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Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications
                (IJERA)              ISSN: 2248-9622         www.ijera.com
                   Vol. 2, Issue 5, September- October 2012, pp.1911-1913
 Aquatic Macrophytes Of Laokhowa Wildlife Sanctuary, Assam,
                           India.
                                       Sanjeeb Kumar Nath
                         Associate Professor, Department Of Botany, Dhing College,
                                       Dhing, Nagaon, Assam.782123.

ABSTRACT
         The wetlands serve as the transitional         category, which are found in abundance in entire
habitats between aquatic and terrestrial                Brahmaputra and Cachar plains.
ecosystems. Wetlands play a significant role in                   Wetlands are potentially rich in aquatic
maintaining the diversity of life on earth. These       resources, which play a significant role in
wetlands thus offer suitable habitats for aquatic       maintaining biodiversity. They offer habitats
life forms. The wetlands not only play a major          suitable for support growth of a variety of aquatic
role in recharging the water regime but also act        life forms. In the recent past, biotic pressures on
as shock absorber for the resuming flood and            these ecosystems have increased tremendously and
erosion. The wetlands of north eastern region of        it has been realized that many species of both plant
India is unique in terms of diversity of aquatic        and animals would be lost prior to their
macrophytes. Laokhowa Wildlife sanctuary                understanding in modern times. A systematic effort
located in central Assam of Nagaon district             to study the flora of these fragile ecosystems and its
harbours diverse form of aquatic macrophytes.           proper assessment is, therefore, necessary for their
Laokhowa Wildlife Sanctuary is intersected by a         better understanding and its proper management for
large number of water bodies locally known as           the welfare of mankind. The documentation and
Beel, duba, nala etc and thus created many              proper assessment of aquatic biodiversity is
submerged and swampy saturated areas                    dependent on respective areas and their habitat. It is
throughout the sanctuary. During the rainy              more fruitful to conserve biodiversity in smaller
season almost all become full of water and the          areas, as it will provide the necessary information
water decreases slowly as winter begins. These          for formulating policies and programs for its
wetlands are habitats and are food source for           effective management and conservation.
many herbivores as well serves as the breeding                    Macrophytes are an important component
grounds for various avifauna, fish fauna and            of an aquatic ecosystem and play a major role in
other aquatic lives. Moreover these wetlands act        primary productivity of the aquatic ecosystem.
as source of livelihood and bioresource for             Wetlands are not only important life-support system
human population. Therefore, it is necessary for        for the flora and fauna alone but also act as sources
their better understanding and its proper               of livelihood and bioresource to supplement their
management. A detail study of macrophytic               day-to-day life for human populations surrounding
diversity of the wetlands of Laokhowa Wildlife          them. Literatures relating to aquatic and wetland
Sanctuary was conducted and recorded along              flora have been carried out by several workers in
with seasonal variation. Altogether 373 plants          various parts of the country (Biswas & Calder,
belonging to 258 genera and 93 families were            1937; Mirashi, 1954; Sen & Chatterjee, 1959;
recorded from the study site.                           Maheswari, 1960; Subramanyam, 1962; Vyas 1964;
                                                        Singh& Tomar 1982; Srivastava et al., 1987;
INTRODUCTION:                                           Samant et al., 1988 and others). Although a
          There are many internationally accepted       considerable amount of on general floristics of
definitions on wetlands of which International          Assam has been done by several workers (Kanjilal
Union for Conservation of Nature and Natural            1934-40; Rao&Rabha 1966), there are only a few
Resources (IUCN) and International Biosphere            contributions on aquatic and wetland flora of Assam
Programme (IBP) are important and accepted more         as a whole ( Pathak, Biswas & Boissya 1987;
in Indian situation. With slight modification of        Baruah and Baruah 2000).
IUCN definition, Assam Remote Sensing
Application Center defined wetlands “All the            STYDY AREA AND LOCATION:
submerged or water saturated lands, natural or                    Laokhowa Wildlife Sanctuary is situated in
manmade, inland or coastal, permanent or                the Nagaon district of Assam, India between the
temporary, static or dynamic, vegetated or non-         latitudes 26030 /N to 26032 / N and longitude 92040 /
vegetated, which necessarily have a land-water          E to 92047 / E in the flood plains of the river
interface”. As such the beel, jalah, doba, doloni,      Brahmaputra. Laokhowa Wildlife Sanctuary (LWS)
pitoni, etc. in Assam could easily be put in wetland    covers a geographical area of 70.1 sq. Km. About 20
                                                        per cent of the Sanctuary is occupied by aquatic or



                                                                                             1911 | P a g e
Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications
                (IJERA)              ISSN: 2248-9622         www.ijera.com
                   Vol. 2, Issue 5, September- October 2012, pp.1911-1913
swampy areas. The swampy and aquatic areas are             Polygonum hydropiper, Polygonum barbatum,
located throughout the sanctuary.                          Polygonum viscosum, Carex spiculata, Saccharum
                                                           spontaneum, Cynodon dactylon, Arundo donax,
MATERIALS AND METHODS:                                     Xanthium indicum, etc.
         The study area was explored thoroughly            (7) Woody species: -Barringtonia acutangula,
and detailed observation on the vegetation and flora       Lagerstroemia reginae etc.
was made. Macrophytes were collected and
preserved according to the conventional herbarium          ACKNOWLEDGEMENT:
techniques (Jain and Rao, 1976). Collected materials                The author is thankful to Chief Conservator
were identified with the help of standard literatures      of Forest Assam and Ranger of Laokhowa Wildlife
and confirmed in the Herbarium of Botanical Survey         Sanctuary for their kind permission to undertake the
of India, Eastern circle, Shillong (Assam). The            research work. Thanks are due to authorities of
cause of degradation of the phytodiversity and their       B.S.I. Shillong for providing access to the
conservation strategies are also worked out.               herbarium facilities.

RESULTS AND DISCUSSION:                                    REFERENCES
         The flora of the study area is mixed type           1.    Baruah, P.P. & Baruah, C.K. 2000. Study
with both aquatic and terrestrial macrophytes.                     of the hydrophytic flora of Kaziranga
Majority of the plants are aquatic herbs intermixed                National Park, Assam, India. Annals of
with terrestrial herbs shrubs and climbers in the                  Forestry 8(2): 170-178.
study areas. Altogether, 373 species belonging to            2.    Biswas, K.P. & Calder, C.C. 1936. Hand
258 genera and 93 families were collected. The                     Book of Common Water and Marsh Plants
aquatic macrophytes of Laokhowa Wildlife                           of India and Barma. Bishen Singh
Sanctuary can be classified in to seven categories                 Mahendra Pal Singh, Dehradun.
following Kaushik (1969, 1995) and Baruah and                3.    Jain, S. K. and Rao, R.R.1976. A Hand
Baruah (2000) on the basis of their location in the                Book of Field and Herbarium Methods,
water body. The macrophytes are categorizes as                     Today and Tomorrow Printers and
follows.                                                           Publishers, New Delhi.
                                                             4.    Kanjilal, U.N., Kanjilal, P.C., Das, A., De,
AQUATIC MACROPHYTES:                                               R.N. and Bor, N.L. 1934-40. Flora of
(1) Free floating hydrophytes: - The common                        Assam,
plants of this category include Eichhornia crassipes,        5.    Vol 1-5. Govt Press, Shillong. Kaushik,
Pistia stratiotes, Lemna perpusilla, Trapa natans,                 J.P. 1969. The vegetation of marshes,
Azolla pinata, Salvinia natans, Utricularia aurea, etc.            ponds and river banks in Shivpuri, Madhya
(2) Free submerged hydrophytes: - The common                       Pradesh. Bull. BSI, 11:84-88.
plants of this category include Ceratophyllum                6.    Kaushik, J.P. 1995. Aqua- Phytodiversity
demersum.                                                          in Northern Madhya Pradesh. In:
(3) Anchored submerged hydrophytes: - The                          Taxonomy and Biodiversity. Pandey,
plants which are submerged but attached to the                     A.K.(ed). CGS Publishers & Distributors,
ground are under this category, which includes                     Delhi, pp.68-72.
plants like Vallisneria spiralis, Hydrilla verticillata,     7.    Maheswari, J.K. 1960. The vegetation of
Ottelia alismoides, Limnoplila indica, Potamogeton                 marshes, swamps and river sides in
crispus etc.                                                       Khandwa District (M.P.) J. Bombay Nat.
(4) Floating anchored hydrophytes: - The rooting                   Hist. Soc. 57: 371-387.
specimen which are floating by their upper parts             8.    Mirishi, M.V. 1954. Studies on the
constitute this category which includes plants like                hydrophytes of Nagpur. J. Indian bot. Soc.
Nymphaea nauchali, Nymphoides hydrophyllum,                        33:298-308.
Nelumbo nucifera, Ipomoea aquatica etc.                      9.    Rao,     A.S.    &     Rabha,     L.C.1966,
(5) Emergent amphibious hydrophytes: - The                         Contribution to the Botany of Kamrup
common plants of this category includes                            District (Southern Part) Assam Bull.
Monocharia hastata, Panicum paludosum, Leersia                     Bot.Surv. Ind.8 (384): 296-303.
hexandra, Typha elephantine, Polygonum posumbu,              10.   Samant, S.S., Rawal, R.S. & Pangtey,
Ipomoea aquatica, Jussiaea repens, Marsilia                        Y.P.S. 1988. Aquatic and Marshy
quadrifolia etc.                                                   Angiospermic Plants of Nainital, Kumaun
(6) Wetland or swampy hydrophytes: - The plants                    Himalaya. In Khulbe, R.D. (Ed)
growing in marshy or wetlands areas and dominated                  Perspective in Aquatic Biology. Papyrus
by Cyperus brevifolius, C. digtatus, C. imbricatus,                Pub House, New Delhi. p.p.409-416.
C. distans, C. iria, C. rotundus, C. kyllingia,              11.   Sen, D.N. & Chetterjee, U.N.1959.
Aeschynomene indica, Alpinia nigra, Alternanthera                  Ecological studies on aquatic and swampy
sessilis, Frimbristylis dichotoma, Phragmites karka,


                                                                                               1912 | P a g e
Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications
             (IJERA)              ISSN: 2248-9622         www.ijera.com
                Vol. 2, Issue 5, September- October 2012, pp.1911-1913
      vegetation of Gorakhpur. A Survey. Agra
      Uni. Res. (Sci) 8:17-27.
12.   Singh, K.K. & Tomar, R.P.S. 1982 The
      aquatic and marsh land flora of Kheri
      District, Uttar Pradesh. J.Bombay Nat.
      Hist. Soc. 79: 271-274.
13.   Srivastava, A.K., S.N. Dixit & Singh, S.K.
      1987. Aquatic angiosperm of Gorakhpur.
      Indian J. For.10: 46-57.
14.   Subramanayam,       K.     1962.  Aquatic
      Angiosperm. CSIR, New Delhi. Pathak
      K.C., Biswas, S. & Boissya, C.L. 1987.
      Systematic studies on the wetland flora of
      Assam. (Abs.) Proc. 33rd Ann. Confer.
      Assam Sci. Soc., March 6-8, 1987:27.
      AAU, Brand. Jorhat.
15.   Vyas, L.N. 1964, A study of the
      hydrophytes and marsh plants of Alawar. J.
      Indian bot. Soc.43: 17-30.




                                                                           1913 | P a g e

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  • 1. Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.1911-1913 Aquatic Macrophytes Of Laokhowa Wildlife Sanctuary, Assam, India. Sanjeeb Kumar Nath Associate Professor, Department Of Botany, Dhing College, Dhing, Nagaon, Assam.782123. ABSTRACT The wetlands serve as the transitional category, which are found in abundance in entire habitats between aquatic and terrestrial Brahmaputra and Cachar plains. ecosystems. Wetlands play a significant role in Wetlands are potentially rich in aquatic maintaining the diversity of life on earth. These resources, which play a significant role in wetlands thus offer suitable habitats for aquatic maintaining biodiversity. They offer habitats life forms. The wetlands not only play a major suitable for support growth of a variety of aquatic role in recharging the water regime but also act life forms. In the recent past, biotic pressures on as shock absorber for the resuming flood and these ecosystems have increased tremendously and erosion. The wetlands of north eastern region of it has been realized that many species of both plant India is unique in terms of diversity of aquatic and animals would be lost prior to their macrophytes. Laokhowa Wildlife sanctuary understanding in modern times. A systematic effort located in central Assam of Nagaon district to study the flora of these fragile ecosystems and its harbours diverse form of aquatic macrophytes. proper assessment is, therefore, necessary for their Laokhowa Wildlife Sanctuary is intersected by a better understanding and its proper management for large number of water bodies locally known as the welfare of mankind. The documentation and Beel, duba, nala etc and thus created many proper assessment of aquatic biodiversity is submerged and swampy saturated areas dependent on respective areas and their habitat. It is throughout the sanctuary. During the rainy more fruitful to conserve biodiversity in smaller season almost all become full of water and the areas, as it will provide the necessary information water decreases slowly as winter begins. These for formulating policies and programs for its wetlands are habitats and are food source for effective management and conservation. many herbivores as well serves as the breeding Macrophytes are an important component grounds for various avifauna, fish fauna and of an aquatic ecosystem and play a major role in other aquatic lives. Moreover these wetlands act primary productivity of the aquatic ecosystem. as source of livelihood and bioresource for Wetlands are not only important life-support system human population. Therefore, it is necessary for for the flora and fauna alone but also act as sources their better understanding and its proper of livelihood and bioresource to supplement their management. A detail study of macrophytic day-to-day life for human populations surrounding diversity of the wetlands of Laokhowa Wildlife them. Literatures relating to aquatic and wetland Sanctuary was conducted and recorded along flora have been carried out by several workers in with seasonal variation. Altogether 373 plants various parts of the country (Biswas & Calder, belonging to 258 genera and 93 families were 1937; Mirashi, 1954; Sen & Chatterjee, 1959; recorded from the study site. Maheswari, 1960; Subramanyam, 1962; Vyas 1964; Singh& Tomar 1982; Srivastava et al., 1987; INTRODUCTION: Samant et al., 1988 and others). Although a There are many internationally accepted considerable amount of on general floristics of definitions on wetlands of which International Assam has been done by several workers (Kanjilal Union for Conservation of Nature and Natural 1934-40; Rao&Rabha 1966), there are only a few Resources (IUCN) and International Biosphere contributions on aquatic and wetland flora of Assam Programme (IBP) are important and accepted more as a whole ( Pathak, Biswas & Boissya 1987; in Indian situation. With slight modification of Baruah and Baruah 2000). IUCN definition, Assam Remote Sensing Application Center defined wetlands “All the STYDY AREA AND LOCATION: submerged or water saturated lands, natural or Laokhowa Wildlife Sanctuary is situated in manmade, inland or coastal, permanent or the Nagaon district of Assam, India between the temporary, static or dynamic, vegetated or non- latitudes 26030 /N to 26032 / N and longitude 92040 / vegetated, which necessarily have a land-water E to 92047 / E in the flood plains of the river interface”. As such the beel, jalah, doba, doloni, Brahmaputra. Laokhowa Wildlife Sanctuary (LWS) pitoni, etc. in Assam could easily be put in wetland covers a geographical area of 70.1 sq. Km. About 20 per cent of the Sanctuary is occupied by aquatic or 1911 | P a g e
  • 2. Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.1911-1913 swampy areas. The swampy and aquatic areas are Polygonum hydropiper, Polygonum barbatum, located throughout the sanctuary. Polygonum viscosum, Carex spiculata, Saccharum spontaneum, Cynodon dactylon, Arundo donax, MATERIALS AND METHODS: Xanthium indicum, etc. The study area was explored thoroughly (7) Woody species: -Barringtonia acutangula, and detailed observation on the vegetation and flora Lagerstroemia reginae etc. was made. Macrophytes were collected and preserved according to the conventional herbarium ACKNOWLEDGEMENT: techniques (Jain and Rao, 1976). Collected materials The author is thankful to Chief Conservator were identified with the help of standard literatures of Forest Assam and Ranger of Laokhowa Wildlife and confirmed in the Herbarium of Botanical Survey Sanctuary for their kind permission to undertake the of India, Eastern circle, Shillong (Assam). The research work. Thanks are due to authorities of cause of degradation of the phytodiversity and their B.S.I. Shillong for providing access to the conservation strategies are also worked out. herbarium facilities. RESULTS AND DISCUSSION: REFERENCES The flora of the study area is mixed type 1. Baruah, P.P. & Baruah, C.K. 2000. Study with both aquatic and terrestrial macrophytes. of the hydrophytic flora of Kaziranga Majority of the plants are aquatic herbs intermixed National Park, Assam, India. Annals of with terrestrial herbs shrubs and climbers in the Forestry 8(2): 170-178. study areas. Altogether, 373 species belonging to 2. Biswas, K.P. & Calder, C.C. 1936. Hand 258 genera and 93 families were collected. The Book of Common Water and Marsh Plants aquatic macrophytes of Laokhowa Wildlife of India and Barma. Bishen Singh Sanctuary can be classified in to seven categories Mahendra Pal Singh, Dehradun. following Kaushik (1969, 1995) and Baruah and 3. Jain, S. K. and Rao, R.R.1976. A Hand Baruah (2000) on the basis of their location in the Book of Field and Herbarium Methods, water body. The macrophytes are categorizes as Today and Tomorrow Printers and follows. Publishers, New Delhi. 4. Kanjilal, U.N., Kanjilal, P.C., Das, A., De, AQUATIC MACROPHYTES: R.N. and Bor, N.L. 1934-40. Flora of (1) Free floating hydrophytes: - The common Assam, plants of this category include Eichhornia crassipes, 5. Vol 1-5. Govt Press, Shillong. Kaushik, Pistia stratiotes, Lemna perpusilla, Trapa natans, J.P. 1969. The vegetation of marshes, Azolla pinata, Salvinia natans, Utricularia aurea, etc. ponds and river banks in Shivpuri, Madhya (2) Free submerged hydrophytes: - The common Pradesh. Bull. BSI, 11:84-88. plants of this category include Ceratophyllum 6. Kaushik, J.P. 1995. Aqua- Phytodiversity demersum. in Northern Madhya Pradesh. In: (3) Anchored submerged hydrophytes: - The Taxonomy and Biodiversity. Pandey, plants which are submerged but attached to the A.K.(ed). CGS Publishers & Distributors, ground are under this category, which includes Delhi, pp.68-72. plants like Vallisneria spiralis, Hydrilla verticillata, 7. Maheswari, J.K. 1960. The vegetation of Ottelia alismoides, Limnoplila indica, Potamogeton marshes, swamps and river sides in crispus etc. Khandwa District (M.P.) J. Bombay Nat. (4) Floating anchored hydrophytes: - The rooting Hist. Soc. 57: 371-387. specimen which are floating by their upper parts 8. Mirishi, M.V. 1954. Studies on the constitute this category which includes plants like hydrophytes of Nagpur. J. Indian bot. Soc. Nymphaea nauchali, Nymphoides hydrophyllum, 33:298-308. Nelumbo nucifera, Ipomoea aquatica etc. 9. Rao, A.S. & Rabha, L.C.1966, (5) Emergent amphibious hydrophytes: - The Contribution to the Botany of Kamrup common plants of this category includes District (Southern Part) Assam Bull. Monocharia hastata, Panicum paludosum, Leersia Bot.Surv. Ind.8 (384): 296-303. hexandra, Typha elephantine, Polygonum posumbu, 10. Samant, S.S., Rawal, R.S. & Pangtey, Ipomoea aquatica, Jussiaea repens, Marsilia Y.P.S. 1988. Aquatic and Marshy quadrifolia etc. Angiospermic Plants of Nainital, Kumaun (6) Wetland or swampy hydrophytes: - The plants Himalaya. In Khulbe, R.D. (Ed) growing in marshy or wetlands areas and dominated Perspective in Aquatic Biology. Papyrus by Cyperus brevifolius, C. digtatus, C. imbricatus, Pub House, New Delhi. p.p.409-416. C. distans, C. iria, C. rotundus, C. kyllingia, 11. Sen, D.N. & Chetterjee, U.N.1959. Aeschynomene indica, Alpinia nigra, Alternanthera Ecological studies on aquatic and swampy sessilis, Frimbristylis dichotoma, Phragmites karka, 1912 | P a g e
  • 3. Sanjeeb Kumar Nath/ International Journal of Engineering Research and Applications (IJERA) ISSN: 2248-9622 www.ijera.com Vol. 2, Issue 5, September- October 2012, pp.1911-1913 vegetation of Gorakhpur. A Survey. Agra Uni. Res. (Sci) 8:17-27. 12. Singh, K.K. & Tomar, R.P.S. 1982 The aquatic and marsh land flora of Kheri District, Uttar Pradesh. J.Bombay Nat. Hist. Soc. 79: 271-274. 13. Srivastava, A.K., S.N. Dixit & Singh, S.K. 1987. Aquatic angiosperm of Gorakhpur. Indian J. For.10: 46-57. 14. Subramanayam, K. 1962. Aquatic Angiosperm. CSIR, New Delhi. Pathak K.C., Biswas, S. & Boissya, C.L. 1987. Systematic studies on the wetland flora of Assam. (Abs.) Proc. 33rd Ann. Confer. Assam Sci. Soc., March 6-8, 1987:27. AAU, Brand. Jorhat. 15. Vyas, L.N. 1964, A study of the hydrophytes and marsh plants of Alawar. J. Indian bot. Soc.43: 17-30. 1913 | P a g e