaceae Ticodendraceae MALPIGHIALES Achariaceae Balanopaceae Bonnetiaceae Caryocaraceae Chrysobalanaceae (incl. Dichapetalaceae, Euphroniaceae, Trigoniaceae) Clusiaceae (Guttiferae) Ctenolophonaceae Elatinaceae Erythroxylaceae* Euphorbiaceae, s.l. Goupiaceae Humiriaceae Irvingiaceae Ixonanthaceae Lacistemataceae MALPIGHIALES (continued) Linaceae Lophopyxidaceae Malpighiaceae Ochnaceae (incl. Medusagynaceae, Quiinaceae) Pandaceae Passifloraceae (incl. Malesherbiaceae, Turneraceae) Peridiscaceae Phyllanthaceae Picrodendraceae Podostemaceae Putranjivaceae Rhizophoraceae Salicaceae Violaceae EUROSIDS II MALVALES Tapisciaceae Bixaceae BRASSICALES Cistaceae Akaniaceae Dipterocarpaceae [incl. Malvaceae Bretschneideraceae] Mutingiaceae Bataceae Neuradaceae Brassicaceae (Cruciferae) Sarcolaenaceae Caricaceae Sphaerosepalaceae Emblingiaceae Thymelaeaceae Gryrostemonaceae SAPINDALES Koeberliniaceae Anacardiaceae Limnanthaceae Biebersteiniaceae Moringaceae Burseraceae Pentadiplandraceae Kirkiaceae Resedaceae Meliaceae Salvadoraceae Nitrariaceae Setchellanthaceae Rutaceae Tovariaceae Sapindaceae Tropaeolaceae Simaroubaceae 4
  5. 5. 10/14/13 Myrtaceae - Myrtle family (myrtus, Gr. name for myrtle). 120 genera / 3850 species •  trees and shrubs •  with glandular-punctate or pellucid leaves •  Usually epiperigynous flowers with numerous stamens. K 4-5 [3,6] C 4-5 [3,6] A ∞ G (2-5) [(-16)], inferior [rarelyhalf-inferior or superior], with hypanthium. Myrtaceae - Myrtle family (myrtus, Gr. name for myrtle). 120 genera / 3850 species Timber, oil - Eucalyptus spp. edible fruits - Psidium guajava, guava spices - Syzygium aromaticum, cloves, cultivated ornamentals -Callistemon (bottlebrush), Eucalyptus sideroxylon 5
  6. 6. 10/14/13 Leptospermum laevigatum FABALES Fabaceae (Leguminosae) - Bean/Pea family (after faba, Latin name for broad bean). 643 genera / 18,000 species •  trees, shrubs, vines, or herbs, •  stipulate, often compound leaves •  Single, unicarpellous pistil with marginal placentation •  legume (or modified legume) K 5 or (5) [(3-6)] C 5 or (5) [0,1-6, or (1-6)] A 10 or (10) to ∞ [variable] G 1 [2-16], superior, hypanthium sometimes present worldwide distribution ecologically important for nitrogen-fixing rhizobial nodules pulses (e.g., beans, peas, soybeans, etc.), fodder plants, oils, timber trees, gums, dyes, and insecticides. 6
  7. 7. 10/14/13 Fabaceae: 3 subfamilies Caesalpinioideae Flowers zygomorphic; petals distinct; posterior petal inner to laterals; stamens distinct. Mimosoideae Flowers actinomorphic; petals distinct or connate; stamens often ∞, showy; flowers often densely aggregated. Faboideae (=Papilionoideae) Flowers zygomorphic; perianth papilionaceous; posterior petal outer to laterals; stamens connate. Caesalpinioideae Flowers zygomorphic Petals distinct Posterior petal inner to laterals Stamens distinct posterior petal inner to laterals stamens distinct Bauhinia variegata Orchid Tree Bauhinia variegata Orchid Tree 7
  8. 8. 10/14/13 style ovary stipe Bauhinia variegata Orchid Tree

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